identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B187855C42FFFAABACD3DFFC3A6611.text	03B187855C42FFFAABACD3DFFC3A6611.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratoxenus Mann 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Ceratoxenus Mann, 1923</p>
            <p> Ceratoxenus Mann 1923: 360 (original description, included  C. tricornis Mann, 1923 ). </p>
            <p> Xenopelta Mann, 1923 (synonymized by Emerson 1935: 369); Seevers 1957: 106 (redescription as  Xenopelta ). </p>
            <p> Type species:  Ceratoxenus tricornis Mann, 1923 (designation by monotypy) </p>
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	https://treatment.plazi.org/id/03B187855C42FFFAABACD3DFFC3A6611	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
03B187855C43FFFDABACD72BFF4E6541.text	03B187855C43FFFDABACD72BFF4E6541.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratoxenus tricornis Mann 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ceratoxenus tricornis Mann, 1923</p>
            <p>(Figs. 1A–D, 2A–C, 6A, D)</p>
            <p> Ceratoxenus tricornis Mann 1923: 360 (original description); </p>
            <p> Xenopelta tricornis ; Emerson 1935: 369 Seevers 1957: 107 (redescription as  X. tricornis ); Jacobson et al. 1986: 64 (redescription as  X. tricornis ). </p>
            <p>Diagnosis: The species is easily recognized by the presence of long bristles on vertex, head lateral projections composed of two long and broad frontal processes in front of each antennal insertion, transpassing the antennal insertions anteriorly; medial process between lateral projection weak and obtuse, gena straight, not carinate; pronotum with anterior impression longer and slightly narrower than posterior, and anterior margin straight.</p>
            <p>Head with gena straight; long bristles present on vertex; surface roughly scamose; head lateral projections at frons composed of two long and broad frontal processes in front of each antennal insertion, transpassing the antennal insertions anteriorly; medial process acutely projected, not overtaking the level of frontal elevations; frontoclypeal suture with weak and obtuse medial process, between frontal projections (Figs. 1A–B, 2A, C, 6A). Head surface somewhat scamose. Antenna with antennomere 1 cylindrical, with almost same length of antennomeres 2–4 combined; 2–4 longer than wide, subequal in length; 5–7 shorter than previous, longer than wide, about same length; 8 slightly shorter than previous segments and longer than segments 9–10, subquadrate; 9–10 transverse, subequal. Secondary maxilar palpomere slightly rounded and inflated at apex. Postmentum occupying more than half of total head length, longer than wide, slightly longer than gula, entirely covered by bristles.</p>
            <p>Thorax. Pronotal disc light brown with lateral margins darker; anterior impression longer and slightly narrower than posterior, and with anterior margin straight; bearing long bristles marginally; posterior half of pronotal disc with circular impression occupying almost entire pronotal width, with slight circular elevation medially; anterior region slightly elevated behind head, with one row of three bristles; lateral margins parallel with longitudinal row of three long bristles symmetrically distributed (Fig. 1C, 2C). Elytra elongate, wider on posterior region; light-brown, outer margin darker. Legs with pro- and mesocoxa transverse; metacoxa triangular. Tibia slightly longer than femur, covered with long and thick bristles along entire length; outer margins of tibia slightly convex, and inner margin straight; pro-, meso-, and metatarsi increasing in length.</p>
            <p>Abdomen. As described to genus.</p>
            <p> Material examined.   HOLOTYPE. “Kartabo P., Br. Guiana,  Aemerson [col.]”; “ Kartabo, Bartica District, British Guiana, 22.VIII.1920 ”; “WMMann [W. M. Mann], cooln, 1940”; “Type No., 64133, U.S.N.M.”; “USNMENT 01374929”  .  PARATYPE. “Kartabo, British Guiana, 15:X:1920”; “ Coll. By A. Emerson #424”; “FMNHINS 4431876” . </p>
            <p> Remarks.  Ceratoxenus tricornis is characterized especially by the presence of the two long and narrow frontal horns anterior to and transpassing antennal insertions, presence of vertexal bristles, and a straight and uncarinated gena; pronotal impressions are also distinct, with the anterior half longer and just slightly narrower than posterior, and with anterior margin straight. </p>
