taxonID	type	description	language	source
03B2B90D995ED71698D091A3C86BAFC9.taxon	materials_examined	Type species: Cepon elegans Giard & Bonnier, 1887, by subsequent designation of Markham, 1975. Synonym: Merocepon Richardson, 1910 Type species: Merocepon xanthi Richardson, 1910, by original designation.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995ED71698D091A3C86BAFC9.taxon	discussion	Remarks. The type species of Trapezicepon Bonnier, 1900, T. amicorum (Giard & Bonnier, 1888), is quite similar in many characters to species of Cancricepon, especially to C. castroi n. sp. The most obvious difference is that T. amicorum, which has been reported infesting Trapezia cymodoce (Herbst, 1801) (Trapeziidae) (Bonnier, 1900) and Actumnus tomentosus Dana, 1852 [= A. setifer (De Haan, 1835)] (Pilumnidae) (Stebbing, 1910), lacks middorsal projections on any pereomeres of the female. The barbula is similar in both species in that there are two falcate lateral projections on each side but differs in that T. amicorum has a broad, flat median region, whereas members of the genus Cancricepon have a median region with a pair of blunt triangular projections. The males of T. amicorum and Cancricepon spp. are nearly indistinguishable. It appears that these two genera are closely related and molecular data would be helpful in determining if their status as separate genera is warranted. A second species of Trapezicepon, T. domeciae Danforth, 1972, was incompletely described and figured but appears to be congeneric with T. amicorum and therefore also closely related to Cancricepon spp. However, the third described species in Trapezicepon, T. thalamitae Markham, 1985, is clearly not congeneric with T. amicorum nor closely related to any species of Trapezicepon or Cancricepon. This conclusion is based on numerous characters of the female and male of T. thalamitae including: female with large middorsal projections on pereomeres 6 and 7 in T. thalamitae vs. none in T. amicorum, male with head much narrower than first pereomere in T. thalamitae vs. subequal in width in T. amicorum, and male with medial dorsal fusion of pleomeres in T. thalamitae vs. pleomeres completely separated in T. amicorum. In fact, these three characters show that T. thalamitae belongs to Scyracepon Tattersall, 1905, a genus in which females of all species have large middorsal projections on at least pereomeres 6 and 7, males have a head much narrower than the first pereomere, and males have medial dorsal fusion of all pleomeres. We therefore formally transfer T. thalamitae to Scyracepon, as S. thalamitae (Markham, 1985) n. comb. An additional juvenile female found parasitizing an unidentified species of Lambrus Leach 1815 (= Parthenope Weber, 1795) was identified as Trapezicepon sp. by Nierstrasz & Brender à Brandis (1931). However, due to the specimen’s immaturity, its generic placement is unclear and it could belong to any one of a number of keponine genera, including Apocepon Nierstrasz & Brender à Brandis, 1931, whose type species, A. sibogae Nierstrasz & Brender à Brandis, 1931, contains the only other bopyrid specimen (also a female) known to parasitize a host in Parthenopidae.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995DD71098D0972AC9DDAC19.taxon	description	(Fig. 1)? Leidya distorta. — Hay & Shore, 1918: 440 [see Table 1] (not Leidya distorta (Leidy, 1855 )).	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995DD71098D0972AC9DDAC19.taxon	description	Portunicepon rissoi. — Shiino, 1934: 276 [table of characters]. Cancricepon choprai (sic). — Bourdon, 1971: 387 – 389 [mention]. Cancricepon (Grapsicepon) choprai (sic). — Bourdon, 1971: 389 [mention].	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995DD71098D0972AC9DDAC19.taxon	materials_examined	Material examined. Infesting Micropanope cf. sculptipes Stimpson, 1871 (3.8 mm CW, 2.2 mm CL) (UF 31733); mature dextral female, 1.1 mm (UF 42205), mature male, 1.0 mm (UF 42205), USA, Gulf of Mexico, Florida, 28 ° 26 ’ 56 ” N, 84 ° 40 ’ 37 ” W, north-northwest of St. Petersburg, south of Big Bend area, 200 ft depth (= 60.1 m), 24 May 2012, coll. G. Paulay, N. Evans, F. Michonneau. Additional mature male, 0.8 mm (UF 42205; prepared for SEM), same locality as other specimens.