identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BF4239FFF3FFCFEAF8C2659C63FE79.text	03BF4239FFF3FFCFEAF8C2659C63FE79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides Sars 1835	<div><p>Genus Terebellides Sars, 1835 emended by Schüller &amp; Hutchings, 2013</p><p>Type species. Terebellides stroemii Sars, 1835, redescribed in Parapar &amp; Hutchings (2015)</p></div>	https://treatment.plazi.org/id/03BF4239FFF3FFCFEAF8C2659C63FE79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFF3FFC5EAF8C10C9B87FD84.text	03BF4239FFF3FFC5EAF8C10C9B87FD84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides africana Augener 1918	<div><p>Terebellides africana Augener, 1918 stat. prom.</p><p>Figures 2A, 3−7, 11A, 37; Tables 1, 2</p><p>Terebellides Stroemi var. africana Augener, 1918 . Augener (1918): 555 –560, Plate 7, fig. 245, text fig. 96.</p><p>Material examined. Type material. Two syntypes (ZMH V1709, ZMH V1712) (Table 1); lectotype: V1709 (female with intra-coelomic oocytes); paralectotype: V1712 (see Discussion) . Non-type material. Seven specimens (NHMD-231436, NHMD-231441, NHMD-231453, NHMD-231455, NHMD-231458); Atlantide and Galathea expeditions (Kirkegaard, 1959) (Table 1).</p><p>Diagnosis. Body large (20–30 mm in length). Branchial anterior lobe present; posterior ventral lobes much smaller than dorsal ones, without terminal filament and emerging after fusion line of dorsal ones. TC1 notopodia and notochaetae as long as following. Thoracic neurochaetae with rostrum / capitium length about 3/1, and capitium with a first row of 3(4) medium-sized teeth, followed by progressively smaller teeth.</p><p>Description of lectotype</p><p>Measurements and general body features. Complete specimen, 32 mm long and 3.0 mm wide (Fig. 2A, 3 A−C), body tapering posteriorly, with segments increasingly shorter and crowded towards pygidium. Prostomium compact, large tentacular membrane with typical buccal tentacles with expanded tips. SGI forming an expanded lower lip below tentacular membrane (Fig. 3 E−F). Lateral lappets not conspicuous on SGIII–VII (TC1–5), larger in SGIII–V, with low dorsal projections in TC1−3 (Fig. 5A); TC2 dorsally elevated in relation to TC1 and TC3 (Fig. 2A, 5A). Oval-shaped glandular region in CH 3 not observed.</p><p>Branchiae. Branchiae arising as single structure from SGIII, with a single, short, stalked, mid-dorsal branch, typically comma-shaped and reaching TC3 (Fig. 2A, 3 A−C). A short anterior lobe (see below) and two pairs of pos- terior lobes: one pair dorsal (upper), with lobes 1+2 fused along most of their length, one pair ventral (lower), with lobes 3+4 much shorter than dorsal ones, not fused together, near distal end and after a long fusion line of dorsal ones. Anterior projection of dorsal lobes (lobe 5) short, about ¼ of dorsal lobes’ length (Fig. 2A, 3 C−D, 4B). Pointed projection of posterior region of both upper and lower lobes short, a bit longer in lower ones but never forming a long terminal filament. Branchial lamellae with well-developed ciliary rows and ciliated papillae on lamellar outer edge (Fig. 5B).</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX), that of TC1 about as long as following ones (Fig. 3E, 5A), with dorsal projection; notochaetae as simple capillaries as long as following ones. TC2 and TC3 with dorsal projection similar to TC1 (Fig. 5A, C). Notopodia of TC2 shifted dorsally in some specimens (Fig. 5A). Neuropodia as sessile pinnules from TC6 (SGVIII) to pygidium, with uncini in single rows starting from TC7 (SGIX) throughout. First neuropodia at TC6, with five sharply bent, acute tipped, geniculate chaetae in left torus (Fig. 6 A−B); right side torus lost, presumably removed by H. Augener (Fig. 3C). From TC7, neuropodia with up to 18–22 uncini per torus in one row, with long shafted denticulate hooks, rostrum about three times longer than capitium, with three large teeth above main fang, and surmounted by an upper crest of several smaller denticles (Fig. 7 A−C), dorsalmost hooks in anteriormost uncinigers showing a very short rostrum and a less angled capitium in relation to chaetal main axis, resembling aciculae (Fig. 6 C−D).</p><p>Abdomen. Abdomen as long as thorax. Thirty-two to thirty-four pairs of neuropodia as erect pinnules with about 25−30 uncini per torus; uncini with about five teeth above main fang, surmounted by a upper crest of 2 teeth and a variable number of smaller denticles (?) (see Fig. 2E in Augener, 1918).</p><p>Other body features. Single, well-developed nephridial papilla behind TC1 notopodia and nephridial openings dorsal to TC4−5 notopodium (Fig. 5A, C). Pygidium as blunt, funnel-like depression.</p><p>Methyl green staining pattern. Striped through the entire thorax; pattern 7 of Schüller &amp; Hutchings (2010) (Fig.</p><p>37).</p><p>Variability. Type specimens differing in size (paralectotype: 22.0 mm long, 2.0 mm wide), but showing no significant differences in relevant morphological characters. Branchiae clearly smaller in lectotype than in non-type specimens identified by Kirkegaard (1959), likely due to a longer preservation in alcohol.</p><p>Type locality. Equatorial Guinea; 0−30 m depth ( Table 1). The location corresponds to the lectotype (V-1709), while Augener (1918) simply refers to “West Africa” for the paralectotype (V-1712) .</p><p>Distribution and bathymetry. The revision of material from the Atlantide expedition, originally identified as T. stroemii by Kirkegaard (1959), expands the distribution of T. africana stat. prom. to the Gold Coast and Nigeria, from 17 to 66 m depth (Fig. 11A). The specimen NHMD-231441, collected during the Galathea expedition (75 m depth, off Angola), mostly agrees with the diagnosis of T. africana stat. prom., but its very poor condition prevented fully confirming its identity. The specimen NHMD-231455 (Fig. 4C) shows two filiform structures protruding dorsally from TC3 that might correspond to a parasitic copepod.</p><p>Nomenclatural note. Following Alonso-Zarazaga (2007), the term stat. prom. (Lat. status promotus; Eng. promoted status) is used here to qualify T. africana instead of the classical stat. nov. (Lat. status novus; Eng. new status) for referring to the rising of a subspecies (meaning of “variety” following the International Code for Zoological Nomenclature-ICZN, article 45.6.4.) to the species rank. This author also proposed stat. dem. (Lat. status demotes; Eng. demoted status) instead of stat. nov. (i.e., upgrading an infrasubspecific entity to specific or subspecific status, ICZN, article 16A) .</p><p>Remarks. Even though type specimens were not examined under SEM to avoid damage, this species is clearly distinguished from the other described here due to its large body size, a TC1 similar in size to following thoracic notopodia, and branchiae with very short ventral lobes emerging close to the distal end of the dorsal ones.</p><p>Terebellides africana stat. prom. resembles T. stroemii as redescribed by Parapar &amp; Hutchings (2015) in size and general body shape, typical comma-shaped branchiae, and appearance and size of TC1, and mostly differs in branchial features: 1) ventral and anterior lobes clearly smaller, the former showing a different position, and 2) lobes bearing ciliated papillae (Fig. 5 A−B) (absent in T. stroemii, Parapar &amp; Hutchings, 2015: Fig. 6 E−F).</p><p>Terebellides africana stat. prom. resembles T. stroemii sensu Caullery (1944) from Indonesia (St. 271 of Siboga Expedition) in branchiae size and shape, which reach TC3 (Caullery, 1944: Fig. 147A). However, their ventral posterior lobes are much different, and resemble those of a specimen from St. 311 (Caullery, 1944: Fig. 148A) that has also larger anterior branchial lobes. Therefore, we suspect that each of them might correspond to undescribed species.</p><p>Rullier (1964a) reported T. stroemii from Cameroon, near the type locality of T. africana stat. prom. (Equatorial Guinea), which were not assigned to the variety africana due to the shape of TC6 geniculate chaetae (not illustrated in his paper). Even though we were not able to examine this material, deposited in the Muséum national d’Histoire naturelle (Paris), it seems it does not correspond to any of the species described here. Two species described here, T. kirkegaardi sp. nov. and T. longiseta sp. nov., are geographically close to T. africana stat. prom., but they clearly differ in the shape of TC6 geniculate chaetae.</p><p>Terebellides persiae Parapar, Moreira, Gil &amp; Martin, 2016 overall resembles T. africana stat. prom., but differs in having much larger branchiae (particularly the anterior lobe) and a characteristic thoracic dorsal hump (cfr. Fig. 2B &amp; 3A in Parapar et al., 2016b) absent in T. africana stat. prom. Terebellides parvus Solís-Weiss et al., 1991 (Solís-Weiss et al., 1991: Fig. 2a) has also similar branchiae, but differs by having fewer abdominal chaetigers (25 vs. 32−34 in T. africana stat. prom.).