taxonID	type	description	language	source
03BA8793F810FFA1FF0D620F49D0FE4D.taxon	materials_examined	Type species: Cteïpolia sacelli Staudinger, 1896, by monotypy. Notes. (1) The genus name was originally spelled with the letter ï, and this spelling has subsequently been used by Ronkay et al. (1995, 2014) and Gordeeva et al. (2023). However, according to the ICZN (1999) Article 11.2, a scientific name must have been spelled only in the 26 letters of the classic Latin alphabet, which does not include ï. Thus, following the ICZN (1999) Articles 27 and 32.5.2, the letter ï must be amended to i and the correct spelling of the genus name is Cteipolia. (2) In the Introduction chapter of their paper, Gordeeva et al. (2023) state “ the close affinity of Cteipolia with the genera of the Agrochola-Conistra clade of Xyleninae ” while in the Results chapter of the same work, the authors indicate “ the proper taxonomic position of the genus in the tribe Xylenina Guenée, 1837, subtribe Conistrina Beck, 1996 ”. According to the current Noctuidae systematics, the ‘ Agrochola-Conistra generic complex’ (sensu Ronkay et al. (2001 )) belongs to the subtribe Xylenina Guenée of the tribe Xylenini Guenée (Lafontaine & Schmidt 2010; Fibiger et al. 2011; Keegan et al. 2021) while the name Conistrina is a junior synonym of Xylenina. Thus, the correct placement of genus Cteipolia is in the subtribe Xylenina, tribe Xylenini, subfamily Noctuinae. (3) Fibiger et al. (2010) and Gordeeva et al. (2023) erroneously attributed the authorship of C. murina to Eduard Eversmann. For the correct authorship, see Titov et al. (2024). (4) Male genitalia of Cteipolia are very uniform within species-groups and in most cases display no reliable interspecific differences, which makes the species delimitation problematic. Unlike in males, female genitalia are diagnostic in their signum, antrum and apophysis structures. Males of similar species have somewhat differently ciliate antennae and this fact, along with the uniform male genitalia, allows to assume that, similar to some other Noctuinae genera such as Euxoa Hübner, [1821], Agrotis Ochsenheimer, 1816, certain species-groups of Xestia Hübner, 1818 (Noctuini), the tribe Episemini Guenée, 1852, etc., the pre-copulatory isolation mechanisms (species-specific pheromones) in Cteipolia are stronger than the genitalia “ lock- and-key ” barrier (Mikkola 2008).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F810FFA1FF0D620F49D0FE4D.taxon	diagnosis	Diagnosis. The genus is described in details by Ronkay et al. (1995). Species of the genus are relatively small and characteristically looking moths with forewing having almost parallel margins, which is markedly narrower than in similarly small Dasypolia species. The male genitalia of Cteipolia are characterised by the following characters. (1) The distal section of the valva is narrow (narrower than in Dasypolia). (2) The editum is short and narrow and bearing only a proximal setose tubercle-like ampulla whereas in Dasypolia it is extended distally and bears a digitus-like distal ampulla protruding beyond the ventral margin of the valva. (3) The phallus has ribbon-like carinae protruding into the basal section of the vesica, which are invaginated in the phallus tube in the resting condition, whereas the phallus of Dasypolia bears a heavily sclerotised and dentate carinal plate. (4) The vesica is simple, tubular and membranous whereas in Dasypolia, it has one or two diverticula and may also bear clusters of minute graniculi. Compared to Dasypolia s. str., the female genitalia of Cteipolia have an elongate, telescopic ovipositor with long apophyses posteriores (vs. short and conical ovipositor in Dasypolia s. str.), a membranous ductus bursae with a well-developed antrum (whereas in Dasypolia s. str. most of the ductus bursae is sclerotised and dorso-ventrally flattened), and a corpus bursae bearing one or two signa, which are absent in Dasypolia s. str.