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995DD71098D0972AC9DDAC19.taxon	description	Description. Female (UF 42205) length 1.1 mm; maximum width (across pereomere 5) 0.8 mm; head length 0.4 mm; head width 0.6 mm; pleon length 0.5 mm. All segments of body distinct (Fig. 1 A). Head subovate, frontal margin slightly bilobate medially. Frontal lamina large and extending beyond sides and frontal margin of head. Eye pigment on extreme edge of head at junction with frontal lamina. Antennules and antennae with two articles each, terminally setose. Barbula with two slender falcate pointed lateral projections on each side, outer projections larger than inner ones, middle region smooth (Fig. 1 B). Maxilliped with prominent anterior segment, large palp fringed with setae, plectron triangular, short and blunt (Fig. 1 C). Pereon broadest across pereomere 5. Subcircular tergal projections prominent on pereomere 2 – 4, coxal plates rudimentary. Last three pereomeres with overlapping, posteriorly directed mid-dorsal projections (Fig. 1 D). Oostegites almost enclosing brood pouch, oostegite 1 (Fig. 1 E, F) with subcircular anterior article, almost equal in length to posterior article, internal ridge smooth without any tubercles, posterior article triangular. Pereopods subequal in structure, first two and last two pairs smaller than pairs 3 – 5 (Fig. 1 A). All pereopods with elongate ischia, triangular meri, and blunt dactyli. Pleon with six segments, first five pleomeres bearing 5 pairs of biramous pleopods and uniramous lateral plates, all margins slightly undulating, surfaces smooth (Fig. 1 D). Endopodites of pleopods much smaller than exopodites, globular. Terminal pleomere ending in uniramous uropods, similar to but longer than fifth pleopods, surface smooth, margins slightly undulating (Fig 1 A). Male (UF 42205) length 1.0 mm; maximum width (across pereomere 4) 0.3 mm; head length 0.1 mm; head width 0.2 mm. All body regions and segments distinct (Fig. 1 G). Head elliptical, distinctly separated from first pereomere (Fig. 1 G). Large, irregular dark eyes near posterolateral corners. Antennules and antennae visible beyond margins of head in dorsal view, of three and five articles each, respectively; both bearing setae on two distalmost articles (Fig. 1 H). Pereon with mid-ventral tubercles on all segments. Pereomeres 3 – 5 subequal in width, patches of pigmentation on pereomeres 2 – 7 and pleomeres 1 – 2 (Fig. 1 G). First two pereopods larger than other five, all with similar structure (Fig. 1 H, I); all meri and carpi with setae on ventral surface. Propodi of pereopods 2 – 7 distally produced into comb-like surface receiving tips of dactyli (Fig. 1 H) Pleon with 6 pleomeres, each narrower than preceding one. First five pleomeres with low, rounded pleopods, without mid-ventral tubercles. Pleomere 6 without uropods, posterolaterally extended into two rami, each tipped with setae.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995DD71098D0972AC9DDAC19.taxon	discussion	Remarks. These specimens are similar to those described by previous authors (Nierstrasz & Brender à Brandis, 1925; Markham, 1975), although smaller than those specimens for which measurements were given. Both males in the present material were identical in all characters except size. The species was previously known from another species of Micropanope Stimpson, 1871, but this is the first record from M. cf. sculptipes, a species previously unknown to bear any bopyrid. The specimens identified as C. choprae by An (2009) from China do not belong to this species and are described below as C. beibusinus n. sp.