            <p>Distribution. Guyana —Kartabo.</p>
            <p> Biology. The host of  C. tricornis recorded in Mann (1923) was  Nasutitermes guayanae (Holmgren, 1910) , but data in Emerson (1935) leaves little doubt if the host is actually the newly described species,  N. similis Emerson, 1935 . </p>
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	https://treatment.plazi.org/id/03B187855C43FFFDABACD72BFF4E6541	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
03B187855C45FFFEABACD1FBFF61635D.text	03B187855C45FFFEABACD1FBFF61635D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratoxenus akerata Moreno & Zilberman 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ceratoxenus akerata Moreno &amp; Zilberman sp. nov.</p>
            <p>(Figs. 3A–D, 6C, F)</p>
            <p>Diagnosis: The species is recognized by the less developed and widely spaced lateral projections at frons along with a acutely projected medial process between the lateral projections; slightly acute and elevated gena, forming a longitudinal carina behind eyes; pronotum with anterior impression about the same length, and noticeably narrower than posterior, and with anterior margin arcuate.</p>
            <p>Head. Gena slightly elevated behind eyes; head surface finely punctuate; lateral projections at frons less developed and more widely spaced; medial process acutely projected, not overtaking the level of frontal elevations (Fig. 3 A,C; Fig. 6 C,F). Long bristles absent on vertex. Antenna with antennomere 1 cylindrical, with almost same length of antennomeres 2–4 combined; 2–5 longer than wide, subequal in length; 6–8 shorter than previous, longer than wide, all subequal in length; 9–10 slightly shorter than previous subquadrate; 11 conical. Secondary maxilar palpomere slightly rounded and at apex. Postmentum occupying more than half of total head length, longer than wide, slightly longer than gula, entirely covered by long bristles.</p>
            <p>Thorax. Pronotum broader on posterior region; pronotum light-brown with lateral margins darker; anterior impression about the same length and noticeably narrower than posterior, and with anterior margin arcuated, bearing long bristles marginally; lateral margins parallel with long bristles sparsely distributed (Figs. 3C, 6F). Elytra elongate, wider on posterior region; light-brown, outer margin darker. Legs with pro- and mesocoxae transverse; metacoxa triangular. Tibia slightly longer than femur, covered with long and thick bristles along entire length; outer margins of tibia slightly convex, and inner margin straight; pro-, meso-, and metatarsi increasing in length.</p>
            <p>Abdomen. As described to genus. Female unknown.</p>
            <p> Material examined.   HOLOTYPE. Male. PANAMA. “Barro C’do I. [  Barro Colorado Island ], C. Z. [Canal Zone]”; “19:VIII:35, A. Emerson [col.], #1479”; “  Xenopelta cornuta ”; “ Seevers det.”; “  Ceratoxenus akerata Moreno &amp; Zilberman, 2024 ”; “HOLOTYPE”; “ MZSP 21237 ”. </p>
            <p> Etymology. The specific epithet is derived from Greek, the junction of “ἀ-” (a-, “without”), and “ κέρατα ” (kérata, horns). Thus,  “akerata ” means “without horns” in allusion to the less developed frontal projections in the species. Noun in apposition. </p>
            <p> Remarks.  Ceratoxenus akerata sp. nov. differs from  C. tricornis and  C. inflatus by the less developed lateral projections of head and pronotal impressions about the same length; from  C. tricornis (in parenthesis) by the presence of carinete gena (uncarinete) and medial process slightly acutely projected (strongly acutely projected); and from  C. inflatus (in parenthesis) by the smaller and widely spaced lateral projections at frons (larger and closely spaced), medial process acutely projected (rounded), and acutely projected carinate. gena (roundely projected). </p>
            <p>Biology. Host unknown.</p>
            <p> Distribution. Panama —Barro Colorado Island. This species marks a new record to the genus  Ceratoxenus for Panama. </p>