</p><p>The relative size and position of the postero-ventral branchial lobes in relation to the dorsal ones in T. africana stat. prom. resembles those of T. mediterranea Parapar, Mikac &amp; Fiege, 2013, T. guangdongensis Zhang &amp; Hutchings, 2018 and T. yangi Zhang &amp; Hutchings, 2018 . However, T. mediterranea has much larger ventral lobes, a more developed branchial stem, and a larger TC1 with longer notochaetae; T. guangdongensis shows smaller anterior branchial lobes and a very small TC1, and T. yangi clearly differs in having much-developed anterior lobes, similar to those of T. longiseta sp. nov. (see below).</p><p>Terebellides africana stat. prom. resembles T. augeneri sp. nov., but differs in body size (22−32 mm vs. 7−19 mm in T. augeneri sp. nov.) and in the position of the branchial ventral lobes (arising from the base of the dorsal ones in T. augeneri sp. nov.).</p><p>Two more species here described occur in shallow waters: T. congolana sp. nov. and T. kirkegaardi sp. nov. The former resembles T. africana stat. prom. in body size and shape and in the position of branchial ventral lobes but differs in having much longer branchial dorsal lobes that reach TC6 and a TC1 notopodium proportionally larger compared to thorax size. On the other hand, T. kirkegaardi sp. nov. differs from T. africana stat. prom. in having a distinctly smaller body and branchiae with a very long branchial stem, an anterior lobe about half the length of posterior ones, and ventral lobes originating from the base of dorsal ones.</p></div>	https://treatment.plazi.org/id/03BF4239FFF3FFC5EAF8C10C9B87FD84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFF9FFD8EAF8C40B999FFC32.text	03BF4239FFF9FFD8EAF8C40B999FFC32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides augeneri Parapar & Martin & Moreira 2020	<div><p>Terebellides augeneri sp. nov.</p><p>Figures 2B, 8−10, 11A, 37; Tables 1, 2</p><p>urn:lsid:zoobank.org:act: 147575A6-3605-4327-A3AD-F5F1C80A3D0D</p><p>Material examined. Type material. Six specimens: holotype (MNCN 16.01 /18579); five paratypes (MNCN 16.01 /18577, 16.01/18580, 16.01/1858 and NHMD-636919, Table 1) .</p><p>Diagnosis. Small/medium-sized species (7–19 mm in length). Branchial anterior lobe absent; posterior ventral lobes thinner than dorsal ones, with terminal projections, directly emerging from branchial stem. TC1 notopodia and notochaetae as long as following. Thoracic neurochaetae with rostrum / capitium length about 1/1, and capitium with a first row of 5–6 medium sized teeth followed by progressively smaller teeth.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, 19 mm long and 1.5 mm wide; female with oocytes in body cavity (Fig. 2B). Body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; large tentacular membrane surrounding mouth, with typical buccal tentacles and expanded tips (Fig. 8C, 9A). SGI as an expanded structure below tentacular membrane. Lateral lappets on SGIII– VII (TC1–5), larger in SGIV–VI (TC2–4) (Fig. 2B, 8 C–D, 9A).</p><p>Branchiae. Branchiae arising as single structure from SGIII, with a single stalked mid-dorsal branch, one pair of dorsal (upper) not-fused lobes, and a pair of shorter ventral (lower) lobes, neither fused to each other, nor to dorsal ones, with short terminal projections (Fig. 9C). Anterior projection of dorsal lobes (lobe 5) absent (Fig. 2B). Both sides of branchial lamellae having several parallel rows of cilia and low ciliated tufts on lamellar outer edge (Fig. 9 B−C).</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX), those of TC1 about as long as following ones. All notochae- tae as simple capillaries. Neuropodia as sessile pinnules from TC6 (SGVIII) to body end, with uncini in single rows, from TC7 (SGIX) throughout. First neuropodia on TC6, with 4–5 sharply bent, acute tipped, geniculate chaetae with minute teeth forming a well-defined capitium (Fig. 10 C−D). From TC7, neuropodia with up to 5–10 uncini per torus in one row, with long shafted denticulate hooks, rostrum about same length as capitium, with 5−6 medium-sized teeth above main fang, and surmounted by two rows of progressively shorter teeth (Fig. 10 A−B).</p><p>Abdomen. Thirty-one pairs of neuropodia, as erect pinnules with about 20−25 uncini per torus, with four teeth above main fang surmounted by an upper crest of similar sized teeth (Fig. 10 E−F) and several smaller teeth above.</p><p>Other body features. Nephridial (?) papilla behind TC1; nephridial openings dorsal to any anterior thoracic notopodium not seen. Pygidium blunt, as a funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 4 solid; CH 5 to CH 11 striped; near pattern 1 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variability. Body ranging from 7.0 to 19.0 mm in length.</p><p>Type locality. Off Angola; 73 m depth (Table 1) .</p><p>Distribution and bathymetry. Off Angola; 72−83 m depth (Fig. 11A; Table 1).</p><p>Etymology. This species is named after the German scientist and prolific marine zoologist Hermann Augener (1872–1938), as an homage to his numerous contributions to polychaete taxonomy.</p><p>Remarks. The branchiae of T. augeneri sp. nov. resemble those of T. hutchingsae Parapar, Moreira &amp; Martin, 2016 in bearing ventral lobes much less developed than the dorsal ones and arising from the base of the latter, but differ in having wider and longer lobes and in lacking a longer terminal projection. Both species also differ in the size of the ciliature in branchial lamellae outer edge (much lower in T. augeneri sp. nov.), in the shape of thoracic uncini (with teeth surmounting the rostrum much bigger in T. augeneri sp. nov.) and in bearing geniculate chaetae in CH 5 and CH 6 (only in CH 6 in T. augeneri sp. nov.).</p><p>Terebellides ceneresi Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019 has in common with T. augeneri sp. nov. the general branchial shape, with very similar branchial lobes and ciliary tufts on both sides of the outer edge of lamellae and no papillar projections pointing over their edge. Nevertheless, both species differ in the thoracic uncini, both in the shape of main fang (straight in T. augeneri sp. nov.; terminally bent in T. ceneresi) and the denticulation of capitium (5–6 teeth in first row in T. augeneri sp. nov.; 3–4 in T. ceneresi).</p><p>Terebellides shetlandica Parapar, Moreira &amp; O’Reilly, 2016 also bears ventral branchial lobes not fused to dorsal ones and arising directly from the branchial stem. However, it is much smaller than T. augeneri sp. nov. (up to 15 mm; Parapar et al., 2016c), the four posterior lobes are of similar size, and the branchial lamellae do not bear ciliated papillae.</p><p>In African Atlantic waters T. augeneri sp. nov. most closely resembles T. ramili sp. nov. (see below). However, it inhabits much deeper bottoms and is unique among African species because in the thoracic uncini all teeth surmounting the rostrum are very small and of the same size; in all other species, the first-row teeth are always larger than those of posterior rows.</p></div>	https://treatment.plazi.org/id/03BF4239FFF9FFD8EAF8C40B999FFC32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFE4FFDFEAF8C0DB9D1AFD48.text	03BF4239FFE4FFDFEAF8C0DB9D1AFD48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides congolana Parapar & Martin & Moreira 2020	<div><p>Terebellides congolana sp. nov.</p><p>Figures 11A, 12A, 13−15, 37; Tables 1, 2 urn:lsid:zoobank.org:act: F56164BC-1B78-4ECE-ACA5-543E5D75EB90</p><p>Material examined. Type material. Four specimens: holotype (MNCN 16.01 /18582); three paratypes (MNCN 16.01 /18581, 16.01/18583 and 16.01/18584, Table 1) .</p><p>Diagnosis. Body large (31–33 mm in length), with branchial anterior lobe, posterior ventral lobes much smaller than dorsal ones, with short terminal filament, and emerging after fusion line of dorsal ones. TC1 notopodia and notochaetae much longer than following ones. Thoracic neurochaetae with rostrum / capitium length about 1/1, and capitium with a first row of 4–5 big teeth followed by a second row of much smaller, progressively smaller teeth.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, 31.0 mm long and 3.0 mm wide (Fig. 12A); female with oocytes in body cavity. Body tapering posteriorly, with segments increasingly shorter and crowded towards pygidium. Prostomium compact; large tentacular membrane surrounding mouth with typical buccal tentacles with expanded tips (Fig. 13A, E). SGI as an expanded structure below tentacular membrane (Fig. 13 A–B). Lateral lappets on SGIII–VIII (TC1–6), larger in SGIII–VII (TC1−5) (Fig. 13 A−E). Low oval-shaped glandular region in CH 3 in some specimens (Fig. 13B), but not evident.</p><p>Branchiae. Branchiae arising as single structure from SGIII, with a single short-stalked mid-dorsal branch (Fig. 12A, 13 A−B, E), one pair of long dorsal (upper) lobes 50% fused, reaching TC6−7 (Fig. 13C) and one pair of ven- tral (lower) lobes about 1/5 as long as dorsal ones), not fused together and arising from mid-length of dorsal lobes fusion line (Fig. 13 A−B). Anterior projection of dorsal lobes (lobe 5) large, about 1/4 length of posterior dorsal lobes (Fig. 13E). Pointed projection of posterior region only in lower lobes, as a short and thick terminal filament (Fig. 13A). Both sides of branchial lamellae with several parallel bent rows of cilia and ciliated papillae on outer edge (Fig. 14 C−D).