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995BD71298D0955ACE39A9A4.taxon	description	(Figs 2, 3) urn: lsid: zoobank. org: act: 66 A 94 A 51 - 8 CFC- 4479 - 985 D- 9 DCE 9 C 1 D 02 B 1	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995BD71298D0955ACE39A9A4.taxon	materials_examined	Material examined. Infesting female Trapezia septata Dana, 1852 (11 mm CW, 9 mm CL) (UF 22339); mature dextral holotype female (4.5 mm), mature allotype male (1.3 mm, prepared for SEM) (UF 42206), Australia, Western Australia, Ningaloo Reef, Wreck Zirv, reef front, 22 ° 36 ’ 30 ” S, 113 ° 37 ’ 30 ” E, in Pocillopora sp., 10 m, May 2009, coll. J. Cale, L. Plaisance. Infesting female Trapezia lutea Castro, 1997 (8.4 mm CW, 5.9 mm CL) (UF 39611); mature sinistral paratype female (3.7 mm), mature paratype male (1.6 mm) (UF 42207), Maldives, 0.3 km south of Magoodhoo Island, 10 – 30 m forereef, in Pocillopora sp., 03 ° 04 ’ 29 ” N, 72 ° 57 ’ 59 ” E, 10 May 2014, coll. J. Moore. Infesting male T. lutea (15.7 mm CW, 12.3 mm CL) (UF 27019); mature sinistral paratype female (7.2 mm), mature paratype male (1.9 mm) (UF 42208), Japan, Okinawa Prefecture, Okinawa Island, White Beach, 1 – 14 m, 26 ° 17 ’ 44 ” N, 127 ° 54 ’ 22 ” E, 17 July 2010, coll. N. Evans, F. Michonneau, G. Paulay, T. Naruse, Y. Ise. Infesting male Quadrella coronata Dana, 1852 (6.9 mm CW, 5.4 mm CL) (UF 4821); juvenile non-type female (3.2 mm), 2 non-type cryptoniscus larvae (1.4 mm each) (UF 42209), Papua New Guinea, 02 ° 22 ’ 06 ” S, 146 ° 17 ’ 19 ” E, 29 June 2003, coll. L. Kirkendale.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995BD71298D0955ACE39A9A4.taxon	description	Description. Female length 3.7 mm excluding uropods; maximum width (across pereomere 3) 3.7 mm; head length 1.1 mm, width 1.2 mm; length of uropods 1.1 mm (measurements based on holotype). All segments distinct; no pigmentation (Fig. 2 A, B). Head ovate, frontal lamina prominent, extending beyond both sides of head and frontal margin, notched medially (Fig. 2 A, B); eyes lacking. Antennules and antennae of three and five articles each, respectively, terminally setose; all articles with scales (Fig. 2 C). Barbula with 2 falcate pointed lateral projections on each side, outer projections larger than inner; median region with a pair of blunt triangular projections (Fig. 2 D). Maxilliped with rounded anterior segment and large tapered palp, inner margin fringed with setae, plectron triangular, short and blunt (Fig. 2 E). Pereon broadest across pereomere 3 (Fig 2 A, B). Coxal plates developed in first 4 pereomeres. Large tergal projections present on pereomeres 2 – 4, with projections on longer side of body larger than those on shorter side. Last three pereomeres with pointed mid-dorsal projections (Fig 2 A, B). Brood pouch completely covered by oostegites, highly vaulted. Oostegite 1 (Fig. 2 F, G) with two segments subequal in length, anterior segment ovate; internal ridge notched near lateral margin, with small tubercle; posterior margin fringed with setae, round posterolateral point extended laterally. Pereopods 1 (Fig. 2 H) and 5 – 7 all small, with blunt dactyli, pereopods 2 – 4 large, with sharp dactyli (Fig. 2 I). All pereopods with stout bases. Pleon of six segments, first five pleomeres with 5 pairs of biramous pleopods and uniramous lateral plates. All pleopods and lateral plates with digitate margins, surfaces covered with tubercles (Fig. 2 J). Uropods uniramous, similar to pleopods (Fig. 2 A, B, J). Allotype male length 1.8 mm; maximum width (across pereomeres 4 – 6) approximately 0.7 mm; head broader than long, length 0.2 mm, width 0.3 mm. All body regions and segments distinct (Fig. 3 A, B). Head elliptical, with rounded anterior margin; small, dark eyes near posterolateral corners (Fig 3 A). Antennules with three articles each, distal margin of all articles with setae (Fig. 3 C). Antennae with five articles each, second through fourth articles with single seta, distal segment with many setae (Fig. 3 C). Pereon with seven segments. Pereomeres 4 – 6 subequal in width, small patches of pigmentation present on some pereomeres and first pleomere (Fig. 3 A). All pereomeres with rounded mid-ventral tubercles (Fig. 3 B). First three pereopods larger than posterior four; dactyli of first three pereopods much larger than others (Fig. 3 D – F), propodi, carpi and meri with ridged pad-like scales and tufts of setae on ventral surfaces (Fig. 3 D – F). Pleon with six pleomeres, lateral margins rounded (Fig. 3 A, B). First five pleomeres with uniramous lateral digitiform pleopods, tips tapered, acute with terminal seta (Fig. 3 G), first pleomere with midventral tubercle (Fig. 3 B). Sixth pleomere (pleotelson) without uropods or posterolateral lobes, anal cone visible in ventral view (Fig. 3 B).	