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F810FFA1FF0D620F49D0FE4D.taxon	distribution	Distribution. The genus is distributed from eastern Turkey and Armenia in the west, through Central Asia and southern Ural Mountains to South Siberia (Transbaikalia) and Pakistani Himalaya in the east and south, respectively (Ronkay et al. 1995, 2014; Gordeeva et al. 2023). Bionomics. Adults of both sexes fly in October; females overwinter and are active in March – April. A male of an unidentified Cteipolia species was observed on 8 March 2021 flying in the day time during a thaw in the Russian part of the Altai Mountains (Titov et al. 2024). Although Gordeeva et al. (2023) stated that ‘ all representatives of the genus are mountain species’, at least two of them, viz. C. isotima Püngeler, 1914 and C. murina (Ménétriés, 1848) are found to occur in flat landscapes with sandy soils in the Balkhash-Alakol Basin in Southeast Kazakhstan and Irgiz River valley in West Kazakhstan, respectively (Titov et al. 2024). Nevertheless, the majority of species are found in rocky steppe habitats in hilly or mountain landscapes at the altitudes ranging from 480 – 2400 m a. s. l. The early stages are unknown except for C. amissa Gordeev, Gordeeva, G. Ronkay & L. Ronkay, 2023, the eggs, larvae and pupa of which are illustrated by Gordeeva et al. (2023). In the laboratory conditions, the authors fed the larvae on the leaves of the genus Salix (S. Gordeev, pers. comm., also see: Gordeeva et al. (2023): plate 37, figs 4 – 7) but the food plant in nature remains unknown and may be different. However, in the majority of places of Cteipolia occurrence, if not all, willow thickets have been observed by the authors of the present paper. The larva of C. amissa is free-living (see: Gordeeva et al. (2023): plate 37, figs 4 – 7), what was the reason for raising the status of Cteipolia to genus level and transferring it into from the subtribe Antitypina into Xylenina (in contrast, larvae of Dasypolia are endophagous and tunnel the stems or rhizomes of their host plants).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA1FF0D66204E18FA3E.taxon	diagnosis	Diagnosis. The species-group is characterised by the narrow, finger-shaped uncus, and the apically tapered harpe in males, and the short and bowl-shaped antrum in females.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA4FF0D66D949A9FEE4.taxon	description	(Figs 1, 2, 17)	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA4FF0D66D949A9FEE4.taxon	materials_examined	Type material. Holotype (Figs 1, 17): female, “ Armenia | Old Jermuk road, | h- 1340 m, 39.6998741 N | 45.5594319 E | 22,25. iii. 2023 | Duda & Saldaitis leg. ” / “ Slide | AV 7520 ♀ | A. Volynkin ” (WIGJ). Paratypes: 6 females, same data as holotype, gen. prep. No.: AV 7519 ♀ (ASV and WIGJ).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA4FF0D66D949A9FEE4.taxon	diagnosis	Diagnosis. The forewing length is 14.0 – 14.5 mm in females. Cteipolia aloyani sp. n. (Figs 1, 2) is most morphologically similar to C. tertia (Figs 3, 4) described from Tajikistan and superficially differs from it in the more brownish forewing ground colour, and the more outbent postmedial line of the forewing. Compared to the geographically closest known congener, the Turkish C. vera (Figs 5, 6), the new species has broader forewing with brown ground colour (it is reddish brown in C. vera), more diffuse transverse lines of the forewing, a longer and thicker reniform stigma with a thinner pale edge, and a thinner and more diffuse discal spot of the hindwing. In