</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX), with dorsal ciliated papillae (Fig. 14 E−F). TC1 much more developed and with longer notochaetae than subsequent ones (Fig. 13B, E). All notochaetae as simple capillaries. Neuropodia as sessile pinnules from TC6 (SGVIII) to body end, with uncini in single rows starting from TC7 (SGIX) throughout. First neuropodia (TC6) with four sharply bent, acute tipped, geniculate chaetae (Fig. 15B) having minute teeth forming an ill-defined capitium. From TC7, neuropodia with 8−14 uncini per torus in one row, with long-shafted denticulate hooks, rostrum about same length as capitium, with 4−5 big teeth above main fang surmounted by several much smaller denticles (Fig. 15 C−E).</p><p>Abdomen. About 40% of body length, with 31 pairs of neuropodia as erect pinnules (Fig. 15G); about 25–30 uncini per torus having four teeth above main fang surmounted by 3–4 teeth in irregular middle row and an upper crest of several smaller teeth (Fig. 15H).</p><p>Other body features. One well-developed nephridial papilla behind each TC1 notopodium (Fig. 14E), and two pairs of nephridial openings in TC4 and TC5 (Fig. 14F). Pygidium blunt, funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 9 solid; CH 10 to CH 18 striped; being CH 10 to CH 11 much more marked than following; J-shaped glandular region marked lateral to CH 3; near pattern 5 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variability. Body between 31 and 33 mm in length. Type locality. Off Congo (Table 1). Distribution and bathymetry. Off Congo; sandy mud; 47−167 m depth (Fig. 11A; Table 1). Etymology. The name of the species refers to the type location, the Republic of Congo. Remarks. Terebellides anguicomus Müller, 1858 sensu Solís-Weiss et al. (1991) resembles T. congolana sp. nov. in having large sized-branchiae reaching TC6 but differs in having a large transverse dorsal hump on TC5 and 17 thoracic chaetigers (18 in T. congolana sp. nov. and most species of the genus, see key below).</p><p>Terebellides congolana sp. nov. also differs from other African species in having posterior ventral branchial lobes that are much smaller and located distally, but the following ones also have this character: T. rigel Schüller &amp; Hutchings, 2013; T. mediterranea Parapar, Mikac &amp; Fiege, 2013; T. akares Hutchings, Nogueira &amp; Carrerette, 2015, T. persiae and T. hutchingsae . However, T. rigel is a deep-water species (&gt; 2000 m) from the Weddell Sea and does not bear long TC1 notopodia and notochaetae; T. akares was described from coral reefs (Lizard Island, NE Australia) and has a distinctly short abdomen; T. persiae is a shallow-water species from the Iranian coast, that has a much more developed anterior branchial lobe and a distinct dorsal hump at TC6; T. hutchingsae is also a shallow subtidal species from the Gulf of Thailand, that bears geniculate chaetae in two chaetigers, TC5 and TC6, instead of only one in TC6. The closest species to T. congolana sp. nov. is T. mediterranea, a shallow-water species (31 m depth) from the Adriatic Sea; it also bears branchiae with a similar shape and very small ventral posterior lobes arising near the distal end of the branchiae, and TC1 notopodia and notochaetae longer than following, but it differs from the new species in having a shorter body (21 mm in length vs. 31 mm in T. congolana sp. nov.) and a much larger tentacular membrane.</p><p>Terebellides europaea Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019, from the French coast of the Bay of Biscay, also shares with T. congolana sp. nov. the long notopodia and notochaetae of TC1, but it differs from the new species in having papillar projections over the edge on anterior branchial lamellae, much bigger ventral branchial lobes and attached to the branchial stem, and in lacking ciliated papillae in branchial lamellae.</p></div>	https://treatment.plazi.org/id/03BF4239FFE4FFDFEAF8C0DB9D1AFD48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFE3FFD2EAF8C0459C68F84D.text	03BF4239FFE3FFD2EAF8C0459C68F84D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides fauveli Parapar & Martin & Moreira 2020	<div><p>Terebellides fauveli sp. nov.</p><p>Figures 11B, 12B, 16–20, 37; Tables 1, 2 urn:lsid:zoobank.org:act: F388A555-F6F0-48A8-93C5-C244DADDA76B</p><p>Material examined. Type material. Thirty-six specimens: holotype (NHMD-636920); thirty-five paratypes (NHMD-231435, NHMD-231442–231445, NHMD-231447, NHMD-231450–231452, NHMD-231461, NHMD- 231463, NHMD-636921 to NHMD-636922). Non-type material. Seven specimens (MNCN 16.01 /18585 to 16.01/18590, Table 1) .</p><p>Diagnosis. Body of medium/large size (15–35 mm in length). Branchiae comma-shaped, with anterior lobe and posterior ventral lobes thinner than dorsal ones, directly emerging from branchial stem and with well-developed thin terminal filament. TC1 notopodia and notochaetae longer than following thoracic ones. Thoracic neurochaetae with rostrum / capitium length about 4/1, and capitium with a first row of 2–3 very large teeth followed by three rows of also large, but smaller teeth.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, 30.0 mm long and 3.0 mm wide (Fig. 12B). Body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; large tentacular membrane surrounding mouth, with some typical buccal tentacles with expanded tips. SGI as expanded structure, below tentacular membrane (Fig. 16 B−C). Low lateral lappets on SGIII–VII (TC1–5) much larger in SGIII–V (TC1−3) and latero-dorsally white in TC3−4 (Fig. 16 A−B).</p><p>Branchiae. Branchiae comma-shaped, arising as single structure from SGIII, consisting of a single short stalked mid-dorsal branch with one pair of long dorsal (upper) unfused lobes, reaching TC6−7 (Fig. 12B, 16 B−C), and one pair of ventral (lower) smaller and thinner lobes (about half length of dorsal), not fused together and emerging after a short fusion line of dorsal lobes (Fig. 19 A−B). Anterior projection of dorsal lobes (lobe 5) about 1/7 length of posterior dorsal lobes (Fig. 16A, C). Pointed projection of posterior region only in lower lobes, as a thin but welldeveloped terminal filament (Fig. 16D, 17 B−C, 19A−B). Both sides of branchial lamellae with rows of cilia (Fig. 17 B−C, 19C); ciliated tufts or papillae not seen.</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX), much more developed and with longer notochaetae in TC1 than in following ones (Fig. 16 C−D, 17A−B, 19D). All notochaetae as simple capillaries. Neuropodia as sessile pin- nules, from TC6 (SGVIII) to pygidium, with around 15 uncini in single rows starting from TC7 (SGIX) throughout. First neuropodia (TC6) with four sharply bent, acute tipped, geniculate chaetae (Fig. 18A) having small minute teeth forming an ill-defined capitium (Fig. 19 E−F). From TC7, neuropodia with 8–14 uncini per torus in one row, with long shafted denticulate hooks, a very long rostrum about four times longer than capitium, with 2(3) very big teeth above main fang, surmounted by two rows of 3−4 also big, but smaller teeth and an upper crest of several smaller denticles (Fig. 18 B−C).</p><p>Abdomen. About 40% of body length, with 29 pairs of neuropodia as erect pinnules, with around 30 uncini per torus having three teeth above main fang surmounted by an upper crest of several minute teeth (Fig. 18D).</p><p>Other body features. One nephridial papilla behind each TC1 notopodium and nephridial openings in TC4 and TC5 notopodia. Pygidium blunt, funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 9 solid; CH 10 to CH 18 striped; being CH 10 to CH 11 much more marked than following; J-shaped glandular region marked lateral to CH 3; near pattern 5 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variability. Body between 15 and 35 mm in length.</p><p>Type locality. Ivory Coast; 80–90 m depth (Table 1) .</p><p>Distribution and bathymetry. Widely distributed along the West African coasts, from Liberia to South Angola; 42−200 m depth (Fig. 11B; Table 1).</p><p>Etymology. The species is named after the French zoologist Pierre Fauvel (1866–1958) for his many contributions to the study of the Polychaeta across world oceans, especially in Europe and Africa.</p><p>Remarks. Three Terebellides species have been previously described with TC1 provided with large notopodia and long notochaetae: T. kobei Hessle, 1917 (Japan), T. californica Williams, 1984 (California), T. mediterranea, and two in this work: T. congolana sp. nov. and T. fauveli sp. nov.</p><p>Terebellides fauveli sp. nov. has shorter TC1 notopodia and notochaetae than T. kobei, lacks a conspicuous triangular dorsal projection in TC3 notopodia (present in T. kobei), has a white lateral colouration in anterior chaetigers (absent in T. kobei) and has a different capitium denticulation, being all teeth small and of similar size in T. kobei, as illustrated by Hessle (1917: textfig. 32, p. 140).