taxonID	type	description	language	source
03B187BCFF948106FF36DE81FDC1F987.taxon	materials_examined	MATERIAL EXAMINED. No specimens examined by authors.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF948106FF36DE81FDC1F987.taxon	distribution	DISTRIBUTION. Endemic to Kyrgyzstan. NOTES. The species was described from Kyrgyzstan, Naryn Oblast, Terek Valley NW Durbeldzin [= Baetov; ~ 41 ° 21 ' N 74 ° 53 ' E (authors note)], 2000 m, 27 – 28. VII 2009 based on 2 males and 10 females (Skalický, 2010).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF948106FF36D853FE70FABA.taxon	materials_examined	MATERIAL EXAMINED. Jalal-Abad Oblast: Chatkal District, vicinity Sumsar, 41 ° 13 ' 54 " N 71 ° 19 ' 00 " E, 24. VI 2004, 33 ex. (A. Rubenets leg., IBIW); 5 km E Shekaftar, S Karyn-Kur, 41 ° 13 ' 28 " N 71 ° 23 ' 26 " E, 990 m, foothill desert, at light, 26 – 27. VI 2017, 1 ex. (S. Litovkin leg., PCL); 2 km S Sumsar, 41 ° 11 ' 52 " N 71 ° 18 ' 27 " E, 1017 m, clayish shore of stream, 5. VI 2018, 1 ex. (S. Litovkin leg., PCL); 5 km W Tash-Kumyr, 41 ° 20 ' 08 " N 72 ° 07 ' 57 " E, 766 m, hilly terrain, dry riverbed, at light, 16 – 17. VII 2017, 2 ♂ (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF948106FF36D853FE70FABA.taxon	distribution	DISTRIBUTION. Kyrgyzstan: North Kirgizia (Ovchinnikov, 1996, as Heterocerus campestris Motsch.); Osh Oblast, Alai Mountain Range, 10 km N Daraut Kurgan, Tengizbai Pass, 3000 – 3800 m (Mascagni, 2003). Augyles flavidus is distributed in Africa, South of Eastern Europe, Middle East, Caucasus, Central Asia, (Mascagni, 2006). Therefore, the records of this species from Siberia and the Russian Far East (Mascagni, 2016) are erroneous and require confirmation. The record of Heterocerus (Littorimus) flavidus from “ Kirg. ” (“ Кирг. ”) (Jacobson, 1913) refers to the Kirghiz steppe. NOTES. The finding of this species on the Tengizbai Mountain Pass at the altitudes of 3000 – 3800 m above sea level (Mascagni, 2003) is the highest registered locality for the family Heteroceridae. Most likely, the record corresponds to the lower range of the indicated heights. A new material is needed to confirm the occurrence of Heteroceridae in such extreme environments.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978105FF36DA1BFE03FDF2.taxon	description	Figs 1 – 3	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978105FF36DA1BFE03FDF2.taxon	materials_examined	MATERIAL EXAMINED. Jalal-Abad Oblast: 5 km E Shekaftar, S Karyn-Kur, 41 ° 13 ' 28 " N 71 ° 23 ' 26 " E, 990 m, foothill desert, at light, 26 – 27. VI 2017, 58 ex. (S. Litovkin leg., PCL); 10 km NW Jalal-Abad, 40 ° 56 ' 00 " N 72 ° 53 ' 20 " E, 917 m, hilly terrain, pistachio woodlands, at light, 15 – 16. VII 2017, 1 ex. (S. Litovkin leg., PCL). Osh Oblast: 1.5 km N Nayman Reservoir, 40 ° 21 ' 45 " N 72 ° 21 ' 49 " E, 1228 m, pasture, at light, 7 – 8. VII 2017, 2 ex. (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978105FF36DA1BFE03FDF2.taxon	distribution	DISTRIBUTION. North Africa, Middle East, Caucasus, Central Asia (Mascagni, 2016), Ciscaucasia in Russia (Sazhnev & Ilyina, 2017). Here this species is recorded from Kyrgyzstan for the first time.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978105FF36D829FBE5FA7D.taxon	materials_examined	MATERIAL EXAMINED. Issyk-Kul Oblast: near Issyk-Kul Lake, 5 km S Balykchi, 42 ° 23 ' 03 " N 76 ° 11 ' 10 " E, 1615 m, at light, 8 – 9. VII 2018, 2 ex. (S. Litovkin leg., PCL). JalalAbad Oblast: 5 km W Tash-Kumyr, 41 ° 20 ' 08 " N 72 ° 07 ' 57 " E, 766 m, hilly terrain, dry riverbed, at light, 15 – 16. VI 2017, 4 ex. (S. Litovkin leg., PCL); same locality data, but 16 – 17. VII 2017, 15 ex. (S. Litovkin leg., PCL); 10 km WNW Tash-Kumyr, 41 ° 23 ' 15 " N 72 ° 06 ' 05 " E, 1012 m, hilly terrain, steppe, at light, 17 – 18. VI 2017, 1 ex. (S. Litovkin leg., PCL); Chatkal District, vicinity Sumsar, 41 ° 13 ' 54 " N 71 ° 19 ' 00 " E, 24. VI 2004, 26 ex. (A. Rubenets leg., IBIW); 5 km E Shekaftar, S Karyn-Kur, 41 ° 13 ' 28 " N 71 ° 23 ' 