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	https://treatment.plazi.org/id/03B187855C45FFFEABACD1FBFF61635D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
03B187855C46FFFFABACD787FDA5609A.text	03B187855C46FFFFABACD787FDA5609A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratoxenus inflatus Moreno & Zilberman 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ceratoxenus inflatus Moreno &amp; Zilberman sp. nov.</p>
            <p>(Figs. 4, 5, 6B, E)</p>
            <p>Diagnosis: The species is recognized by the larger and closely spaced lateral projections at frons; by the rounded triangular medial projection, overtaking the level of frontal elevations; gena elevated, broadly rounded, forming a longitudinal carina behind eyes; pronotum with anterior impression shorter and noticebly narrower than posterior, and with anterior margin arcuated.</p>
            <p>Head. Gena somewhat smooth, broadly elevated behind eyes; head surface finely punctuate; lateral projections at frons larger and closely spaced; medial process roundely projected. Long bristles absent on vertex (Figs. 4C, 6B,E). Antenna with antennomere 1 cylindrical, with almost same length of antennomeres 2–4 combined; 2–4 longer than wide, subequal in length; 5–8 shorter than previous, slightly longer than wide, all subequal in length; 9– 10 slightly shorter than previous, subquadrate; 11 conical. Second maxilar palpomere broadly rounded and inflated at apex. Postmentum longer than wide occupying more than half of total head length,, slightly longer than gula, entirely covered by long bristles.</p>
            <p>Thorax. Pronotum slightly broader on posterior region; pronotum light-brown with lateral margins darker; anterior impression shorter and noticeably narrower than posterior, and with anterior margin strongly arcuate, bearing long bristles marginally; lateral margins parallel with long bristles sparsely distributed (Figs. 4C, 6E). Elytra elongate, wider on posterior region; light-brown, outer margin darker. Legs with pro- and mesocoxae transverse; metacoxa triangular. Tibia slightly longer than femur, covered with long and thick bristles along entire length; outer margins of tibia slightly convex, and inner margin straight; pro-, meso-, and metatarsi increasing in length.</p>
            <p>Abdomen. As described to genus. Male unknown.</p>
            <p> Material examined.   HOLOTYPE. Female. GUYANA. “TOPOTYPE”; “Kartabo, British Guiana, 13: III: 1924”; “Coll. by, A.Emerson, #24.50 [field number]”; “FMNHINS [  Field Museum of Natural History ] 4518561”. </p>
            <p> Etymology. The epithet is derived from Latin,  inflatus , essentially meaning “inflate”, in allusion to the bloated elevated gena, and to the conspicuously rounded lateral projections of head. Noun in masculine. </p>
            <p> Remarks. The specimen studied contains the Emerson’s label “24.50” dated “ 13.III.1924 ”, and seemingly several termitophiles were collected by him in this date in different nests of  Nasutitermes guyanae . The specimen here studied was not referenced in Emerson's synonyms paper, but he mentioned one specimen labeled as "24.61" as "  Xenopelta cornuta " from this same date and locatity. Probably, these two specimens belong to the same species (  C. inflatus ), and based on these data we deduce that the host of  C. inflatus is  Nasutitermes guyanae , as this species was the only termite with termitophiles Emerson (1935) recorded at that particular place and date. It is noteworthy that the author erroneous synonym would stand along with any likely output, since there is no possibility of any of the abovementioned specimens belong to the genus  Xenopelta . For comparison with other species, please see the other remarks above. </p>
            <p> Biology. Probably  Nasutitermes guayanae (see remarks section). </p>
            <p>Distribution. Guyana —Kartabo.</p>
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	https://treatment.plazi.org/id/03B187855C46FFFFABACD787FDA5609A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
03B187855C48FFF0ABACD442FCA86084.text	03B187855C48FFF0ABACD442FCA86084.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopelta Mann 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Xenopelta Mann, 1923</p>
            <p> Xenopelta Mann, 1923: 356 (original description, included species  X. cornuta Mann, 1923 ); Seevers 1957: 106 (redescription). </p>