</p><p>Terebellides californica was described from shelf and slope depths in the Pacific Ocean (Oregon to Western Mexico) and is characterized by the author only by the “ very well developed first chaetiger with greatly prolonged fine notosetae ” (Williams, 1984: p. 128), which is clearly longer than the one in T. fauveli sp. nov. and lacks any special development in TC3 and body colour pattern. This species was later redescribed by Hilbig (2000) as having a “ trilobed structure of the peristomium ” as potential new diagnostic character, which is not present in T. fauveli sp. nov. However, in our opinion, the relevance of this character should be considered with caution as its presence in T. californica has not been sufficiently assessed.</p><p>Terebellides mediterranea most closely resembles T. fauveli sp. nov. in branchial shape. However, T. fauveli sp. nov. bears larger branchiae that reach TC6 (instead of TC3−4) and has posterior branchial lobes with terminal filament. Moreover, both species differ in number and size of the denticulation of thoracic uncini, with 3−4 mediumsized teeth above main fang and surmounted by a row of 6−7 slightly shorter tooth in T. mediterranea (see Parapar et al., 2013: Fig. 7B).</p><p>Terebellides anguicomus and T. lanai Solís-Weiss et al., 1991 resemble T. fauveli sp. nov. in the shape (commalike) and size of the branchiae, also reaching TC6. However, T. lanai bears branchial lobes fused for most of their length, and T. anguicomus has a large transverse dorsal hump on TC5, 17 thoracic chaetigers (instead of 18) and a shorter TC1.</p><p>While some of the French species recently described by Lavesque et al. (2019), such as T. lilasae Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019, have similar general branchial shape as in T. fauveli sp. nov., the capitium denticulation of the thoracic uncini is very different; in fact, the shape of the thoracic uncini in T. fauveli sp. nov. has not been recorded so far in any other species of the genus.</p><p>Among the West African species, T. fauveli sp. nov. apparently presents the widest bathymetric and geographic distribution. Non-type specimens collected off Congo (N’Kossa gas field, Table 1) show subtle differences in the shape and size of the terminal filament of lower branchial lobes, size of TC1 and thoracic uncini teeth (cfr. Fig. 17−18 vs. 19−20). Therefore, we cannot discard they may represent a different, close taxon. More extensive sam- pling across West Africa and further morphological and molecular analyses may help to test the present wide distribution of T. fauveli sp. nov. and the possible existence of several cryptic species.</p></div>	https://treatment.plazi.org/id/03BF4239FFE3FFD2EAF8C0459C68F84D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFE9FFE9EAF8C41B99CEFC54.text	03BF4239FFE9FFE9EAF8C41B99CEFC54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides kirkegaardi Parapar & Martin & Moreira 2020	<div><p>Terebellides kirkegaardi sp. nov.</p><p>Figures 11B, 21A, 22–24, 37; Tables 1, 2</p><p>urn:lsid:zoobank.org:act: 82974B42- 2924-48 A8-BD1A-254FED0F881B</p><p>Material examined. Type material. Holotype (NHMD-231439); paratypes: 37 specimens (NHMD-636923 to NHMD-636925), all from the same original vial as the holotype (Table 1).</p><p>Diagnosis. Body of medium/large size (15–30 mm in length). Branchiae with a long and thick stalk; anterior lobe large; posterior ventral lobes about as long as dorsal ones, with short terminal filaments, emerging directly from branchial stem. TC1 notopodia and notochaetae shorter than following thoracic ones. Thoracic neurochaetae with rostrum / capitium length about 2/1, and capitium with a first row of 2–5 medium-sized teeth followed by progressively smaller ones.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, 28.0 mm long and 2.5 mm wide (Fig. 21A, 22 A−B); body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; large tentacular membrane surrounding mouth, with typical buccal tentacles with expanded tips (Fig. 22 A−Β). SG1 as an expanded structure below tentacular membrane (Fig. 22A, 24A). Lateral lappets and ovalshaped glandular region in TC3 not present.</p><p>Branchiae. Branchiae arising as single structure from SG3, consisting of a single very long (about same length as dorsal posterior lobes) stalked mid-dorsal branch (Fig. 21A, 22 A−D, 24A), with one pair of dorsal (upper) un- fused lobes reaching TC4 or TC6 (if branchial stalk bent backwards); a pair of ventral (lower) lobes about as long as dorsal ones but thinner, not fused together and arising directly from stalk (Fig. 21A, 22 A−D, 23A, 24A). Ante- rior projection of dorsal lobes (lobe 5) very well developed, about half length of posterior dorsal lobes (Fig. 21A, 22 A−D, 23A, 24A). Pointed projection of posterior region of both upper and lower lobes much longer in ventral ones, becoming a terminal filament (Fig. 23A). Both sides of branchial lamellae ciliated, arrangement in rows and/ or tufts or papillae not seen.</p><p>Thorax. Eighteen pairs of notopodia (SG3−20); that of TC1 shorter than subsequent ones (Fig. 24 A−B), with notochaetae as simple capillaries, shorter in number and length than following ones. Neuropodia as sessile pinnules, from TC6 (SG8) to pygidium, with uncini in single rows starting from TC7 (SG9) throughout. First neuropodia (TC6) with 4–5 sharply bent, acute-tipped, geniculate chaetae (Fig. 23B, 24 C−D) having minute teeth forming a very low marked capitium (Fig. 24D). From TC7, neuropodia with 11–17 uncini per torus in one row (Fig. 24E) with long shafted denticulate hooks, rostrum about two times longer than capitium, with 2−5 big teeth above main fang, and surmounted by a crest of smaller denticles (Fig. 24 F−G).</p><p>Abdomen. About 40% of body length, with 31 pairs of neuropodia as erect pinnules with about 20–25 uncini per torus having four teeth above main fang surmounted by a crest of several teeth of same and/or smaller length (Fig. 23D).</p><p>Other features. Papilla on first thoracic notopodia not seen; button-hole-like pairs of dorsal nephridial openings on, at least, each SG7 (TC5) notopodium (Fig. 23 B−C); second pair not seen. Pygidium blunt, as funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 3 solid; CH 4 to CH 9 striped; fading from CH 10 to CH 12; J-shaped glandular region marked lateral to CH 3; pattern 5 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variations. Terebellides kirkegaardi sp. nov. is a medium-sized species (15−30 mm long). Eggs were visible in body cavity of females across all length range. Most specimens are in good conditions, with bodies and branchiae complete.</p><p>Type locality. Cameroon; 8–9 m depth (Table 1) .</p><p>Distribution and bathymetry. Victoria (aka Limbé, Cameroon); 8−9 m depth (Fig. 11B; Table 1).</p><p>Etymology. This species is named in honour of Danish researcher Jørgen Bagger Kirkegaard (1920−2006), who participated in the Galathea and Atlantide Deep Sea Expeditions. His 1958 publication on West African sedentary polychaetes included most of the specimens studied in the present work.</p><p>Remarks. Terebellides kirkegaardi sp. nov. is mostly characterised by its long branchial stalk and well-developed anterior triangular lobe, coupled with a TC1 notopodia shorter than following, and very low marked capitium teeth in TC6 thoracic uncini (only detected with SEM). The branchial appearance is similar across most specimens studied (cfr. Fig. 21A, 22 A–D, 24A)—but see Fig. 22 E–F, 23A—giving rise to a characteristic body profile that differs from any other known species of the genus.</p><p>Terebellides kirkegaardi sp. nov. most closely resembles Terebellides canopus Schüller &amp; Hutchings, 2013, Terebellides mira Schüller &amp; Hutchings, 2013 (both from deep waters off South Western Atlantic Ocean), and Terebellides sepultura Garraffoni &amp; Lana, 2003 (from Brazilian coasts). However, they lack the anterior projection of branchial lobes and posterior lobes are free from each other. Terebellides mira also bears a branchial stem that is much longer than the proper branchial lobes. Two specimens from Myanmar, identified as Terebellides cf. woolawa Hutchings &amp; Peart, 2000 by Parapar et al. (2016a), also show long branchial stems, but differ from T. kirkegaardi sp. nov. in having an even more developed, non-triangular anterior lobe, as well as smaller postero-ventral lobes. Furthermore, T. woolawa as described by Hutchings &amp; Peart (2000) bears a similar anterior lobe, but the branchial stem is much shorter.</p></div>	https://treatment.plazi.org/id/03BF4239FFE9FFE9EAF8C41B99CEFC54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFD5FFECEAF8C7419A23FEA7.text	03BF4239FFD5FFECEAF8C7419A23FEA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides longiseta Parapar & Martin & Moreira 2020	<div><p>Terebellides longiseta sp. nov.</p><p>Figures 11C, 21B, 25−27, 37; Tables 1, 2 urn:lsid:zoobank.org:act: 33AEE210-CA77-488F-A43B-2FDC23375CBE</p><p>Material examined. Type material. Holotype (NHMD-231459); paratypes: five specimens (NHMD-231437, NHMD-231448 and NHMD-636926). Non-type material. One specimen (MNCN 16.01 /18591, Table 1) .