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995BD71298D0955ACE39A9A4.taxon	etymology	Etymology. This species is named after Dr. Peter Castro (Professor Emeritus, California State Polytechnic University, Pomona) in honor of his contributions to trapeziid taxonomy.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D995BD71298D0955ACE39A9A4.taxon	discussion	Remarks. This new species can be distinguished from its closest relative, C. savignyi, by several characters of the female: all pereomeres with asymmetrical dorsolateral bosses (at least those on pereomere 2 symmetrical in C. savignyi), maxilliped palp elongate and tapered (maxilliped palp stout and rounded at tip in C. savignyi), endopods of pleopods less than 1 / 3 as large as exopods (endopods of pleopods 1 / 3 to 1 / 2 as long as exopods in C. savignyi). Additional differences between C. castroi n. sp. and other species of Cancricepon are given in the key to species below. Differences between the males of the two species are difficult to determine, as the sole male described by Stebbing (1910) had unusual pleopod structures and may not have been fully mature. No bopyrids were previously known from either Trapezia septata or T. lutea. No species of Quadrella Dana, 1851, has ever previously been reported to bear bopyrids, although Q. coronata is the type host for Sacculina pilosella Van Kampen & Boschma, 1925 (Cirripedia: Rhizocephala) (Van Kampen & Boschma 1925). Australia, the Maldives, and Papua New Guinea are new localities for species of Cancricepon.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9959D71C98D090DDCEF6AECC.taxon	description	(Fig. 4)	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9959D71C98D090DDCEF6AECC.taxon	materials_examined	Material examined. Infesting Leptodius affinis (De Haan, 1835) (8.6 mm CW, 5.7 mm CL) (UF 29451); mature dextral female (damaged, ca. 3.0 mm excluding pleon), mature male (prepared for SEM, 0.95 mm) (UF 42216), Taiwan, Keelung area, near bridge to Chao-Jing Station, 0 - 2 ft (= 0 – 0.6 m) intertidal, 25 ° 08 ’ 31 ” N, 121 ° 48 ’ 11 ” E, 2 July 2011, coll. M. Bemis, M. Malay, J. Huang.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9959D71C98D090DDCEF6AECC.taxon	description	Description. Female total length 4.1 mm (including pleon and uropods), maximal width 2.5 mm across third pereomere, head length 0.8 mm (including frontal lamina), head width 1.0 mm. All body regions and segments distinct. No pigmentation (Fig. 4 A, B). Head subovate, wider than long, anterior margin slightly bilobate and posterior margin convex; large frontal lamina extending beyond both sides of head, anterior margin with two small notches; eyes absent (Fig. 4 A). Antennules of three articles each, with stout basis and second article, distal article minute; antennae smaller than antennules, also of three articles each, all small; antennules and antennae with setae on distal two articles, all articles with scales (Fig. 4 C). Barbula with two falcate projections on each side, outer projections larger than inner ones and highly curved; pair of triangular projections medially (Fig. 4 D). Maxilliped with two segments, anterior segment much larger than posterior one (Fig. 4 E); large and curving palp with terminal setae (Fig. 4 F). Pereon with distinct segments, broadest across pereomere 3 (Fig 4 A); all pereomeres except first with middorsal projections, but pereomeres 2 – 4 with small round projections, pereomeres 5 – 7 with posteriorly pointed projections (Fig. 4 A, J). First four pereomeres with rounded dorsolateral bosses and weakly developed tergal projections (Fig. 4 A, I). Brood pouch completely covered by oostegites (Fig. 4 B). First oostegite (Fig. 4 G, H) with two subequally long articles, posterior article with irregular margin, internal ridge almost smooth, bearing a few small low projections near outer margin. Fifth oostegite with terminal setae and rounded tubercles on surface, covered with scales (Fig. 4 L). Pereopods subequal in structure, large knob-like structure on each ischium; posterior pereopods larger (Fig. 4 I, K, L) Pleon of six distinct pleomeres, first five with long lateral plates and biramous pleopods; lateral plates and pleopods with digitate