            <p> Type species:  X. cornuta Mann, 1923 (designation by monotypy) </p>
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	https://treatment.plazi.org/id/03B187855C48FFF0ABACD442FCA86084	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
03B187855C4BFFF5ABACD3DFFDA56661.text	03B187855C4BFFF5ABACD3DFFDA56661.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopelta cornuta Mann 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Xenopelta cornuta Mann, 1923</p>
            <p>(Figs. 7–9)</p>
            <p> Xenopelta cornuta Mann, 1923: 358 (original description); Seevers 1957: 107 (redescription of a  Ceratoxenus specimen as  X. cornuta ). </p>
            <p>Diagnosis: Same as for genus with the inclusion of color data. Overall color dark-brown, outer margins of elytra darker, and sternites slightly lighter. White areas of membrane between sclerites.</p>
            <p>Head finely punctuate on surface; eyes occupying less than half of head length; vertexal bristles absent. Antennomeres longer than wide, antennomere 1 with about same length of 2–5 combined; antennomeres 2–7 decreasing in length; 8–10 subequal in length, subquadrate; last antennomere conical. Antennae covered with thick and sparse bristles on antennomeres 1–6; antennomeres 7–11 densely covered with short bristles. Mandibles toothless, broader at base, slightly constricted medially. Postmentum finely punctuate. Gula narrow and convergent in first half and slightly divergent along their entire length.</p>
            <p>Thorax. Pronotum with lateral margins parallel, each with three long bristles symmetrically distributed in longitudinal row; posterior margin broadly arcuate medially, with straight angles. Elytra elongate, wider on posterior region; light-brown, outer margin reddish-brown; punctuate on surface; lateral margins of elytra slightly rounded; anterior margin weakly prominent; distal margins slightly declivous and convergent; angles slightly rounded; elytral surface with two long bristles in longitudinal row; apices rounded. Pro-, meso-, and metatarsi increasing in length; tarsomeres bearing sparse bristles.</p>
            <p>Abdomen with tergites III–VI transverse, with anterior margin straight and heavily sclerotized; posterior margin bearing transversal rows of long bristles. Tergite VIII wider than long; lateral margins rounded; posterior margin weakly arcuate; bearing four long bristles on surface symmetrically distributed. Sternite VIII subquadrate, wider than long; with bristles symmetrically distributed on surface. Tergite IX subtriangular, with long bristles in a longitudinal row parallel to lateral margin. Tergite X with posterior margin rounded, slightly narrower than anterior margin. Male unknown.</p>
            <p> Material examined.   HOLOTYPE. Female. "  Kartabo , Bartica District, British Guiana, 13. X. 1920 "; "appendages mounted on balcam slides"; WMMann [W. M. Mann], colln., 1940"; "Type No. C 4158, U.S. N.M"; "USNMENT 01374928" (pinned and parts on slides)  .   PARATYPE. “  Kartabo , Bartica District, British Guiana, 11.X.1920 ”; “  Xenopelta cornuta , paratype Mann”; “FMNHINS [Field Museum of Natural History] 4431875”, pinned  . </p>
            <p> Remarks. Species of  Xenopelta can be separated from the other genera of Neotropical  Termitogastrina by the frons abruptly elevated between antennal insertions and the posterior region of pronotum strongly elevated. An undescribed species of  Ceratoxenus was erroneously identified as  Xenopelta cornuta by Emerson (1935) leading the author to synonymize  Ceratoxenus with  Xenopelta . For the comparison with the genus  Ceratoxenus , see description and remarks in the  Ceratoxenus section. </p>
            <p> Biology. The host species is  Nasutitermes guayanae . </p>
            <p>Distribution. Guyana —Kartabo.</p>
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	https://treatment.plazi.org/id/03B187855C4BFFF5ABACD3DFFDA56661	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno, Carlos;Zilberman, Bruno	Moreno, Carlos, Zilberman, Bruno (2024): Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini). Zootaxa 5474 (1): 35-49, DOI: 10.11646/zootaxa.5474.1.3, URL: http://dx.doi.org/10.11646/zootaxa.5474.1.3