</p><p>Diagnosis. Body medium/big sized (up to 21 mm in length). Anterior branchial lobe large; posterior ventral lobes thinner than dorsal ones, with long terminal filament, directly emerging from branchial stem. TC1 about as long as following ones; all thoracic notochaetae very long and numerous, with rostrum vs capitium length of about 1/1, and capitium with a first row of 3 big teeth followed by much smaller ones.</p><p>Description based on holotype</p><p>Measurements and general body features. Incomplete specimen, broken at TC10; 8.0 mm long and 3.5 mm wide (Fig. 21B, 25A). Prostomium compact; large tentacular membrane surrounding mouth; buccal tentacles lost. SGI as an expanded structure below tentacular membrane. Lateral lappets on SG3–6 (TC1–4), especially large in SG3 (TC1) forming a fan-shaped antero-dorsal expansion (Fig. 25 B–C, arrow).</p><p>Branchiae. Branchiae arising as single structure from SG3, with a single stalked mid-dorsal branch having one pair of dorsal (upper) lobes (1+2), not fused along their length, and one pair of ventral (lower) lobes (3+4) of similar length but much thinner, neither fused together nor to dorsal ones (Fig. 21B, 25B, D). Anterior lobe (lobe 5) well developed, triangular, about half to 1/3 length of posterior dorsal lobes (Fig. 25C). Pointed projection of both upper and lower posterior lobes much longer in ventral ones, becoming a long terminal filament (Fig. 25B, D, 26 B−C). Both sides of branchial lamellae with several parallel rows of cilia and low ciliated tufts, giving a zig-zag appearance to outer edge of dorsal posterior lobe branchial lamellae under stereomicroscope (Fig. 25B, D).</p><p>Thorax. Incomplete, with ten pairs of notopodia from SG3 to SG12 (TC1 to TC10). TC1 notopodia well developed, as long as subsequent ones (Fig. 25 B−C, 26A). All notochaetae simple capillaries with textured surface, very long (near body width at same level) and numerous (12–15 per notopodium). Neuropodia as sessile pinnules from TC6 (SG8), with uncini in single rows starting from TC7 (SG9) throughout. First neuropodia (TC6/TU1) with 8–9 sharply bent, acute-tipped, geniculate chaetae, having minute teeth forming a capitium visible under SEM. From TC7, neuropodia with about 20–25 partially extruded uncini per torus in one row, with long shafted denticulate hooks, rostrum about same length as capitium, with three big teeth above main fang surmounted by an upper crest of several much smaller denticles (Fig. 27 A−B).</p><p>Abdomen. Not present in holotype (see below) .</p><p>Other features. Nephridial papilla and nephridial openings not seen (see below).</p><p>Methyl green staining pattern. Not determined; only one complete specimen available but ventrally contract- ed.</p><p>Variability. Only one complete specimen (NHMD-231448), 21 mm long, 1.5 mm wide, with 30 abdominal chaetigers. Specimen NHMD-636926 similar to holotype, with a conspicuous fan-shaped antero-dorsal expansion, branchiae with only ventral posterior lobes, very thin and with about 25 lamellae each, and nephridial openings in TN4−5. One specimen from NHMD-231448 with a scar caused by a copepod attachment near to TC1 notopodium; another complete specimen from same vial (with about 30 abdominal segments) with neuropodia as erect pinnules and about 25–30 uncini per torus having uncini with four teeth above main fang surmounted by one/two smaller ones and an upper crest of smaller teeth (Fig. 27 C–D). Pygidium blunt, as funnel-like depression. Specimens from NHMD-231437 badly preserved, but characteristically long, with abundant notochaetae on last half of thorax and branchiae with terminal filament in posterior ventral lobes and lamellae border conspicuously sinuous, similar to holotype.</p><p>Type locality. Equatorial Guinea; 260–650 m depth (Table 1) .</p><p>Distribution and bathymetry. From Equatorial Guinea to South Angola in SW Africa; 235–650 m depth (Fig. 11C; Table 1).</p><p>Etymology. The epithet refers to the very long and numerous chaetae present in thoracic notopodia.</p><p>Remarks. The conspicuous, long thoracic notochaetae are the most distinctive character of T. longiseta sp. nov. Other species with long notochaetae are Terebellides klemani Kinberg, 1867 (Brazil) and Terebellides irinae Gagaev, 2009 (Arctic Basin), but they are thinner and fewer, 2–4 in T. klemani, and 5–10 in T. irinae . In addition, T. klemani bears TC1 notochaetae shorter than those of following chaetigers; the branchiae of T. irinae lack anterior branchial lobes and posterior lobes originate from a thin and long stalk and are free along their entire length, the ventral being much shorter than the dorsal (see Gagaev, 2009: Fig. a–b).</p><p>Terebellides gentili Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019 also bears conspicuously long notochaetae (not explicitly mentioned in the description but see Lavesque et al., 2019: Fig. 10A). Nevertheless, this species clearly differs from T. longiseta sp. nov. in having fewer and thinner notochaetae and conspicuous papillar projections on the outer edge of the branchial lamellae, that are absent in T. longiseta sp. nov.</p><p>Terebellides yangi resembles T. longiseta sp. nov. in having greatly developed anterior branchial lobes but differs in length and number of thoracic notochaetae (far more numerous and longer in the African species).</p><p>On the other hand, several African non-type specimens identified as T. jorgeni may correspond to T. longiseta sp. nov. (see note below). One specimen from St. 133 showed a scar due to a parasitic copepod, similar to those reported for specimens of T. shetlandica by Parapar et al. (2016c). This was the only parasitized specimen observed across all material examined in this work.</p></div>	https://treatment.plazi.org/id/03BF4239FFD5FFECEAF8C7419A23FEA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFD0FFE3EAF8C26C9B8AFEA4.text	03BF4239FFD0FFE3EAF8C26C9B8AFEA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides nkossa Parapar & Martin & Moreira 2020	<div><p>Terebellides nkossa sp. nov.</p><p>Figures 11C, 28A, 29–32, 37; Tables 1, 2 urn:lsid:zoobank.org:act: 8C262B7A-F8EC-4ABC-A6AE-538652C71D1B</p><p>Material examined. Type material. Fifty specimens: holotype (MNCN16.01 /18594); forty-nine paratypes (MNCN16.01 /18592, 16.01/18593 and 16.01/18595 to 16.01/18610). Non-type material. Five dried-out specimens (MNCN16.01 /18611 and MNCN16.01 /18612); one specimen (NHMD-231434) (Table 1) .</p><p>Diagnosis. Body small sized (5–15 mm in length); semi-transparent body wall. Branchial anterior lobe absent; posterior lobes not fused; ventral ones thinner and shorter than dorsal ones, with long terminal filament, directly emerging from branchial stem. TC1 notopodia and notochaetae smaller than following ones. Thoracic neurochaetae with rostrum / capitium length about 1/1 and capitium with a first row of 4–7 medium-sized teeth followed by progressively shorter ones.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, with coelomic cavity filled with oocytes, 12 mm long and 1.0 mm wide (Fig. 28A, 29A); body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; tentacular membrane surrounding mouth, with a low number of buccal tentacles (Fig. 29A). SGI as an expanded structure below tentacular membrane (lower lip) (Fig. 29A, 30A). Lateral lappets weakly developed on SGIII–VII (CH 1–5).</p><p>Branchiae. Branchiae arising as single structure from SGIII, with a single stalked mid-dorsal branch having of one pair of dorsal (upper) lobes not fused along their lengths, and a pair of ventral (lower) lobes much shorter, neither fused together nor to dorsal ones (Fig. 28A, 29A, 30 A−B, 32A). Anterior projection of dorsal lobes (lobe 5) not present. Pointed projection of posterior region of both upper and lower lobes much longer in ventral ones, becoming a long terminal filament (Fig. 30 A−B, 32A). Both sides of branchial lamellae with several parallel bent rows of cilia and several ciliated tufts near outer edge (Fig. 30 C−D).</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX); those in TC1 slightly smaller than subsequent ones (Fig. 30A, E−F, 32A). Neuropodia as sessile pinnules, from TC6 (SGVIII) to pygidium, with uncini in single rows start- ing from TC7 (SGIX) throughout. First neuropodia (TC6) with 3–4 sharply bent, acute-tipped, geniculate chaetae with well-developed minute teeth forming a capitium (Fig. 31A). From TC7, neuropodia with up to 7–9 uncini per torus in one row (Fig. 31B), with long-shafted denticulate hooks, rostrum about same length as capitium, with 4−7 medium-sized teeth above main fang, surmounted by 4−6 slightly shorter teeth (Fig. 31C, 32 C−D).</p><p>Abdomen. 29−30 pairs of neuropodia, as erect pinnules with about 20 uncini per torus (Fig. 31 D−E) having three teeth above main fang surmounted by two teeth above one in middle, an upper crest of two teeth and a variable number of smaller teeth (Fig. 31F).