the female genitalia, C. aloyani sp. n. (Fig. 17) is distinguished from C. tertia (Fig. 18) by the shorter ovipositor, the longer apophyses anteriores, the posteriorly broader antrum with a V-shaped ventral margin (it is rounded in the congener), the shorter ductus bursae, and the narrower signum bursae. Compared to C. vera (illustrated by Ronkay et al. (1995) and Gordeeva et al. (2023 )), the female genitalia of C. aloyani sp. n. have a somewhat longer ovipositor, considerably longer apophyses anteriores, a markedly shorter, bowl-shaped antrum (it is elongate and with parallel margins in the congener), a longer ductus bursae, and a single and small signum bursae whereas the corpus bursae of C. vera bears two long and thick signa. The male is unknown.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA4FF0D66D949A9FEE4.taxon	distribution	Distribution. The new species is currently known only from its type locality in south-western Armenia. Bionomics. Female adults were attracted to artificial light in the evening twilight, which corresponds to the behavior of another species in the genus, C. isotima observed earlier (Titov et al. 2024). The collection site is situated on southern slopes of the Vardenis Range in the Arpa River Canyon in its middle flow (Fig. 29). It is characterised by rich flora and very mosaic vegetation. Ecosystems of the canyon were classified according to EUNIS habitats’ classification (Davies et al. 2004) adapted to Armenia by Fayvush & Aleksanyan (2016). Along the banks of Arpa river Mesotrophic, the vegetation of slow-flowing rivers (C 2.33) with Potamogeton spp., Myriophyllum spicatum L., etc. is presented. The main vegetation type in the bottom of the gorge is Armenian willow galleries (G 1.1143) with dominance of Salix spp. such as Salix alba L. and Salix excelsa S. G. Gmel. Besides willows, Populus nigra L. and Populus alba L., are present in the tree stand. Small plots of Salix spp. and Tamarix thickets (F 9.141 and F 9.3142, respectively) with Salix caprea L., S. purpurea L., S. viminalis L., S. wilhelmsiana M. Bieb., as well as Tamarix ramosissima Ledeb. are present in the gorge. Slopes of the gorge are covered with mosaic of the following types of vegenation. (a) Mixed open forests with Juniperus spp. and deciduous trees and shrubs (G 4.92). Besides the dominating Juniperus polycarpa K. Koch, the species characteristic of shibliak and arid open forests are present, namely Amygdalus fenzliana (Fritsch) Lipsky, Paliurus spina-cxhristi Mill., Prunus divaricate (Ledeb.) C. K. Schneid., and Rhamnus pallasii Fisch. & C. A. Mey., as well as single individuals of Celtis caucasica Willd., Fraxinus excelsior L., etc. (b) Traganth communities with dominance of Astragalus microcephalus Willd. (F 7.41211). Besides Astragalus itself, Onosma sericea Willd., Poa bulbosa L., Stachys inflata Benth., etc. are common is this habitat, penetrating also into the plots of mixed forest.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F811FFA4FF0D66D949A9FEE4.taxon	etymology	Etymology. The new species is named after Mr Artak Aloyan (Jeghegnadzor, Armenia), patriot of his country and sincere friend of second and thirds coauthors.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F815FFAAFF0D647148ECFE28.taxon	diagnosis	Diagnosis. Adults of the species-group are distinguished from the C. tertia species-group by the pale, eyespot-like orbicular and reniform stigmata having a thin dark core and a broad pale edge, whereas in the tertia species-group, the dark cores are markedly longer and thicker while the pale edge is thin. The male genitalia ground plans of the two species-groups are alike but in the female genitalia of the C. murina species-group, the antrum is elongate and rather champagne glass-shaped whereas it is short and bowl-shaped in the C. tertia species-group.