taxonID	type	description	language	source
03B087BF607F1F46FF3AFC15A91EF822.taxon	materials_examined	Material examined. 13, 1 Ƥ: (IEBR- 46), Vietnam, Ninh Binh Province, Cuc Phuong National Park, forest, September 2006, leg. Luu Van Hien; 13: (IEBR- 59), Vietnam, Ha Tinh Province, Huong Son District, Son Kim commune, secondary forest, leaves litter, 600 m a. s. l., 3 May 2004, leg. Nguyen Duc Anh; 13: (IEBR- 60), Vietnam, Ha Tinh Province, Huong Son District, Cau Treo, secondary forest, leaves litter, 800 m a. s. l., 30 April 2004, leg. Nguyen Duc Anh; 13, 1 Ƥ: (IEBR- 79), Vietnam, Phu Tho Province, Xuan Son National Park, Forest, 500 m, 12 – 15 June 2004, leg. Nguyen Thi Phuong Lien; 23, 4 ƤƤ (IEBR- 177), Vietnam, Nghe An Province, Pu Mat National Park, Khe Thoi, closed forest, near stream, 18 o 58 ' 17.4 N- 104 o 48 ' 20.9 E, 4 – 10 April 2011, leg. Nguyen Duc Anh; 4 ƤƤ, 1 juvenile (IEBR- 178), same data as sample IEBR- 177.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF607F1F46FF3AFC15A91EF822.taxon	discussion	Remarks. This species was described by Golovatch (1984) and so far is known only from Cuc Phuong National Park (Golovatch, 1984; Korsós & Golovatch, 1989). The new records of S. grandis have expanded its distribution in northern Vietnam. Sellanucheza hoffmani sp. nov. (Figs. 3 – 5. Map 1)	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF607F1F46FF3AFC15A91EF822.taxon	materials_examined	Material examined: HOLOTYPE: 13 (IEBR- 61 H), Vietnam, Kon Tum Province, Loxo Pass, approximately 80 km north of Kontum, secondary forest, 800 m a. s. l., 12 – 19 April 2004, leg. Nguyen Duc Anh. PARATYPES: 43, 2 ƤƤ, (IEBR- 61 P), same data as holotype. NON-TYPES: 23, (IEBR- 181), 5 ƤƤ, 1 juvenile (IEBR- 182), Vietnam, Quang Binh Province, Minh Hoa District, Thuong Hoa commune, Phong Nha-Ke Bang region, 10 – 22 August 2011, leg. Nguyen Duc Anh.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF607F1F46FF3AFC15A91EF822.taxon	etymology	Etymology. This species is dedicated to Prof. Dr. Richard L. Hoffman, a prominent specialist in Diplopoda, who also established genus Szechuanella in 1961, a senior synonym of Sellanucheza Enghoff et al., 2004. Diagnosis. The new species is close to its congener, S. grandis, in body form, but differs distinctly in gonopod characters, including a long, basal laminiform process of the postfemoral region, and a very long basal spine of the lamina medialis, and in having sternite V with two separated, small, setiferous cones between coxae 4.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF607F1F46FF3AFC15A91EF822.taxon	description	Description. Size: body ca. 52 mm (male), 59 mm (female) in length, width of midbody pro- and metazona 4.5 mm (male), 5.4 (female) and 5.3 mm (male), 6.4 mm (female), respectively. Holotype 52 mm long and width of pro- and metazona 4.5 mm and 5.4 mm, respectively. Coloration: castaneous brown in general, especially antenna, metaterga. Prozona, pleura and antennomeres 6 & 7 darker while other body parts including paraterga, sterna and legs yellowish (Figs. 3 A – D). Head: a bit smaller than collum, sparsely setose even on labrum. Frons slightly convex. Epicranial suture distinctly evident. Antenna slender, claviform, relatively long, ca. reaching to the end of body ring 4 laterally. Most antennomeres subequal in length, except for antennomeres 1 and 7 (Figs. 3 A – B). Collum: marginally broader than body rings 2 – 5, finely smooth without any granulation or rugosity, with traces of a row of 3 + 3 setae near front margin. Axial line obviously distinct. Paratergum modestly-developed, sub-semicircular. Body rings: submoniliform, body rings 2 – 5 slightly smaller than 6, parallel-sided on body rings 6 – 15, thereafter tapering gradually onward. Both metaterga and prozona smooth, shining. Metaterga without any traces of setae. Metatergal transverse sulcus present from body ring 2, however, well-developed from body ring 5, extending to base of paraterga, which are keel-like but poorly developed on body rings 5 – 19 while modestly developed on body rings 2 – 4. Ozopore formula normal. Pores located at lateral sides of paraterga, but visible dorsally. Axial line thin, but obviously distinct, running from head to the end of body. Stricture dividing