</p><p>Other features. Paired, mulberry-like nephridial papilla behind first thoracic notopodia and dorsal to each SGVI and VII (CH 4 and CH 5) notopodium (Fig. 30A, E−F, 32A−B). Pygidium blunt, as funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 4 solid; CH 5 to CH 11 striped; CH 12 and CH 13 only neuropodia slightly stained; pattern 1 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variability. Most specimens of T. nkossa sp. nov. are complete, with well-defined branchiae, ranging from 5 to 15 mm long, showing intra-coelomic oocytes from 9-mm long specimens (Fig. 29D). The digestive tract main parts, especially the hind stomach, can be easily seen by transparency through the delicate body wall (Fig. 29C). The only specimen from the Atlantide cruise (NHMD-231434), identified as T. stroemii by Kirkegaard (1959), comes from shallow waters (108 m depth) off Sierra Leone, measures 7 mm in length and overall agrees with T. nkossa sp. nov. description, except in having much shorter, comma-shaped branchiae, both features not present in our specimens of this species, and therefore it was labelled as T. af. nkossa .</p><p>Type locality. Gabon; 43 m depth (Table 1) .</p><p>Distribution and bathymetry. Gabon and Democratic Republic of Congo; 43−250 m depth (Fig. 11C; Table 1).</p><p>Etymology. The species is named after the N’Kossa gas field located off Congo, where most specimens were collected.</p><p>Remarks. The diagnostic characters above mentioned clearly distinguishes T. nkossa sp. nov. from all other species here described. However, it shares some branchial features with six previously described species: Terebellides mundora Hutchings &amp; Peart, 2000 (Australia), Terebellides atlantis Williams, 1984 (New England slope), Terebellides parapari Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019, T. sepultura, T. irinae and T. shetlandica .</p><p>Terebellides mundora differs from T. nkossa sp. nov. in having branchiae with posterior ventral lobes almost as long as dorsal ones and thoracic uncini with rostrum shorter than capitium, while the new species has ventral lobes much shorter and rostrum longer than capitium; T. sepultura has a very large branchial stalk (short in T. nkossa sp. nov.), and T. irinae has very long thoracic notopodial chaetae (short in T. nkossa sp. nov.).</p><p>The Atlantic species T. atlantis, T. shetlandica and T. parapari closely resemble T. nkossa sp. nov. in their small body size ( T. atlantis: 8–18 mm; T. shetlandica: 06– 19 mm; T. parapari: 8–23 mm and T. nkossa sp. nov.: 5–15 mm) and several other morphological features. Thus, even though the brief original description of T. atlantis (Williams, 1984) does not allow accurate comparisons, the Icelandic specimens described by Parapar et al. (2011) differ from T. nkossa sp. nov. in the size and shape of dorsal branchial lobes vs. ventral ones (which are all highly deciduous in the former) and the shape of thoracic uncini, which show the capitium endowed with minute and numerous teeth (type 4 in Parapar et al., 2016c) (long and scarce in T. nkossa sp. nov., type 3 in Parapar et al., 2016c). Terebellides shetlandica and T. parapari most closely resemble T. nkossa sp. nov. in general branchial shape with ventral branchial lobes only fused at base with dorsal ones and provided with a terminal filament and thoracic uncini. Furthermore, all three species were found at a similar depth range: 100–160 m for T. shetlandica, 120 m for T. parapari and 43–250 m for T. nkossa sp. nov. Nevertheless, T. nkossa sp. nov. mostly differs from T. shetlandica in the number of uncini per abdominal neuropodium (6–7 vs. ca. 20, respectively), and from T. parapari in having smaller TC1 notopodial and notochaetae than following (much longer in T. parapari) and having ciliated papillae dorsal to thoracic notopodia and ciliature in branchial lobes (absent in T. parapari).</p><p>Other species with four-lobed branchiae are: Terebellides ehlersi McIntosh, 1885 (Fiji Islands), Terebellides intoshi Caullery, 1944 (Indonesia) and Terebellides lobatus Hartman &amp; Fauchald, 1971 (New England). Nevertheless, all of them are much larger than T. nkossa sp. nov. (30−40 mm vs. 5−15 mm) and T. intoshi also bears two chaetigers with geniculate chaetae (cfr. Imajima &amp; Williams, 1985), instead of one.</p></div>	https://treatment.plazi.org/id/03BF4239FFD0FFE3EAF8C26C9B8AFEA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFDFFFE4EAF8C2719A98FED3.text	03BF4239FFDFFFE4EAF8C2719A98FED3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides ramili Parapar & Martin & Moreira 2020	<div><p>Terebellides ramili sp. nov.</p><p>Figures 11D, 28B, 33, 34, 37; Tables 1, 2 urn:lsid:zoobank.org:act: 97DCCE83-A2BC-48EF-B84A-BB20123827AF</p><p>Material examined. Type material. Four specimens: holotype (MNCN16.01 /18615); three paratypes (MNCN 16.01 /18613, 16.01/18614 and 16.01/18616). Non-type material. One specimen (NHMD-231438, Table 1) .</p><p>Diagnosis. Body small/medium sized (18.5 mm in length); TC1–5 ventrally whitish. Branchial anterior lobe absent; posterior ventral lobes thinner than dorsal ones, with terminal filament and directly emerging from branchial stem. TC1 notopodia and notochaetae as long as following ones. Thoracic neurochaetae with rostrum/capitium length about 1/1, and capitium with a first row of 5−6 small teeth followed by similar sized ones.</p><p>Description based on holotype</p><p>Measurements and general body features. Complete specimen, 18.5 mm long and 1.5 mm wide; female with oocytes in body cavity (Fig. 28B, 33 A−B). Body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; short tentacular membrane surrounding mouth, devoid of buccal tentacles. SGI as an expanded structure below tentacular membrane (Fig. 33A). Lateral lappets on SGIII–VII (TC1–5), larger in SGIII–IV (Fig. 33B).</p><p>Branchiae. Branchiae arising as single structure from SGIII, with a single stalked mid-dorsal branch (Fig. 28B) having one pair of dorsal (upper), non-fused lobes and one pair of shorter ventral (lower) lobes, neither fused together nor to dorsal ones, with terminal filament (Fig. 28B, 33 A−B). Anterior projection of dorsal lobes (lobe 5) absent. Both sides of branchial lamellae with cilia; rows and/or tufts or papillae not seen.</p><p>Thorax. Eighteen pairs of notopodia (SGIII–XX), with those of TC1 as long as subsequent ones. Neuropodia as sessile pinnules, from TC6 (SGVIII) to pygidium, with uncini in single rows starting from TC7 (SGIX) throughout. First neuropodia (TC6) with 4–5 sharply bent, acute-tipped, geniculate chaetae (Fig. 34A) showing minute teeth forming a capitium. From TC7, neuropodia with up to 6–10 uncini per torus in one row, with long shafted denticulate hooks, rostrum about same length as capitium, with 8–10 small teeth above main fang, surmounted by two rows of several teeth of same size (Fig. 34 B–D).</p><p>Abdomen. Thirty-one neuropodia as erect pinnules, with about 25−30 uncini per torus having four teeth above main fang surmounted by an upper crest of 4–5 teeth and a variable number of smaller teeth (Fig. 34 E–F).</p><p>Other body features. Pairs of nephridial papillae both at TC1; dorsal paired nephridial openings in anterior thoracic notopodia not seen. Pygidium blunt, as funnel-like depression. Five anterior thoracic chaetigers (TC1−5) ventro-laterally whitish (Fig. 33A); last two (TC4–5) more pigmented.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 3 solid; CH 4 to CH 12 striped; being CH 4 and CH 5 much more marked than previous ones, and CH 12 faded; J-shaped glandular region slightly marked lateral to CH 2 and CH 3; near pattern 5 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Variability. Body 6.0 mm long in the only complete (and probably immature) paratype (MNCN16.01/18613). Specimen NHMD-231438 (Fig. 33C), collected during the Galathea expedition and originally identified as T. stroemii by Kirkegaard (1959), has been identified here as T. af. ramili . It is a 22-mm long female with oocytes that was collected nearby the type location and at similar depths (Table 1). However, its identity cannot be fully confirmed as colouration of TC4–TC5 is different than others being more widely distributed in TC4 and much less in TC5 (Fig. 33C). Moreover, this specimen bears shorter branchial ventral lobes than in T. ramili sp. nov. and lacks terminal filament.</p><p>Type locality. Off Congo; 190 m depth (Table 1) .</p><p>Distribution and bathymetry. Off Congo and Namibia; 190−537 m depth (Fig. 11D; Table 1).</p><p>Etymology. The species is named after Dr. Francisco Ramil (Universidade de Vigo, Spain), for his numerous contributions to the taxonomy of marine invertebrates, especially Cnidarians, in many world seas but specifically in the Iberian Peninsula, Antarctica and the Atlantic African littoral.