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F81BFFA8FF0D67D94F57FE82.taxon	description	(Figs 7, 8, 19)	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F81BFFA8FF0D67D94F57FE82.taxon	materials_examined	Type material. Holotype (Figs 7, 19): female, “ 19. III. 2024, | SE Kazakhstan, | Zhetysu Region, | Sholak Mts, 750 m, | 43 ° 54 ' 57.6 '' N | 77 ° 47 ' 47.8 '' E, | S. V. Titov leg. ” / “ Slide | AV 7573 ♀ | A. Volynkin ” (WIGJ, ex CAV). Paratypes: 2 females, same data as holotype, gen. prep. No.: AV 7551 ♀ (CAV).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F81BFFA8FF0D67D94F57FE82.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 14.0 mm in females. Cteipolia belyalovi sp. n. (Figs 7, 8) is reminiscent of C. murina (Figs 11, 12), C. isotima (Fig 9, 10) and C. amissa (Fig. 13) but can be distinguished by its somewhat narrower forewing with a more elongate apex, the paler, ochreous brown forewing ground colour with intense irroration of brown scales, and the more distinct hindwing discal spot. The female genitalia of the new species (Fig. 19) differ clearly from the similar congeners (Figs 20 – 24) in the presence of a single signum bursae (vs. two signa in C. murina, C. isotima and C. amissa). Additionally, compared to C. murina and C. isotima, the new species has a slightly broader posterior end of the antrum. The male is unknown.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F81BFFA8FF0D67D94F57FE82.taxon	distribution	Distribution and bionomics. The new species is currently known only from its type locality in Southeast Kazakhstan. The collecting site is situated in the Sholak Mountains, which are dry, old, decaying, low massif at the western extremity of the Altyn-Emel Range belonging to the Dzhungar Alatau mountain range (Fig. 30). The mountains have dry stony steppe slopes dominated by Artemisia L. spp. and with scarce low shrubs of Spiraea hypericifolia L., Rosa platyacantha Schrenk, R. alberti Regel, Lonicera microphylla Roem. & Schult., Caragana balchaschensis (Kom.) Pojark., Cerasus tianshanica Pojark., Salix songarica Andersson, S. tenuijulis Ledeb., and S. wilhelmsiana M. Bieb. (Goloskokov 1984).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F81BFFA8FF0D67D94F57FE82.taxon	etymology	Etymology. The new species is dedicated to the memory of the late Oleg V. Belyalov (Almaty, Kazakhstan), close friend of Sergey Titov, excellent wildlife photographer, documentary filmmaker and cameraman, botanist and zoologist, and populariser of the knowledge of the wildlife of Kazakhstan.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F818FFA9FF0D66144B95FD90.taxon	description	(Figs 9, 10)	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F818FFA9FF0D66144B95FD90.taxon	materials_examined	Type material examined. Lectotype (designated by Ronkay et al. (1995) as ‘ Holotype’): male, “ Tien- schan | (Dscharkent) | Rückbeil 1903 ” / pink label “ Type | isotima Püng. ♂ ” / “ Zool. Mus. | Berlin ” / “ Préparation | № M. B. 296 | Ch. Boursin ” (MfN); paralectotype (designated by Ronkay et al. (1995) as ‘ Paratype’): female (Figs 9, 20), “ Tien- schan | (Dscharkent) | Rückbeil 1903 ” / pink label “ Cotype | isotima Püng. ♀ | abgebildet Iris 1914 / Püng. ” / “ isot / 1904 v. R. Tancré Püng. ” / “ my section II | Hmpsn. i. l. | 7 - 1914 ” / “ Zool. Mus. | Berlin ” / “ 4301 ♀ | gen. prep. No. | det. L. Ronkay ” (MfN). Additional material examined: 1 male, 4. X. 2015, SE Kazakhstan, Almaty [currently Zhetysu] Region, Dzhungar Alatau, Cherkasay Valley, 1580 m, 44 ° 46 ' 33 '' N 78 ° 55 ' 59 '' E, S. V. Titov & M. Černila leg. (CAV); 2 males, 05. X. 2023, E Kazakhstan, Monrak Ridge, N slope, 24 km W of