26 " E, 990 m, foothill desert, at light, 26 – 27. VI 2017, 85 ex. (S. Litovkin leg., PCL). Osh Oblast: Ozgur environs, bank of Akbura River, 40 ° 24 ' 52 " N 72 ° 52 ' 23 " E, 1127 m, near small pool, 6. VII 2017, 1 ex. (S. Litovkin leg., PCL); Nayman Reservoir, 40 ° 20 ' 31 " N 72 ° 22 ' 22 " E, 1200 m, muddy shore, 21 – 23. VI 2017, 10 ex. (S. Litovkin leg., PCL); 1.7 km N Nayman Reservoir, 40 ° 21 ' 53 " N 72 ° 21 ' 39 " E, 1240 m, small residual pools in dry streambed, 8. VII 2017, 1 ex. (S. Litovkin leg., PCL); same locality data, but 27. V 2018, 1 ex. (S. Litovkin leg., PCL); 1.5 km N Nayman Reservoir, 40 ° 21 ' 45 " N 72 ° 21 ' 49 " E, 1228 m, pasture, at light, 8 – 9. VII 2017, 7 ex. (S. Litovkin leg., PCL); Almalyk, 40 ° 23 ' 04 " N 72 ° 42 ' 31 " E, 1290 m, pasture, at light, 2 – 3. VII 2018, 20 ex. (S. Litovkin leg., PCL). Talas Oblast: Kirov Reservoir, ~ 1 km N Kyzyl-Adyr, 42 ° 38 ' 10 " N 71 ° 36 ' 03 " E, 891 m, 11. VI 2016, 1 ex. (D. Palatov leg., IBIW).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978105FF36D829FBE5FA7D.taxon	distribution	DISTRIBUTION. Kyrgyzstan: North Kirghizia, Issyk-Kul hollow, Inner and Central Tien-Shan (Ovchinnikov, 1996, as Heterocerus laevigatus Panz.); Issyk-Kul Lake environs (Skalický, 2010). It is a widespread Holarctic species, also known from the Oriental region (Charpentier, 1979; Mascagni, 2003, 2016; King & Lago, 2012). NOTES. The note “ Kirg. ” in the Jacobson's Catalog (1913) refers to the Kirghiz steppe.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978104FF36DEC0FEC1F99D.taxon	materials_examined	MATERIAL EXAMINED. Issyk-Kul Oblast: Issyk-Kul District, Grigorievka N IssykKul Lake, 11. VII 1984, 1 ex. (M. Danilevsky leg., MSPU); near Issyk-Kul Lake, 5 km S Balykchi, 42 ° 23 ' 03 " N 76 ° 11 ' 10 " E, 1615 m, at light, 8 – 9. VII 2018, 1 ex. (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF978104FF36DEC0FEC1F99D.taxon	distribution	DISTRIBUTION. Kyrgyzstan: areas around Fergana Valley (Ovchinnikov, 1996). This species is widespread in the Palearctic region and reaching to Afrotropical region (Mascagni, 2016). But the Heterocerus flexuosus species complex is require revision. of Shekaftar): 1 – body, dorsal view; 2 – 3 – aedeagus; 4, 5 – Heterocerus mus, male (vicinity of Tash-Kumyr): 4 – body, dorsal view; 5 – tegmen. Scale bars for Figs 1, 4 = 1 mm, for Figs 3, 5 = 0.1 mm.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DA15FD9CFE2D.taxon	materials_examined	MATERIAL EXAMINED. Issyk-Kul Oblast: near Issyk-Kul Lake, 5 km S Balykchi, 42 ° 23 ' 03 " N 76 ° 11 ' 10 " E, 1615 m, at light, 8 – 9. VII 2018, 5 ex. (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DA15FD9CFE2D.taxon	distribution	DISTRIBUTION. Kyrgyzstan: North Kirgizia and Issyk-Kul hollow (Ovchinnikov, 1996). Europe, Turkey, Iran, Central Asia, Siberia (Mascagni 2006, 2016; Efimov & Litovkin, 2015; Prokin et al., 2016). NOTES. The records “ Kirgisia ” (Zaitsev, 1908) and “ Kirg. ” in the Catalog of Jacobson (1913) refer to the Kirghiz steppe.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36D912FD9CFCE8.taxon	materials_examined	MATERIAL EXAMINED. Issyk-Kul Oblast: Issyk-Kul District, Grigorievka N IssykKul Lake, 11. VII 1984, 1 ex. (M. Danilevsky leg., MSPU). Jalal-Abad Oblast: 5 km W Tash-Kumyr, 41 ° 20 ' 08 " N 72 ° 07 ' 57 " E, 766 m, hilly terrain, dry riverbed, at light, 15 – 16. VI 2017, 2 ex. (S. Litovkin leg., PCL); 5 km E Shekaftar, S Karyn-Kur, 41 ° 13 ' 28 " N 71 ° 23 ' 26 " E, 990 m, foothill desert, at light, 26 – 27. VI 2017, 1 ex. (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36D912FD9CFCE8.taxon	distribution	DISTRIBUTION. Kyrgyzstan: North Kirghizia (Ovchinnikov, 1996). Europe, Middle East, Caucasus, Central Asia, Siberia, Mongolia (Efimov & Litovkin, 2015; Mascagni, 2016). NOTES. The records “ Kirgisia ” (Zaitsev, 1908) and “ Kirg. ” in the Catalog of Jacobson (1913) refer to the Kirghiz steppe.