</p><p>Remarks. The conspicuous white colouration of T. ramili sp. nov. resembles that of the boreal Terebellides gracilis Malm, 1874, recently reported from Iceland by Parapar et al. (2011) and the Adriatic Sea by Parapar et al. (2013). The type material redescribed by Parapar et al. (2011) shows a homogeneous colour pattern between TC1 and TC4, while in T. ramili sp. nov. it appears from TC1 to TC5 being even more conspicuous in TC4 and TC5. Furthermore, TC1 notopodia and notochaetae in the Mediterranean specimens of T. gracilis are smaller than in other chaetigers (similar throughout or slightly larger in T. ramili sp. nov.) and the thoracic uncini show large anterior teeth in the capitium (Parapar et al., 2011: Fig. 10C). These teeth are closer to Type 1, as it happens in T. longiseta sp. nov. and T. kirkegaardi sp. nov., or Type 2, as in T. fauveli sp. nov., and different to the Type 4 present in T. ramili sp. nov. Moreover, T. ramili sp. nov. also differs from the three aforementioned species in the branchial shape, size of TC1 and ventral colour pattern (see above).</p></div>	https://treatment.plazi.org/id/03BF4239FFDFFFE4EAF8C2719A98FED3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFD8FFFDEAF8C2B99A24FC70.text	03BF4239FFD8FFFDEAF8C2B99A24FC70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides	<div><p>Terebellides sp. 1</p><p>Figures 11D, 35 A–B, 37; Tables 1, 2</p><p>Material examined. One specimen (NHMD-231440, Table 1).</p><p>Description</p><p>Measurements and general body features. Complete and well preserved specimen, 29 mm long and 2.5 mm wide (Fig. 35 A−B); body tapering posteriorly with segments increasingly shorter and crowded towards pygidium. Prostomium compact; very large tentacular membrane surrounding mouth, with typical buccal tentacles with expanded tips. SGI as an expanded structure below tentacular membrane. Lateral lappets on SGIII–VII (TC1–5).</p><p>Table 1. Locality and collecting data, museum registration numbers and reference to figures, of all Terebellides specimens described in this work. Country names and coordinates are copied from original museum labels.</p><p>…continued on the next page</p><p>Table 1. (continued)</p><p>…continued on the next page</p><p>Table 1. (continued) Angola (Off 130 “ “ 22-12-1950 17º13’S 11º42’E 75 1 NHMD-231444 – paratypes Cunene river) “ 131 “ “ 22-12-1950 17º13’S 11º42’E 100 1 NHMD-231445 – paratypes “ 132 “ “ “ 17º13’S 11º27’E 200 1 NHMD-231447 – paratypes MNCN non-type; Congo 16-2 this work N’Kossa 2003 5°16.839’S 11°33.966’E 150 1 – 16.01/18585 SEM MNCN non-type; “ 17-1 “ “ “ 5°16.365’S 11°33.655’E 110 1 – 16.01/18586 SEM MNCN “ NK0300E(B) “ “ 2015 5º16.4240’S 11º34.0065’E 158 2 – non-type 16.01/18587 MNCN non-type; 19C–F, “ N1S 0500S “ “ “ 5º19.4803’S 11º35.8724’E 137 1 – 16.01/18588 SEM 20A–F MNCN “ N1S 300S “ “ “ 5º20.7549’S 11º36.3353’E 136 1 – non-type 16.01/18589 MNCN non-type; “ NK0500S(A) “ “ “ 5º16.7202’S 11º33.8665’E 167 1 – 19A–B 16.01/18590 SEM Terebellides kirkegaardi sp. nov. original new campaign/oil country station-sample reference date longitude latitude depth specs registration registration Remarks Figures field number number Cameroon Kirkegaard 21A, (Bota at 61 Galathea 01-12-1950 n.d. n.d. 8–9 1 NHMD-231439 – holotype (1959) 22B Victoria) NHMD- “ “ “ “ “ “ “ “ 35 “ paratypes 22A, C-F 636923 NHMD- paratype; “ “ “ “ “ “ “ “ 1 “ 23A–D 636924 SEM NHMD- paratype, “ “ “ “ “ “ “ “ 1 “ 24A–G 636925 SEM</p><p>…continued on the next page</p><p>Table 1. (continued) NHMD- “ “ “ “ “ “ “ “ 1 “ paratypes 636926 MNCN 26A–D, Congo 11_2 this work N’Kossa 2003 5°22.708S 11°27.627E 545 1 – non-type; SEM 16.01/18591 27A–D Kirkegaard Angola 135 Atlantide 17-03-1946 7º55’S 12º38’E 235–460 2 NHMD-231437 – paratypes (1959) Kirkegaard “ 133 “ 1950 17º13’S 11º27’E 320 2 NHMD-231448 -– paratypes 25C (1959) Terebellides nkossa sp. nov. original new station- campaign/oil country reference date longitude latitude depth specs registration registration Remarks Figures sample field number number non-type; Kirgegaard Sierra Leone 51 Atlantide 31-1-1945 7º14’N 12º57’W 108 1 NHDM-231434 – labelled as T. (1959) af. nkossa MNCN Gabon AYM 34-1 this work Ayol 2011 1º19.82’ 8º43.8’ 43 3 – paratype 16.01/18592 MNCN “ AYM 34-2 “ Ayol “ “ “ “ 3 – paratype 16.01/18593 MNCN “ AYM 34-2 “ Ayol “ “ “ “ 1 – holotype 28A, 29A 16.01/18594 MNCN “ AYM 35-1 “ Ayol “ 1º20.3’ 8º44.0’ 44 7 – paratype 29B–C 16.01/18595 MNCN “ “ “ Ayol “ “ “ “ 1 – paratype; SEM 30A–F 16.01/18596 MNCN “ AYM 35-2 “ Ayol “ “ “ “ 5 – paratype 16.01/18597 MNCN “ AYM 36-1 “ Ayol “ 1º20.6’ 8º44.1’ 44 4 – paratype 16.01/18598 MNCN “ AYM 36-2 “ Ayol “ “ “ “ 1 – paratype; SEM 32A–D 16.01/18599 MNCN “ “ “ Ayol “ “ “ “ 5 – paratype 16.01/18600 MNCN “ AYM 37-1 “ Ayol “ 1º21.8’ 8º44.8’ 45 1 – paratype; SEM 31A–F 16.01/18601 MNCN “ “ “ Ayol “ “ “ “ 1 – paratype 29D 16.01/18602 MNCN “ “ “ Ayol “ “ “ “ 7 – paratype 16.01/18603 MNCN “ AYM 37-2 “ Ayol “ “ “ “ 7 – paratype 16.01/18604 Banc du MNCN “ BDP 55-1 “ “ 0º37.5’ 8º45.6’ 47 1 – paratype Prince 16.01/18605 …continued on the next page</p><p>Table 1. (continued)</p>Terebellides ramili sp. countrystation- samplereferencecampaign/oil fielddatelongitudelatitudedepthspecsoriginal registration numbernew registration numberRemarksFiguresCongo2This workN’Kossa18-11-20005º17.419’S11º34.379’E1801CEAB,129GMNCN 16.01/18613paratype“7““18-11-20005°16.425’S11°33.765’E2001CEAB.129DMNCN 16.01/18614paratype; SEM34A–F“9““17-11-20005°16.285’S11°33.568’E1901CEAB.AP.129FMNCN 16.01/18615Holotype28B, 33A–B““““17-11-2000“““2“MNCN 16.01/18616paratypeCongo75Kirkegaard (1959) Galathea07-12-19505º41’S11º38’E2051NHMD-231438– non-type; labelled T. af. ramili33C nov.<p>Terebellides sp. 2 original new station- campaign/oil country reference date longitude latitude depth specs registration registration Remarks Figures sample field number number</p><p>…continued on the next page</p><p>Table 1. (continued) South Africa Kirkegaard (Off Cape 165 Galathea 03-01-1951 n.d. n.d. 110 1 NHMD-231449 – dried out 35C–D (1959) Point)</p><p>Branchiae. Branchiae arising as single structure from SGIII, consisting of a single stalked mid-dorsal branch having one pair of dorsal (upper) lobes not fused along their length, and one pair of ventral (lower) lobes of same length, neither fused together nor to dorsal ones, arising from branchial stem. Anterior projection of dorsal lobes (lobe 5) small. Pointed projection of posterior region of both upper and lower lobes forming a short terminal papilla (Fig. 35 A−B).</p><p>Thorax. Eighteen pairs of notopodia (SGIII−XX), with those in TC1 smaller than subsequent ones. Neuropodia as sessile pinnules, from TC6 (SGVIII) to pygidium, with uncini in single rows starting from TC7 (SGIX) throughout. First neuropodia (TC 6) with eight sharply bent, acute-tipped, geniculate chaetae; second and subsequent neuropodia with up to 14−16 uncini per torus.</p><p>Abdomen. Thirty-one pairs of neuropodia as erect pinnules, with ca. 25 uncini per torus.</p><p>Other body features. A very small nephridial papilla behind first thoracic notopodia; two pairs of simple nephridial openings dorsal to each SGVI and VII (TC 4 and TC 5) notopodium. Pygidium blunt, as a funnel-like depression.</p><p>Methyl green staining pattern. Anterior CH 1 to CH 3 solid; CH 4 to CH 18 striped; more pronounced in CH 4 to CH 11; near pattern 1 of Schüller &amp; Hutchings (2010) (Fig. 37).</p><p>Distribution and bathymetry. Congo; 200 m depth (Fig. 11D; Table 1).</p><p>Remarks. The specimen was not examined under SEM, preventing adequate observations of relevant characters (i.e., ciliation of branchial lamellae, teeth size and shape on thoracic uncini) and, thus, formal description. It is characterised by the great development of the tentacular membrane, a trait shared with T. californica and T. mediterranea which, in turn, differ from Terebellides sp. 1 in having long TC1 notopodium and notochaetae, both smaller in this specimen. Terebellides resomari Lavesque, Hutchings, Daffe, Nygren &amp; Londoño-Mesa, 2019 also shares this character, but differs from Terebellides sp. 1 in having papillar projections over the margin of the branchial lamellae and a well-developed fifth branchial lobe.</p></div>	https://treatment.plazi.org/id/03BF4239FFD8FFFDEAF8C2B99A24FC70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFC1FFFDEAF8C71D990BF904.text	03BF4239FFC1FFFDEAF8C71D990BF904.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides	<div><p>Terebellides sp. 2</p><p>Figures 11D, 35 C–D; Tables 1, 2</p><p>Material examined. One specimen (NHMD-231449, Table 1).</p><p>Description. Complete specimen, not well-preserved, 11 mm long and 1.0 mm wide (Fig. 35 C–D), mature female, with numerous intra-coelomic oocytes. Anterior thoracic segments (TC1−5) ventrally white. General bran- chial structure not well defined because the state of preservation, not clearly corresponding to any species described in this work.</p><p>Distribution and bathymetry. South Africa (off Cape Point); 110 m depth (Fig. 11D; Table 1).</p><p>Remarks. Terebellides stroemii was widely reported by Day (1967) along the coasts of South Africa, including Cape Point, nearby where this specimen was collected. This author provided a brief description of his material but did not mention explicitly where it was collected. However, these specimens clearly differ from Terebellides sp. 2 at least in two characters: larger body size (up to 70 mm) and a much larger shelf-like mouth lower lip (Day, 1967: Fig. 36.1. f-g). Moreover, Day’s drawings show well-developed anterior and posterior branchial lobes, posterior ventral ones being shorter than dorsal ones and not fused to the latter (although reported as “partially fused” in the description); such drawings illustrate the TC6 geniculate chaetae and the thoracic and abdominal uncini but, unfortunately, these characters could not be examined in detail in our specimen without causing irreversible damage. This set of characters does not allow us to assign this specimen neither to any of the above described species nor to Day’s material.</p></div>	https://treatment.plazi.org/id/03BF4239FFC1FFFDEAF8C71D990BF904	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