margins, surfaces with small tubercules (Fig. 4 L – P); endopodites less than half as large as associated exopodites. Sixth pleomere with uniramous uropods, margins digitate, longer than pleopod 5 (Fig. 4 Q). Male length 0.95 mm, maximal width across fourth pereomere 0.3 mm, head length 0.1 mm, head width 0.16 mm (Fig. 4 R). Head elliptical, anterior margin rounded, posterior margin truncate (Fig. 4 R); conspicuous dark eyes near posterolateral margin (Fig. 4 R). Head and first pereomere separated (Fig. 4 R). Antennules with two articles each, first article with two setae, second article with 6 – 8 setae. Antennae with three articles each, first two articles with one seta, third article with 8 setae. Pereon with pigment on pereomeres 2 – 7 and pleomeres 1 – 2 (Fig. 4 R). All pereopods subequal in structure, first four pereopods larger than last three pairs. Carpi and meri of pereopods with one seta on ventral surface, propodi produced into plates covered in scales for receiving tips of dactyli. All pereomeres with midventral tubercles. Pleon of six segments, first pleomere with midventral tubercle (Fig. 4 S). First five pleomeres with low, rounded pleopods (Fig. 4 S), sixth pleomere weakly produced into two rami, each terminated with tuft of setae (Fig. 4 S), anal cone present medially (Fig. 4 S).	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9959D71C98D090DDCEF6AECC.taxon	discussion	Remarks. An et al. (2012) described C. multituberosum as having all pereomeres with middorsal triangular projections; the female from Leptodius affinis lacks a middorsal projection on the first pereomere. The present female specimen also differs from C. multituberosum in having a maxilliped with palp, whereas the maxilliped of C. multituberosum lacks a palp. The male of C. multituberosum has midventral tubercles on the first two pleomeres, whereas the present male has one only on the first pleomere. In all other aspects the present material agrees with C. multituberosum and, in the absence of more material from this host, we tentatively assign them to C. multituberosum. Leptodius affinis, has not previously been reported as a host for bopyrids.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9957D71898D0902ECE25AF64.taxon	description	(Fig. 5) urn: lsid: zoobank. org: act: B 6 E 28 ACF-FA 65 - 429 A- 8944 - 076 E 9 FA 423 B 4	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9957D71898D0902ECE25AF64.taxon	materials_examined	Material examined. Infesting female Scalopidia spinosipes Stimpson, 1858, mature sinistral female holotype (6.8 mm) (MBMCAS CIEO 625301), mature male allotype (2.5 mm) (MBMCAS CIEO 625302), China, Beibu Gulf, Stn. 6253, 19 ° 00 ’ 00 ” N, 107 ° 30 ’ 00 ” E, 67 m, 10 Feb 1960, coll. Z. Fan; infesting female S. spinosipes, mature sinistral female paratype (5.6 mm) (MBMCAS CIEO 625601), mature male paratype (1.9 mm) (MBMCAS CIEO 625602), China, Beibu Gulf, Stn. 6256, 17 ° 30 ’ 00 ” N, 107 ° 30 ’ 00 ” E, 67.7 m, 13 July 1960, coll. Z. Fan.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9957D71898D0902ECE25AF64.taxon	description	Description. Holotype female length 6.8 mm, maximal width 5.0 mm across third pereomere, head length 1.7 mm (include frontal lamina), head width 1.8 mm. All body regions and segments distinct; no pigmentation (Fig. 5 A, B). Head subovate, anterior margin bilobate and posteriorly curved; prominent frontal lamina extending beyond both sides of head and frontal margin; eyes lacking. Antennules and antennae of three and four articles each, respectively, distal articles small; setae on most articles (Fig. 5 C). Maxilliped with large, broad palp, setae on inner margin (Fig. 5 D). Barbula with two large falcate projections laterally, outer projections larger than inner ones; pair of triangular projections medially (Fig. 5 E). Pereon with seven segments, broadest across pereomere 3 (Fig 5 A); first four pereomeres with round dorsolateral bosses, pereomeres 2 and 3 with tergal projections (Fig. 5 A). Brood pouch completely covered by oostegites (Fig. 5 B). Last three pereomeres with middorsal projections, on fifth and sixth pereomeres rounded and directed posteriorly, on seventh pereomere acute and directed anteriorly (Fig. 5 A). First