Karabulak vill., N 47 ° 33 ' 54.0 '' E 84 ° 21 ' 18.4 '', 735 m, gorge with scree slopes, S. V. Titov leg. (CAV); 5 males, 2 females, 9. X. 2023, E Kazakhstan, East Kazakhstan Reg., Saur Mts, 753 m, 3 km SE of Saryterek vill., 47 ° 27 ' 46.6 '' N 85 ° 12 ' 41.5 '' E, S. V. Titov leg. (CAV); 1 male, East Kazakhstan, Saur Mts, 20 km S of Zaisan town, Bol’shoi Zhemenei River, 1600 m, 47 ° 17 ' N 84 ° 54 ' E, 5. X. 2018, P. Gorbunov leg. (PGC); 1 female, same data as previous but S. Melyakh leg. (SMC); 2 males, East Kazakhstan, Manrak Mts, 20 km SW of Tugyl settl., Kusty River bank, 900 m, 47 ° 34 ' N 84 ° 04 ' E, 6. X. 2018, P. Gorbunov leg. (BBС); 5 males, East Kazakhstan, Tarbagatai Mt. Range, Bazar River bank, 32 km S of Kyzykesik, 1000 m, 47 ° 35 ' N 82 ° 04 ' E, 7. X. 2018, P. Gorbunov leg. (BBС, SMC).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F818FFA9FF0D66144B95FD90.taxon	discussion	Remarks. (1) In the original description of the species, Püngeler (1914) cited a pair of specimens therefore Titov et al. (2024) treated the type specimens are syntypes. However, Püngeler labelled the male specimen as “ Type ” and the female as “ Cotype ”, and Ronkay et al. (1995) considered those specimens as holotype and paratype, respectively, and, following the ICZN (1999) Article 74.5, this act constitutes a valid lectotype and paralectotype designation. (2) Gordeeva et al. (2023) defined the type locality of C. isotima Püngeler, 1914, “ Dscharkent ” as situated in Kyrgyzstan (spelled by the authors as “ Kirghisia ”) while Zharkent is in fact a well- known city in the southern foothills of the Dzhungar Alatau Range in Southeast Kazakhstan.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F818FFA9FF0D66144B95FD90.taxon	distribution	Distribution. Currently known from Southeast and East Kazakhstan and found in the Dzhungar Alatau and Saur-Tarbagatai mountain ranges, and south-eastern Balkhash-Alakol Basin (Püngeler 1914; Titov et al. 2024).	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F819FFA9FF0D61464F01FAB6.taxon	diagnosis	Diagnosis. Members of the species-group are superficially reminiscent of the C. tertia species-group with their large and dark orbicular and reniform stigmata having thin and sometimes indistinct pale edges, but the male genital capsule differs clearly from two other species-groups in the genus by the broad, swollen uncus having lateral triangular appendages and bearing a cluster of setae, and the apically dilated harpe. In contrast to the male genitalia, the female genitalia of the C. mimetica / sacelli species-group display no fundamental differences from the C. murina species-group and are characterised only by the presence of a pair of elongate signa bursae. However, the similarly elongate signa are found in C. vera belonging to the C. murina species-group but in this species they are more asymmetrical than in all members of the C. mimetica / sacelli species-group, females of which are known. Notes. In their primary revision of the genus, Ronkay et al. (1995) illustrated only two female adults of C. sacelli and a drawing of the male genitalia, which is very similar to C. isotima and C. murina and therefore the conspecificity of this specimen to the female lectotype of C. sacelli is doubtful. This assumption is supported by the fact that subsequently, Gordeeva et al. (2023) illustrated a male adult, which is similar to the female lectotype of C. sacelli, but no photos of the male genitalia of the specimen were provided by the authors, which probably means that the male specimen illustrated remains undissected. The habitus of the lectotype of C. sacelli, as well as its female genitalia, are very similar to those of the members of the C. mimetica species-group, which have previously been known only from males, therefore C. sacelli is placed herein in this group. However, clarification of the relationships of this species requires the examination of the genitalia of the reliably identified male specimen.