pro- and metazona thin and narrow, striate (Figs. 3 C, 4 A – B). Sterna: sparsely setose, cross-impression moderate in terms of both longitudinal and transversal suture, without any modifications except two separated small setiferous cones between coxae 4. (Figs. 3 D, 4 C) Legs: slender, not very long, subequal in length with body height. Tarsal brushes present until body ring 10, then thin and missing onward, however, only well-developed on mostanterior body rings (Fig. 5 A). Legs without coxal swollen and adenostyle as well. Telson: Epiproct short, with two small caudolateral tubercles. Tip with four spinneret cones (Fig. 4 E). Paraproct subsemicircular with two setae on disk. Hypoproct subtriangular with two distolateral setae (Fig. 4 D). Gonopod: (Figs. 4 F – H, 5 B – D) Coxite (co) slender, short, about half of femorite in length, with densely setose part distoventrally. Telopodite subfalcate. Prefemur densely setose as usual, and separated from femorite by transversal sulcus laterally. Femorite (fe) long, marginally expanded distally with distal part slightly twisted ventrad, and ventrally curved simultaneously. No demarcation between femorite and postfemoral region (solenophore, sph). The latter modestly twisted ventromesad, with a long basal laminiform process (lp). Both lamina medialis (lm) and lamina lateralis (ll) well-developed. Solenophore with an additional long spine (sp) at base of lamina medialis. Seminal groove running entirely on mesal side, distodorsally, before entering flagiliform solenomere (sl) which is completely sheathed by solenophore.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF607F1F46FF3AFC15A91EF822.taxon	discussion	Remarks. Hoffman (1961) established a new genus Szechuanella with the type species S. tenebra from Szechuan Province, China. Unfortunately, this name had been pre-occupied by Szechuanella Lu, 1959 with type species S. szechuanensis Lu 1959 for trilobites from China (Shelley et al., 2000). Consequently, Enghoff et al. (2004) substituted a nomen novum, Sellanucheza. However, Özdikmen (2007) later proposed a new replacement, Cemsunguria, without any comments on the previous substitution of Sellanucheza. Thus, under provision of Article 23 of the International Code of Zoological Nomenclature, the name Cemsunguria becomes a junior synonym of Sellanucheza. This genus currently consists of 4 species including a new species here, and mostly distributed in southern China, and Vietnam so far (Map 1).	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF60781F4AFF3AFDE5AAABFA1B.taxon	materials_examined	Material examined. HOLOTYPE: 13 (IEBR- 128 H), Vietnam, Ninh Binh Province, Cuc Phuong National Park, forest, 15 September 2006, leg. Luu Van Hien. PARATYPES: 23, 3 ƤƤ (IEBR- 128 P), same data as holotype; 3 3, (IEBR- 129), Vietnam, Hai Phong Province, Cat Ba National Park, primary forest, 10 August 2006, leg. Nguyen Tri Tien.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF60781F4AFF3AFDE5AAABFA1B.taxon	etymology	Etymology. This species is dedicated to the late Dr. C. A. W. Jeekel, a prominent specialist in Diplopoda, who established the genus Tonkinosoma.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF60781F4AFF3AFDE5AAABFA1B.taxon	diagnosis	Diagnosis. This species differs from its congener Tonkinosoma flexipes Jeekel, 1953, in following characters: sternite V with two independent small lamina between coxae 4, and without any modification between coxae 5; gonopod femorite much broader and constricted distally; solenophore with both well-developed lamina medialis and lamina lateralis.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF60781F4AFF3AFDE5AAABFA1B.taxon	description	Description. Size: body 42.6 – 47.5 mm (male), 57 mm (female) in length, width of midbody pro- and metazona 4.2 – 4.3 mm (male), 4.8 (female) and 5.3 – 5.8 mm (male), 7.2 mm (female), respectively. Holotype 47.5 mm long and width of midbody pro- and metazoan 4.3 mm and 5.8 mm, respectively. Coloration: body in general reddish brown; legs, antenna and sterna yellowish brown; paraterga, marginally rear part of metaterga and pleura paler (Figs. 6 A – C). Head: a bit smaller than collum, modestly setose on labrum, but sparsely on area between antennal sockets. Frons slightly