taxonID	type	description	language	source
03BA87CCFF95676E2AB8EA77D82794FD.taxon	diagnosis	Emended diagnosis. Mouth cone with 9 individual outer oral styles with broader base and thin and flexible anterior part; 14 trichoscalids each originating from trichoscalid plate and accompanied anteriorly by two pointed filamentous appendages; 16 placids with broader midventral placid, neighboured by two narrower placids and alternatingly a broader and a narrower placid; all placids with knobby projections (= condyles) in one or two rows in narrower placids and two or three rows in broader placids; midventral placid with three apical condyles, two in intermediate row, and 3 – 6 in basal row; ventromedial to midventral free flap and primary pectinate fringe of segment 1 at least partly reduced; acicular spine middorsally on segments 1 – 9, midterminally on segment 11 (= midterminal spine), lateroventrally on segments 1 – 9, laterodorsally on segment 10 (male only in species with irregularly distributed or almost no cuticular hairs), and lateral accessorily on segment 11 (= lateral terminal accessory spine); cuspidate spines present; type- 3 sensory spot ventrolaterally next to lateral terminal accessory spine and subdorsally on segment 11; type- 6 sensory spot elevated above trunk surface giving the impression to be half-drowned in the trunk cuticle; ventromedial appendage at least on two segments of segments 5 – 8 in female; area of micropapillae ventromedially on segment 9 in female of species with irregularly distributed or almost no cuticular hairs.	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFF94676E2AB8EF9BD8C4936F.taxon	description	(Figs 4 – 7; Tables 1, 2, 7)	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFF94676E2AB8EF9BD8C4936F.taxon	diagnosis	Diagnosis. Neck placids with condyles in two rows in narrower placids and 2 – 3 rows in broader placids; each narrower placid with one apical and one basal condyle; primary pectinate fringe of all segments reduced, free flap with numerous short spinose or broad extensions on most parts of a segment; acicular spine middorsally on segments 1 – 9 (and on segment 10 in male only), midterminally on segment 11, laterodorsally on segment 10 (in male only), and lateroventrally on segments 1 – 9; cuspidate spine ventrolaterally on segment 5 and lateral accessorily on segment 8; type- 6 sensory spot ventromedially on segments 1 (slightly more paraventrally), 2, 4, 6, 8 and 9, lateroventrally on segment 10, sublaterally on segments 3, 5 – 7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 2, 3 and 7 – 9, and paradorsally on segments 1 (almost subdorsally), 4 – 6, 8 and 9; ventromedial appendage on segments 7 and 8 and area of micropapillae ventromedially on segment 9 in female only.	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFF94676E2AB8EF9BD8C4936F.taxon	materials_examined	Material examined. Holotypic male (MIMB 32339) and paratypic female (MIMB 32340) mounted for light microscopy (Fig. 1; Table 1).	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFAE67542AB8EF25DDDE9377.taxon	description	(Figs 29 – 31; Tables 1, 2, 7)	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFAE67542AB8EF25DDDE9377.taxon	diagnosis	Diagnosis. Neck placids with condyles in one basal row in narrower placids and three rows in broader placids; narrower placids with one basal condyle; broader placids with two condyles each in apical, intermediate, and basal row; midventral placid with three apical and basal condyles and two in intermediate row; acicular spine middorsally on segments 1 – 10, midterminally on segment 11, lateroventrally on segments 1 – 9, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 5 and 9, lateral accessorily on segments 1, 4 and 8, and middorsally on segments 5 and 7; type- 5 (?) sensory spot ventromedially on segment 11; type- 6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 2 – 9, lateroventrally on segment 10, sublaterally on segments 1 – 3, 5 – 7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 1 – 9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2, 4 – 6, 8 and 9.	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFAE67542AB8EF25DDDE9377.taxon	materials_examined	Material examined. Holotypic male (USNM 1025879) mounted laterally (Fig. 1; Table 1).	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFA5675F2AB8EFE3DA70910D.taxon	description	(Fig. 32; Tables 1, 7)	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFA5675F2AB8EFE3DA70910D.taxon	materials_examined	Material examined. One recently moulted male (USNM 1209806) mounted for light microscopy and previously identified as a juvenile stage of Condyloderes paradoxus from the Bay of Bengal, India (Fig. 1; Table 1).	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
03BA87CCFFA5675F2AB8EFE3DA70910D.taxon	description	Description. For some basic measurements see Table 7. Trunk. The trunk is divided into 11 segments. Whereas the cuticle of the first trunk segment is ring-like without any articulation, the cuticle of segments 2 – 10 possesses both a midventral and a lateral articulation resulting in a tergal and two sternal plates (Fig. 32 A). In segment 11, a single sternal and tergal plate exist although the sternal plate reveals a paraventral fold on both sides (Fig. 32 A). The anterior margin of the first trunk segment is smooth (Fig. 32 A). The trunk cuticle and the pachycyclus are only very weakly developed (Fig. 32 A). At the posterior end of all segments, the free flap overlapping the subsequent segment terminates in a primary pectinate fringe. The free flap is missing where the spines originate. Surface characters of the cuticle like hairs were not recognizable. Also, type- 6 sensory spots were extremely difficult to trace if at all because of the thin cuticle. Type- 3 sensory spots occur ventrolaterally next to the lateral terminal accessory spine and subdorsally on segment 11. Segments 1 – 10 bear a middorsal, segments 1 – 9 on each side a lateroventral spine, segment 10 also possesses a subdorsal to laterodorsal spine, and segment 11 has a spine in a lateral accessory position (= lateral terminal accessory spine) and as the midterminal spine (Fig. 32 A, C; Table 7); the spines are hirsute and acicular. A short hirsute cuspidate spine with a basally thickened and an apically thinner part appears ventrolaterally on segment 5 but displaced towards the ventromedial position (Fig. 32 A), in a lateral accessory position on segment 8, as well as sublaterally on segments 3, possibly 4 and 9 (Fig. 32 A; Table 7). Gonads are well developed and extend until the middle of the trunk revealing sperm in the posterior segments (Fig. 32 B). Comparison with previous description. Opposite to Higgins (1969) who identified the single specimen as a juvenile of Condyloderes paradoxus, we regard the specimen as a recently moulted male of a new species. The specimen reveals a thin cuticle typical for a juvenile or a recently moulted adult, a free flap only occurring in adults but not in juveniles, and a well developed gonad with sperm characteristic of a male. The specimen is assumed to belong to a new species, because it lacks the cuspidate spine ventrolaterally on segments 2 and 9 and in a lateral accessory position on segment 4, whereas C. paradoxus possesses a cuspidate spine in these positions. In addition, the specimen reveals a sublateral cuspidate on segments 3, possibly 4 and 9 lacking in C. paradoxus, and the ventrolateral cuspidate spine on segment 5 is displaced towards the ventromedial position in comparison with C. paradoxus. Sublateral cuspidate spines do not occur in any other known species of Condyloderes (Table 7; Higgins 1969; Adrianov et al. 2002; Martorelli & Higgins 2004; Sørensen et al. 2010 b; Adrianov & Maiorova 2016). The specimen is not described formally as a new species, because only a single specimen is available, many cuticular characters are hard to recognize if at all also because of the thin cuticle, the specimen is covered with detritus hampering observation, and the preparation is not too satisfying.	en	Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi, Higgins, Robert P. (2019): Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov. Zootaxa 4561 (1): 1-91, DOI: 10.11646/zootaxa.4561.1.1
