taxonID	type	description	language	source
03BD87FBE62C7176FF08FBE3B8F3C9B2.taxon	materials_examined	Type species: Phipicobia pygiptilae sp. nov., designated here.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62C7176FF08FBE3B8F3C9B2.taxon	diagnosis	Diagnosis. PHYSOGASTRIC FEMALE. Gnathosoma. Hypostomal apex rounded with shoulders. Lateral hypostomal teeth absent. Peritremes M-shaped. Borders between chambers poorly visible both in median and lateral branches. Movable cheliceral digits edentate on distal tip. Stylophore rounded posteriorly. Idiosoma. Propodonotal shield divided longitudinally, forming one central and two narrow lateral sclerites bearing bases of setae vi, ve, si and se. Six pairs of propodonotal setae present, arranged 2 - 1 - 1 - 1 - 1. Setae vi bases situated anterior to level of setae ve. Hysteronotal shield absent. Hysteronotal setae long, d 1 situated equidistant to e 2 and d 2. Alveoli of setae 1 a- 1 a and 3 a- 3 a separated. Pseudanal setae absent. Genital series with 2 pairs of setae. Aggenital series with 3 pairs of setae. Genital, pygidial and opisthosomal lobes absent. Legs. Legs I thicker than II, II thicker than III – IV. Apodemes Ia slightly divergent, in distal part parallel, provided with small outgrowth but not strict thornlike protuberances. Legs with full complement of solenidia. Physogastric form worm-shaped in outline.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62C7176FF08FBE3B8F3C9B2.taxon	description	MALE: Unknown.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62C7176FF08FBE3B8F3C9B2.taxon	etymology	Etymology. The generic name Phipicobia is a compilation of the Greek letter phi (φ) representing the golden section (sectio aurea), which value is approximately 1.61803 and the name of the genus Picobia. Differential diagnosis. Phipicobia gen. nov. is most similar to the genus Gunabopicobia Skoracki and Hromada, 2013. In females of both genera, the hypostomal apex has shoulders, the movable cheliceral digits are edentate on the distal tips, the stylophore is rounded posteriorly, the propodonotal shield is divided longitudinally into two narrow lateral shields bearing setal bases vi, ve, si and se, and the large central sclerite and the legs are with full set of solenidia. Females of Phipicobia gen. nov. differ from Gunabopicobia by the peritremes with poorly visible borders between chambers both in the median and lateral branches, setal alveoli 1 a- 1 a not coalesced, absence of the pseudanal setae, and by the presence of two pairs of the genital setae. In females of Gunabopicobia, chambers of the peritremes are clearly visible both in the medial and lateral branches, setal alveoli 1 a- 1 a are coalesced, pseudanal setae are represented by two pairs and the genital setae are absent. Habitat, host range and distribution. Monotypic genus with Phipicobia pygiptilae sp. nov. living inside the quills of body feathers of Pygiptila stellaris (Spix) (Passeriformes: Thamnophilidae) in Guyana.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7174FF08FF7BBCC2CA6E.taxon	description	(Figs. 1 – 5) PHYSOGASTRIC FEMALE. Total body length 930 (985 – 1000 in 9 paratypes). Gnathosoma. Hypostomal apex rounded with shoulders (Fig. 3). Infracapitulum apunctate. Peritremes M-shaped. Borders between chambers poorly visible. Each medial branch with about 8 – 9 chambers, each lateral with 5 – 7 chambers (Fig. 4). Stylophore 185 (180 – 195) long, punctate in anterior part. Idiosoma. Propodonotal shield divided into 3 sclerites, unpaired central and 2 narrow punctate lateral sclerites bearing bases of setae vi, ve, si and se. Setae vi bases situated anterior to ve bases. Length ratio of setae vi: ve: si 1: 1 – 1.2: 2.4 – 3.3. Bases of setae se situated anterior to c 1. Hysteronotal shield absent. Setae d 1 bases situated equidistant to e 2 and d 2. Setae d 2, d 1 and e 2 subequal in length. Pygidial and genital regions strongly striated. Lengths ratio of setae f 1: f 2 