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F819FFACFF0D667249B3F89B.taxon	description	(Figs 15, 16, 26 – 28)	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F819FFACFF0D667249B3F89B.taxon	materials_examined	Type material examined. Photographs of the holotype (Fig. 16): “ Kazakhstan, Prov. Almaty, | Zailiisky [Transili] Alatau, 15 km S Issyk | 1750 – 1950 m, 77 ° 42 ['] E, 43 ° 13 ' N, | 3 – 4. X. 1994, leg. Gy. Fábián | & Gy. M. László ” / red framed label “ Holotypus [in red] | Dasypolia (Cteïpolia) [handwritten] | mimetica sp. n. [handwritten] | det. L. Ronkay [in red] / 94 [handwritten] ” / “ 4901 ♂ | gen. prep. No. | det. L. Ronkay ” (HNHM). Additional material examined. KAZAKHSTAN: 1 male, vic. of Vernyi [modern Almaty], Semirechye Reg., Kvochkin [leg.] [in Russian] / 24. X. 1907, gen. prep. No.: AV 7552 ♂ (ZISP); 1 male, 4. X. 2015, Almaty Region, Transili Alatau Mts, Big Almaty Lake, 43 ° 03 ' N 76 ° 58 ' E, 2523 m, S. V. Titov & M. Černila leg., gen. prep. No.: AV 1611 (CAV); 5 males, Almaty Reg., Transili Alatau Mts, 15 km S Issyk, Issyk Lake, 1710 m, 30. IX., 6 and 8. X. 2002, B. Benedek & T. Csővári leg. (BBC); 1 female, Almaty Region, Temerlik [Mts], 15 km NWN of Kegen settl., 43 ° 03 ' N 79 ° 24 ' E, [2600 m], 10. X. 2015, P. Gorbunov leg., gen. prep. No.: TB 2120 (BBC). Note. The species was described from four males from Southeast Kazakhstan and a single male from Kyrgyzstan (Ronkay et al. 1995). The female of the species is illustrated and diagnosed for the first time in the present paper.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F819FFACFF0D667249B3F89B.taxon	diagnosis	Diagnosis of female. The forewing length is 11.0 – 13.0 mm in males and 14.0 mm in the female. Cteipolia mimetica (Fig. 15) is externally reminiscent of C. sacelli (Fig. 14) but has a narrower forewing with markedly shorter subbasal dash and an elliptical circular stigma, which is long dash-shaped in the congener. Additionally, the hindwing of C. mimetica is darker than in the congener. Compared to C. sacelli (Fig. 25), the female genitalia of C. mimetica (Fig. 26) have longer apophyses anteriores, broader antrum with a broader sclerotised plate, and a somewhat shorter but broader corpus bursae.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
03BA8793F819FFACFF0D667249B3F89B.taxon	distribution	Distribution and bionomics. The species is known from northern and central parts of the Tien Shan Mts (Southeast Kazakhstan and Kyrgyzstan) (Ronkay et al. 1995). In the vicinities of the Big Almaty Lake in the Transili Alatau Range, C. mimetica was collected at an altitude of ca. 2500 m in the subalpine zone with scarse trees of Picea schrenkiana tianschanica (Rupr.) Bykov, thickets of bushes of Juniperus L., Salix L., Lonicera L., Cotoneaster Medik., etc., and subalpine meadow patches (Fig. 31). In the Temerlik Mts (SE Kazakhstan), the species was collected at an altitude of ca. 2600 m in the subalpine zone with a mosaic of Picea schrenkiana tianschanica sparse forest, subalpine meadows, and thickets of shrubs of the genera Rosa L., Spiraea L., and Lonicera L. (Fig. 32). Acknowledgements The authors are deeply grateful to Dr. George Fayvush (A. L. Takhtajan Institute of Botany NAS RA, Yerevan, Armenia) for the invaluable advices and consultations concerning habitats characterisation of the type locality of C. aloyani sp. n.; Anna A. Ivashchenko (Institute of Zoology, Almaty, Kazakhstan) for the data on the flora of the Sholak Mountains; László and Gabor Ronkay (Budapest, Hungary) for