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DF4AFC78FA92.taxon	description	Figs 4 – 5	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DF4AFC78FA92.taxon	materials_examined	MATERIAL EXAMINED. Jalal-Abad Oblast: 5 km W Tash-Kumyr, 41 ° 20 ' 08 " N 72 ° 07 ' 57 " E, 766 m, hilly terrain, dry riverbed, at light, 16 – 17. VII 2017, 2 ♂, 4 ♀ (S. Litovkin leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DF4AFC78FA92.taxon	distribution	DISTRIBUTION. This species was described from Chad (Charpentier, 1965) and distributed in Afrotropical region, Middle East, Central Asia, Pakistan and India (Skalický, 2001; Mascagni, 2016). NOTES. Here this species is recorded from Kyrgyzstan for the first time.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36D850FE03FBF1.taxon	materials_examined	MATERIAL EXAMINED. Issyk-Kul Oblast: Issyk-Kul District, Grigorievka N IssykKul Lake, 11. VII 1984, 1 ex. (M. Danilevsky leg., MSPU).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36D850FE03FBF1.taxon	distribution	DISTRIBUTION. Kyrgyzstan: North Kirghizia and areas around Fergana Valley (Ovchinnikov, 1996, as Heterocerus parallelus Kryn.). Europe, Lebanon, Central Asia, southern Siberia, Mongolia (Mascagni, 2016). NOTES. The record of this species from “ Kirgiz. ” in the Catalog of Jacobson (1913) refers to the Kirghiz steppe.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DE68FCFFF9FB.taxon	materials_examined	MATERIAL EXAMINED. No specimens examined by authors.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF918103FF36DE68FCFFF9FB.taxon	distribution	DISTRIBUTION. Kyrgyzstan: environs of Issyk-Kul Lake (Skalický, 2010). Oriental Region, Hong Kong, Nepal, Pakistan, Turkmenistan (Mascagni, 2016). NOTES. The species was recorded from Turkmenistan without an exact locality (Mascagni 2006, 2016). Issyk-Kul Lake in Kyrgyzstan is the northernmost finding of the species and located at a significant distance from the main known range.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36D925FE51FCD5.taxon	description	2006, 2016). The record “ Kirg. ” for “ Heterocerus (Littorimus) hispidulus Kies. ” in the Jacobson's Catalog (1913) refers to the Kirghiz steppe (Kazakhstan or Russia). Records from the south Siberia and the Russian Far East (Jacobson, 1913; Egorov, 1989; Kirejtshuk, 2001) should be attributed to other species of the Augyles cribratellus species group (sensu Charpentier, 1965). The record of this species from Issyk-Kul hollow in Kyrgyzstan (Ovchinnikov, 1996) require verification.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36D8C2FD05FB44.taxon	description	(e. g. Jacobson, 1913) are doubtful. This species can be confused with other Augyles species and we exclude it from the fauna of Kyrgyzstan.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36DA1AFC14FDD9.taxon	materials_examined	MATERIAL EXAMINED. Thailand: Prov. Nakhon Ratchasima, 14 ° 16 ' 40 " N 102 ° 25 ' 29 " E, 250 m, 9 – 13. VII 2013, 1 ♂ (A. Korshunov leg., PCL).	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36DA1AFC14FDD9.taxon	distribution	DISTRIBUTION. Kyrgyzstan: environs of Issyk-Kul Lake (Skalický, 2010). Oriental region, South China, Nepal, Pakistan (Mascagni, 2016). NOTES. Issyk-Kul Lake in Kyrgyzstan is the northernmost finding of the species and located at a significant distance from the main known range. This species is firstly recorded herein from Thailand.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36DA1AFC14FDD9.taxon	description	DOUBTFUL RECORDS FROM KYRGYZSTAN	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
03B187BCFF908102FF36D829FBD6FC7E.taxon	description	(Mascagni, 2006, 2003, 2016). The record “ Kirg. ” for this species in the Jacobson's Catalog (1913) refers to the Kirghiz steppe. It was recorded from Kyrgyzstan (North Kirghizia, IssykKul hollow and Inner Tien-Shan) by Ovchinnikov (1996), but these records require verification.	en	Litovkin, S. V., Sazhnev, A. S. (2018): The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist 372: 25-32, DOI: 10.25221/fee.372.2, URL: http://dx.doi.org/10.25221/fee.372.2