1: 2.6 – 3.4, h 1: h 2 1: 1.5, f 1: h 1 1: 4, f 2: h 2 1: 1.7. Length ratio of g 1: g 2 1: 2 – 4. Aggenital setae ag 1 and ag 2 situated almost at same transverse level, ag 3 posterior to ag 1 and ag 2. Length ratio of ag 1: ag 2: ag 3 1: 1.1 – 1.3: 1.5 – 1.7. Legs. All coxal fields strongly sclerotized, sparsely punctate. Setae 3 c 4.5 – 5 times longer than 3 b. Antaxial and paraxial members of claws pairs III – IV subequal in size and shape. Setae tc’ and tc’ ’ of legs III – IV subequal in length. Lengths of setae: vi 30 (25 – 45), ve 40 (35 – 45), si (95 – 120), c 2 220 (200 – 210), se 180 (160 – 205), c 1 210 (200 – 230), d 2 255 (215 – 150), d 1 245 (230 – 270), e 2 230 (205 – 255), f 1 60 (50 – 70), f 2 195 (170 – 195), h 1 (195), h 2 (290), ag 1 95 (100 – 130), ag 2 110 (105 – 130), ag 3 170 (155 – 170), g 1 45 (40 – 65), g 2 20 (10 – 20), tc’ III – IV (65 - 70), tc ” III – IV (70 – 75), l’RIII 20 (20 – 30), l’RIV 15 (15 – 20), 3 b 20 (20 – 25), 3 c 125 (105 – 120), 4 b 25 (20 – 30), 4 c 115 (100 – 130). MALE. Unknown.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7174FF08FF7BBCC2CA6E.taxon	etymology	Etymology: The specific epithet pygiptilae derives from the generic name of the host.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7174FF08FF7BBCC2CA6E.taxon	materials_examined	Type material. Female holotype (physogastric form) and 9 female paratypes (physogastric forms) from quills of body feathers of Pygiptila stellaris (Spix) (USNM 637200) (Passeriformes: Thamnophilidae), GUYANA: Upper Takutu - Upper Essequibo, Upper Rewa River, 10 August 2006, coll. B. K. Schmidt. Material deposition. Holotype female and 4 female paratypes are deposited in the USNM, 5 female paratypes in AMU.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7174FF08FB6AB9BFC874.taxon	discussion	Remark. Up to now, the genus Rafapicobia has included five species described from birds of the orders Passeriformes (Furnariidae, Mimidae, Muscicapidae) (4 species) and Gruiformes (Rallidae) (1) (Skoracki et al. 2014). Gradually increasing the number of known species in this genus makes the previous diagnosis (Skoracki 2011) is no longer sufficient. It mainly concerns such characters as the hypostomal apex shape (which is tapering), hysteronotal shields (present or absent), presence of small thorn-like protuberances on apodemes I (present or absent) and the presence of 3 pairs of aggenital setae and 1 pair of genital setae. Homology of setae ag 2 and g seems to be unclear. In females of R. zirnitra Skoracki and R. toxostoma Sikora, Fajfer andt Skoracki homology of setae ag 2 does not rise doubts, whilst in R. melzeri Skoracki, Unsoeld, Skorupski and Kavetska, R. dendrocaptesi Skoracki and Solarczyk and R. lepidocolaptesi Skoracki and Solarczyk, R. thamnophili sp. nov., and R. milenskyi sp. nov. arrangement of setae ag 2, their thickness and length suggest they are rather genital than aggenital setae.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7172FF08F990BF5EC80D.taxon	description	(Figs. 6 – 10) PHYSOGASTRIC FEMALE. Total body length 535 (555 – 565 in two paratypes). Gnathosoma. Hypostomal apex tapering, sharp-ended (Fig. 8). Infracapitulum punctate. Borders between chambers both in medial and lateral branches of peritremes invisible (Fig. 9). Stylophore 130 (120 – 125) long, apunctate. Idiosoma. Propodonotal shield divided into three sclerites, 2 narrow lateral, bearing bases of setae si and se, punctate along outer edge and unpaired central shield bearing bases of setae vi, ve and c 1, punctate on whole surface. Lateral sclerites fused with central part unilaterally or bilaterally in some specimens. Setae vi bases situated anterior to ve. Length ratio of vi: ve: si 1: 2 – 3: 2.6 - 3. Bases of setae se and c 1 situated at same transverse level. Two hysteronotal shields restricted to small punctate areas bearing bases of setae d 1. Setae d 1 bases situated closer to e 2 than d 2. Length ratio of setae d 2: d 1: e 2 0.9 – 1.5: 1: 1 – 1.5. Pygidial shield strongly sclerotized, densely punctate, bearing bases of setae f 1 and f 2. Length ratio of setae f 1: f 2 1: 3 – 5, h 1: h 2 1: 3.9 – 4.5, f 1: h 1 1: 2 – 3.5, f 2: h 2 1: 2.7 – 4.3. Agenital plates punctate, bearing bases of setae ag 1. Length ratio of setae ag 2: g 1: 1 – 2. Two pairs of subequal in length pseudanal setae present. Length ratio setae ag 1: ag 2: ag 3 10: 1: 10. Most of dorsal setae knobbed. Legs. All coxal fields strongly sclerotized, apunctate. Apodemes I with small thorn-like protuberances. Setae 3 c 1.6 – 2.8 times longer than 3 b. Antaxial and paraxial members of claws pairs III – IV subequal in size and shape. Setae tc’ and tc’ ’ of legs III – IV subequal in length. Lengths of setae: vi 35 (35 – 40), ve 95 (75 – 80), si 100 (95 – 110), c 2 155 (125 – 135), se 150 (120 – 140), c 1 185 (170 – 195), d 2 130 (130 – 145), d 1 145 (105 – 135), e 2 150 (130 – 165), f 1 20 (20 – 35), f 2 95 (70 – 85), h 1 70 (65), h 2 275 (295 – 300), ag 1 155 (140 – 145), ag 2 15 (15 – 20), ag 3 150 (150 – 155), g 20 (15 – 30), ps 1 and ps 2 10, l’RIII 20 (20 – 25), l’RIV 20 (10 – 20), 3 b 20 (30 – 40), 3 c 55 (65 – 75), 4 b 35 – 45, 4 c 80 (70). MALE. Unknown.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7172FF08F990BF5EC80D.taxon	etymology	Etymology. The specific epithet thamnophili derives from the generic name of the type host.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7172FF08F990BF5EC80D.taxon	materials_examined	Type material. Female holotype (physogastric form) and 4 female paratypes (2 physogastric and 2 nonphysogastric forms) from quills of body feathers of Thamnophilus insignis Salvin et Godman (USNM 639376) (Passeriformes: Thamnophilidae), GUYANA: Cuyuni- Mazaruni, Paruima, ca. 19 km N, at Mt. Holitipu, 25 February 2007, coll. B. K. Schmidt. Non-type material. From Thamnophilidae in GUYANA: 4 females (non-physogastric forms) from Myrmoborus leucophrys (Tschudi) (USNM 637174), Upper Takutu – Upper Essequibo, Lower Rewa River, 23 August 2006, coll. C. M. Milensky; 2 females (1 physogastric and 1 non-physogastric) from Myrmeciza ferruginea (St. Müller) (USNM 637194), Upper Takutu - Upper Essequibo, Upper Rewa River, 9 August 2006, coll. B. K. Schmidt; 12 females (10 physogastric and 2 non-physogastric) from Myrmotherula longipennis Pelzeln (USNM), Upper Takutu – Upper Essequibo, Lower Rewa River, 19 August 2006, coll. C. M. Milensky; 6 (5 physogastric and 1 non-physogastric) females from Hypocnemis cantator (Boddaert), Upper Takutu – Upper Essequibo, Lower Rewa River, 23 August 2006, coll. C. M. Milensky. Material deposition. Holotype female and 2 female paratypes are deposited in USNM, 2 female paratypes in AMU. Additional material, 2 females from Myrmoborus leucophrys, 1 female from Myrmeciza ferruginea, 6 females from Myrmotherula longipennis and 3 females from Hypocnemis cantator are deposited in USNM, 2 females from Myrmoborus leucophrys, 1 female from Myrmeciza ferruginea, 6 females from Myrmotherula longipennis and 3 females from Hypocnemis cantator are deposited in AMU.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE62E7172FF08F990BF5EC80D.taxon	diagnosis	Differential diagnosis. This new species is most similar morphologically to Rafapicobia dendrocolaptesi Skoracki and Solarczyk, 2012 described from Dendrocolaptes platyrostris Spix from Paraguay and D. picumnus Lichtenstein from Argentina (Skoracki and Solarczyk 2012). In females of both species, the hysteronotal shields are present and restricted to small punctate areas bearing bases of setae d 1, setae ag 2 are short (10 – 15 and 15 – 20 in R. dendrocolaptesi and R. thamnophili sp. nov., respectively), most of dorsal setae are beaded, all shields are punctate and antaxial and paraxial members of claws pairs III – IV are subequal. Females of R. thamnophili sp. nov. differ from R. dendrocolaptesi by the propodonotal shield generally divided into three parts, two narrow lateral sclerites bearing bases of setae si and se, punctate along the outer edge and the central shield bearing bases of setae vi, ve and c 1 (in some specimens lateral sclerites are fused with the central part unilaterally or bilaterally) and lengths of setae ve 75 – 95, d 2 130 – 145 and h 1 65 – 70. In females of R. dendrocolaptesi the propodonotal shield is entire and lengths of setae are ve 45 – 60, d 2 165 – 190 and h 1 10.