pictures of the types of C. vera and C. mimetica; and Sergey Gordeev and Tatyana Gordeeva (IGEB, Ulan-Ude, Russia) for valuable discussion and pictures of the Cteipolia amissa paratype. Sergey Melyakh (Yekaterinburg, Russia) is acknowledged for data from their research collections provided. Anton Volynkin sincerely thanks Wolfram Mey (MfN, Berlin, Germany) for his kind assistance during the work at MfN collection. Juozas Dûda and Aidas Saldaitis express their sincere thanks to Gayane Karagyan and Tigran Ghrejyan (Scientific Center of Zoology and Hydroecology NAS RA) for their multiple help in organizing the entomological expedition in Armenia. Juozas Dûda and Aidas Saldaitis’ special thanks go to Alena Kazakevich (Lida, Belarus) and Elena Kazyrytskaya (Darginiai, Lithuania) for their help in preparing the manuscript. Sergey Titov acknowledges Andrey M. Shapovalov (Institute of Zoology, Almaty, Kazakhstan) for organising the collecting trip in the Sholak Mountains; the Bayanaul State National Nature Park (Kazakhstan) (director Suyindik B. Akhmetzhanov, deputy director for science Yergalym U. Kayerdenov, senior researcher Dinara N. Kunshuakova, Sergey M. Reznichenko (former research fellow), the late Venera T. Tuleubekova (former research fellow), and the late Kuantay K. Abukenov (former head of the Department of Science, Ecological Education and Tourism )) for kind assistance in organising the research in the Bayanaul Mountains; Timur N. Smagulov, director of Institute of Archaeological Research, Margulan University (Pavlodar, Kazakhstan), for organising the collecting trip in the Saur and Monrak Mountains; Alexander Belousov and Yulia Belousova (Taldykorgan, Kazakhstan) for their kind hospitality and assistance in organising the collecting trips in the south-eastern Balkhash Basin and Dzhungar Alatau Mountains. Sergey Titov is also deeply grateful to the tragic deceased Sergey Ruzanov, director of the Cherkasai Tourist Center, for his warm welcome and kind assistance in organising the research in the Dzhungar Alatau Mountains. The study of Sergey Titov was funded by Science Committee of the Ministry of Education and Science of the Republic of Kazakhstan (Program No. BR 21882199, “ Inventory of wild animals of arid territories of the Balkhash-Alakol basin with an assessment of threats to their conservation and sustainable use. ”). The study of Alexey Matov was performed in the frames of the research project № 122031100272 - 3 “ Systematics, morphology, ecophysiology and evolution of insects ”, and the research was completed using equipment of the Core Facilities Centre “ Taxon ” at the Zoological Institute of the Russian Academy of Sciences. The study of Pavel Gorbunov was partly financed by the project FUWU- 2022 - 0007 “ Interaction of natural and anthropogenic factors in the formation of biological diversity of the Ural flora and fauna ” of the Institute of Plant and Animal Ecology of the Ural branch of the RAS. The study of Juozas Dûda, Aidas Saldaitis and Mark Kalashian was carried out in the framework of the Agreement of Scientific cooperation between Lithuanian Nature Research Centre and Scientific Center of Zoology and Hydroecology NAS RA. Anton Volynkin, Balázs Benedek, and Balázs Tóth, while performing this work, received no specific grant from any funding agencies.	en	Volynkin, Anton V., Dûda, Juozas, Saldaitis, Aidas, Benedek, Balázs, Titov, Sergey V., Kalashian, Mark, Matov, Alexey Yu, Gorbunov, Pavel Yu., Tóth, Balázs (2024): On the taxonomy of the genus Cteipolia Staudinger (Lepidoptera: Noctuidae: Noctuinae) with descriptions of two new species from Transcaucasia and Central Asia. Ecologica Montenegrina 77: 146-160, DOI: 10.37828/em.2024.77.15, URL: https://doi.org/10.37828/em.2024.77.15