convex. Epicranical suture obviously distinct. Antenna short, claviform, ca. reaching to body ring 3 laterally. Antennomere 1 <7 <6 <2 = 5 (Fig. 6 A). Collum: slightly broader than body ring 2, subtrapeziform, surface shining, smooth, without any rugosity or wrinkles, and with traces of 2 rows of 4 + 4 and 3 + 3 setae in front margin and in middle, respectively. Paratergum subtriangular, poorly-developed. Axial line wanting. Body rings: Width of body ring 4 <3 <2 = 5, parallel-sided on body rings 5 – 16, thereafter tapering gradually posterior. Surface smooth, shining, with traces of a row of 3 + 3 setae before transverse sulcus, which starts on metatergum 5, however, very superficial and vague (Fig. 7 A). Paraterga modestly developed, set lower than metatergal surface (Fig. 6 A & C). Caudal corner acute, surpassing rear contour (Figs. 6 A & C, 7 A). Pleurosternal carinae like small, oblique keels, present until body ring 10, and missing on subsequent ones (Fig. 6 A & C). Axial line thin, visible. Stricture dividing pro- and metazona thin, narrow and striate. Sterna: modestly setose, cross-impression moderate in terms of both longitudinal and transverse suture, without any modification except two independent lamina between coxae 4 (Fig. 7 B). Legs: rather short, but slender, about 1.1 times (male) as long as body height. No adenostyle or any other modification. Tarsi without setose brushes. Telson: Epiproct short, broadly truncated with four spinnerets at tip (Fig. 7 D). Paraproct sub-semicircular with two setiferous knobs on disk. Hypoproct subtriangular with two distolateral setiferous knobs (Fig. 7 C). Gonopods: (Figs. 7 E – F, 8 A – C) relatively simple. Coxite subcylindrical, slender and as long as femorite. Prefemur subrectangular, densely setose as usual, set off from femorite by a transversal suture (su) laterally. Femorite (fe) stout, expending in middle with a smaller distal part, without any femoral processes. Postfemoral region (= solenophore, sph) separated from femorite by an oblique sulcus laterally, curved down ventrally, and strongly twisted ventromesad. Both lamina medialis (lm) and lamina lateralis (ll) present. The former with a triangular lobe (tl) at distal part. Tip of solenophore acute. Seminal groove running entirely on mesal side before entering flagiliform solenomere, which is almost sheathed by solenophore.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
03B087BF60781F4AFF3AFDE5AAABFA1B.taxon	discussion	Remarks. Jeekel (1953) proposed a monotypic genus Tonkinosoma for a new species Tonkinosoma flexipes Jeekel, 1953, from Mauson Mt. (Lang Son province). The generic diagnosis was given to its species as well. Hoffman (1961), when he created a new genus Szechuanella for a new species S. tenebra from China, discussed and compared both genera, Tonkinosoma and Szechuanella. He suggested that S. tenebra and T. flexipes are very similar in gonopod structure except for the degree of curvature of the solenophore, a presence of pleurosternal carinae in Tonkinosoma, and also in small detail of the solenophore structure. Later Hoffman (1963) placed a species Nedyopus variatus Attems, 1953 into his genus Szechuanella, and maintained his separation of the two genera Tonkinosoma and Szechuanella. However, based on fresh material, the two genera are obviously distinguished by a presence / absence of additional processes in the postfemoral region of the gonopod, femorite enlarged or not, and the presence of pleurosternal carinae as well. In brief, the genus Tonkinosoma Jeekel, 1953 currently comprises two species and is distribute in the north of Vietnam (Map 1). MAP 1. Distribution of all Tonkinosomatini species. 1: Wushan, Szechuan Province, China; 2: Sapa, Lao Cai province, Vietnam; 3: Mau Son Mt., Lang Son Province, Vietnam; 4: Xuan Son National Park (NP), Phu Tho Province, Vietnam; 5: Cat Ba Island, Hai Phong Province, Vietnam; 6: Cuc Phuong NP, Ninh Binh Province, Vietnam; 7: Pu Mat NP, Nghe An Province, Vietnam; 8: Huong Son, Ha Tinh Province, Vietnam; 9: Phong Nha – Ke Bang NP, Quang Binh Province; 10: Loxo pass, Kon Tum Province, Vietnam.	en	Nguyen, Anh D. (2011): A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam. Zootaxa 3036: 58-68, DOI: 10.5281/zenodo.278746