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE6287170FF08F974BD4ACDCE.taxon	description	(Figs. 11 – 15) NON-PHYSOGASTRIC FEMALE. Total body length 430 – 470 in two paratypes. Gnathosoma. Hypostomal apex tapering (Fig. 13). Infracapitulum apunctate. Borders between chambers in medial and lateral branches invisible (Fig. 14). Stylophore 120 (115) long. Idiosoma. Propodonotal shield shirt-shaped, punctate on whole surface, bearing bases of setae vi, ve, si and se. Setae strongly knobbed. Setae vi situated anterior to ve bases. Length ratio of setae vi: ve: si 1: 2 – 2.3: 2.5 – 2.8. Hysteronotal shield restricted to small punctate areas bearing bases of setae d 1. Length ratio of setae f 1: f 2 1: 1, h 1: h 2 1: 6.7, f 1: h 1 1: 2 – 3.5, f 2: h 2 1: 20 – 23. Length ratio of g: ag 2 about 1: 1 – 1.5. Two pairs of subequal in length pseudanal setae present. Length ratio of ag 1: ag 2: ag 3 11.3 – 17: 1: 8.7 – 13. Legs. All coxal fields strongly sclerotized, I – II apunctate, III – IV sparsely punctate. Setae 3 c 2.8 – 3 times longer than 3 b. Antaxial and paraxial members of claws pairs III – IV subequal in size and shape. Setae tc ” of legs III – IV 1.8 – 2.2 times longer than tc’. Lengths of setae: vi 20 (20), ve 40 (30 – 45), si 45 (50 – 55), c 2 115 (95 – 130), se 90 (90), c 1 170 (175 – 180), d 2 125 (115 – 130), d 1 105 (135), e 2 140 (140), f 1 15 (10 – 20), f 2 15 (10 – 15), h 1 30 (35), h 2 235 – 295, ag 1 170 (150 – 165), ag 2 15 (10 – 15), ag 3 155 (130), g 10, ps 1, 2 10 (7), tc’ 30 (20), tc ” 50 (45), l’RIII 20 (20), l’RIV (20), 3 b 20 (25), 3 c 60 (70), 4 b 25, 4 c 55 (80). PHYSOGASTRIC FEMALE: as non-physogastric form, body bulb-shaped in outline. MALE. Unknown.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE6287170FF08F974BD4ACDCE.taxon	etymology	Etymology. This species is named in honor of the collector of the host specimen, C. M. Milensky.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE6287170FF08F974BD4ACDCE.taxon	materials_examined	Type material. Female holotype (non-physogastric form) and 2 female paratypes (1 non-physogatric females and 1 physogastric forms) from quill of body feathers of Conopophaga aurita (Gmelin) (USNM 637100) (Passeriformes: Conopophagidae), GUYANA: Upper Takutu - Upper Essequibo, Upper Rewa River, August 2006, coll. C. M. Milensky. Type deposition. Holotype female is deposited in USNM, 2 female paratypes (non-physogastric and physogastric forms) in AMU.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
03BD87FBE6287170FF08F974BD4ACDCE.taxon	diagnosis	Differential diagnosis. This new species is most similar morphologically to Rafapicobia dendrocolaptesi Skoracki and Solarczyk, 2012. In females of both species, the propodonotal shield is shirt-shaped, bearing bases of setae vi, ve, si and se and punctate on whole surface, the hysteronotal shields are present and restricted to the small punctate areas bearing bases of setae d 1 and setae ag 2 are short (10 – 15). Females of R. milenskyi sp. nov. differ from R. dendrocolaptesi by the length of setae si 45 – 55, c 2 95 – 130, se 90, d 2 115 – 130, f 2 10 – 15 and h 1 30 – 35. In females of R. dendrocaptesi the lengths of these setae are si 75 – 80, c 2 155 – 190, se 130 – 140, d 2 165 – 190, f 2 100 – 135 and h 1 10.	en	Glowska, Eliza, Schmidt, Brian K. (2014): New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) parasitizing antbirds and gnateaters (Passeriformes: Thamnophilidae, Conopophagidae) in Guyana. Zootaxa 3861 (2): 193-200, DOI: 10.11646/zootaxa.3861.2.7