03BF4239FFC3FFFEEAF8C00F9DD4FC7C.text	03BF4239FFC3FFFEEAF8C00F9DD4FC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Terebellides	<div><p>Key to intertropical South Atlantic species of Terebellides</p><p>The following key comprises the eight species described here, together with those previously described or recorded from Western Atlantic (Williams, 1984; Solís-Weiss et al., 1991; Garraffoni &amp; Lana, 2003; Díaz-Díaz &amp; Liñero- Arana, 2003), Caribbean Sea (Williams, 1984) and southern Brazil (Bremec &amp; Elías, 1999; Garraffoni &amp; Lana, 2003; Schüller &amp; Hutchings, 2012). No material of these species was examined; characters were obtained from bibliography. It is mostly based on branchial characters, number of thoracic chaetigers and location of geniculate chaetae, but also on the thoracic colouration, presence of dorsal hump, and length of thoracic notochaetae.</p><p>Terebellides banalis Schüller &amp; Hutchings, 2012 and Terebellides bulbosa Schüller &amp; Hutchings, 2012 are not considered here because the original descriptions do not provide detailed information on branchiae. In T. banalis, only one type specimen has branchiae, consisting of a stout stem with four very small and stout branchial lobes (Schüller &amp; Hutchings, 2012: Fig. 7A, 8B), suggesting it may be a juvenile. The species is characterized by having well-developed S2, ventrally wider than S3 (TC1), and by TC6 cane-shaped geniculate chaetae (Schüller &amp; Hutchings, 2012: Fig. 8E). Terebellides bulbosa bears a branchial stem lacking well defined lobes and shows very large lappets in the first four chaetigers, followed by a series of laterally expanded segments before a typical Terebellides - like posterior thorax (Schüller &amp; Hutchings, 2012: Fig. 10).</p><p>1. First unciniger in TC5; 17 or 18 thoracic uncinigers.......................................................... 2</p><p>- First unciniger in TC6; 18 thoracic uncinigers............................................................... 3</p><p>2. 17 thoracic uncinigers............................................................. T. anguicomus Müller, 1858</p><p>- 18 thoracic uncinigers....................................................... T. crux Schüller &amp; Hutchings, 2013</p><p>3. Anterior dorsum with a distinct hump on TC7............................................................... 4</p><p>- Anterior dorsum smoothly curved........................................................................ 5</p><p>4. Ventral branchial lobes more slender than dorsal ones; lateral lappets from TC3 to TC5.................................................................................. T. carmenensis Solís-Weiss, Fauchald &amp; Blankensteyn, 1991</p><p>- Both branchial lobes similar in size; lateral lappets from TC5 to TC7..................... T. totae Bremec &amp; Elías, 1999</p><p>5. TC1 to TC5 ventrally whitish (Fig. 33 A–B)........................................... Terebellides ramili sp. nov.</p><p>- TC1 to TC4 similar in colour as following chaetigers......................................................... 6</p><p>6. Anterior branchial lobe well developed.................................................................... 7</p><p>- Anterior branchial lobe absent or much shorter than posterior lobes............................................. 13</p><p>7. Branchial stem long, about as long as posterior dorsal lobes (Fig. 21A)................. Terebellides kirkegaardi sp. nov.</p><p>- Branchial stem much shorter than posterior dorsal lobes....................................................... 8</p><p>8. TC1 notopodia and notochaetae much more developed than following........................................... 9</p><p>- TC1 notopodia and notochaetae similar to following........................................................ 10</p><p>9. Ventral posterior branchial lobes length about 10% of dorsal ones, located after the fusion line of the latter (Fig. 12A, 13A)............................................................................ Terebellides congolana sp. nov.</p><p>- Ventral posterior branchial lobes length about 50% of dorsal ones, emerging directly from the branchial stem (Fig. 12B, 16C)............................................................................. Terebellides fauveli sp. nov.</p><p>10. Thoracic notochaetae almost as long as thoracic width; TC1 lateral lappet forming a fan-like dorsal expansion (Fig. 21B, 25 B–C)..................................................................... Terebellides longiseta sp. nov.</p><p>- Thoracic notochaetae much shorter than thoracic width; TC1 lateral lappet not forming a fan-like dorsal expansion....... 11</p><p>11. TC1 notochaetae short, scarce................................. T. lanai Solís-Weiss, Fauchald &amp; Blankensteyn, 1991</p><p>- TC1 notochaetae similar to following ones................................................................ 12</p><p>12. 20–26 abdominal chaetigers.......................... Terebellides parvus Solís-Weiss, Fauchald &amp; Blankensteyn, 1991</p><p>- At least 30 abdominal chaetigers...................................................... T. klemani Kinberg, 1867</p><p>13. Branchial lobes neither fused nor comma-shaped........................................................... 14</p><p>- Branchial lobes at least partially fused, comma-shaped....................................................... 19</p><p>14. Branchial posterior ventral lobes with long terminal filament (Fig. 28A, 30 A–B)............. Terebellides nkossa sp. nov.</p><p>- Branchial posterior ventral lobes lacking long terminal filament............................................... 15</p><p>15. Anterior thoracic chaetigers narrow, laterally expanded like a partially compressed accordion.................................................................................... Terebellides concertina Schüller &amp; Hutchings, 2012</p><p>- Anterior thoracic chaetigers different..................................................................... 16</p><p>16. Lower buccal lip and tentacular membrane greatly expanded.................................................. 17</p><p>- Lower buccal lip and tentacular membrane different......................................................... 18</p><p>17. Branchial lobes completely free from each other, bearing few and loose lamellae; upper lip of tentacular membrane small................................................................... Terebellides diva Schüller &amp; Hutchings, 2012 Branchial lobes not fused but well defined, with numerous and well-packed lamellae; upper lip of tentacular membrane much developed (Fig. 35 A–B)................................................................... Terebellides sp. 1</p><p>18. Pygidium with two prominent lateral papillae.......................... Terebellides gingko Schüller &amp; Hutchings, 2012</p><p>- Pygidium lacking papillae......................................... Terebellides sepultura Garraffoni &amp; Lana, 2003</p><p>19. Branchial posterior ventral lobes similar in shape to dorsal ones, about half longer, located at base of dorsal ones (Fig. 2B, 8E); thoracic uncini capitium with first row of 5–6 teeth, similar in size to following ones (Fig. 10 A–B)................................................................................................. Terebellides augeneri sp. nov.</p><p>- Branchial posterior ventral lobes much shorter than dorsal ones................................................ 20</p><p>20. Branchial posterior ventral lobes near distal end of dorsal ones (Fig. 2A, 4A, 5A); thoracic uncini capitium with first row of 1–2 teeth, much bigger than following (Fig. 7A).......................... Terebellides africana Augener, 1918 stat. prom.</p><p>- Branchial posterior ventral lobes emerging at base of dorsal ones.......... Terebellides malvinensis Bremec &amp; Elías, 1999</p></div>	https://treatment.plazi.org/id/03BF4239FFC3FFFEEAF8C00F9DD4FC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Parapar, Julio;Martin, Daniel;Moreira, Juan	Parapar, Julio, Martin, Daniel, Moreira, Juan (2020): On the diversity of Terebellides (Annelida, Trichobranchidae) in West Africa, seven new species and the redescription of T. africana Augener, 1918 stat. prom. Zootaxa 4771 (1): 1-61, DOI: 10.11646/zootaxa.4771.1.1