oostegite (Fig. 5 F, G) with two articles of approximately equal length, anterior article subovate, posterior article strongly tapered distolaterally, internal ridge with one larger thin projection nearly extending to the posterior margin of posterior article, several smaller subacute projections medially. Pereopods subequal in structure, propodus small, posterior pereopods larger (Fig. 5 H, I). Pleon of six pleomeres, first two and last two with smooth dorsal surfaces, pereomeres 3, 4 medially indented and bilobed. First five pleomeres with uniramous lateral plates and biramous pleopods; lateral plates and pleopods with densely digitate margins and surfaces with small tubercules (Fig. 5 A, B, J); endopodites smaller than associated exopodites (Fig. 5 J). Sixth pleomere with uniramous uropods, similar in structure to exopodite of pleopod 5 (Fig. 5 A). Allotype male length 2.5 mm, maximal width across fourth pereomere 0.9 mm, head length 0.2 mm, head width 0.4 mm (Fig. 5 K, L). Head an oblong ellipse, anterior and posterior margins rounded, minute dark eyes near posterior edge (Fig. 5 K); head and first pereomere separated (Fig. 5 K). Antennules with three articles each, distal two articles with setae; antennae with four articles each, distal three articles covered with setae (Fig. 5 M). Pereon of seven segments, fourth pereomere widest. All pereomeres with round midventral tubercles (Fig. 5 L). Pereopods subequal in structure, median three pairs slightly larger than others. Carpi, meri and propodi of pereopods with scales on ventral surfaces (Fig. 5 N). Pleon of six segments, without midventral tubercles. First five pleomeres with lateral tuberculate pleopods (Fig. 5 L), sixth pleomere (pleotelson) with truncate posterior margin, lacking uropods or posterolateral lobes (Fig. 5 L).	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9957D71898D0902ECE25AF64.taxon	etymology	Etymology. The species name is derived from the type locality, the Beibu Gulf, whose name in Chinese is Běibù Wān or " Northern Bay " of the South China Sea. 1 Bourdon (1971) gave Côte d’Ivoire and Dahomey (= Benin) as the type locality for C. anagibbosus in different parts of the paper; Côte d’Ivoire is the correct type locality, based on the data labels associated with the type specimens (MNHN-IU- 2007 - 3702, MNHN-IU- 2007 - 3703). 2 Markham’s (1979) Bermuda record is highly speculative as he reported only a single specimen of P. herbstii with an empty branchial swelling.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
03B2B90D9957D71898D0902ECE25AF64.taxon	discussion	Remarks. The type specimens of this species were originally identified as C. choprae, despite the enormous geographical distance separating them from any previously identified specimens of that species in the western Atlantic. The Chinese specimens can be separated from C. choprae by several characters of the females, including head with frontal lamina extending posteriorly on the sides of the head approximately half way (extending nearly to the posterior margin of the head in C. choprae), pereomeres 2 and 3 with completely to partly distinct median region (no regional separation in C. choprae), seventh pereomere middorsal projection acute (rounded in C. choprae), and first oostegite internal ridge with numerous tapered projections (few low rounded projections in C. choprae). Males of the two species can also be separated in that those of C. beibusinus n. sp. have a truncate pleotelson that lacks posterolateral lobes and a prominent anal cone (lobes and anal cone present in C. choprae); additionally, it appears that the number of pleonal midventral tubercles is different between the species, as there are none in C. beibusinus n. sp., but Markham’s (1975) illustration of a male C. choprae shows what appears to be midventral tubercles on pleomeres 1 – 3, although these were not mentioned in his text. This new species can also be separated from all other species in the genus by the characters given in the key below.	en	An, Jianmei, Wang, Mingxiao, Boyko, Christopher B., Williams, Jason D. (2019): New hosts and localities for species of Cancricepon Giard & Bonnier, 1887 (Isopoda: Epicaridea: Bopyridae) with description of two new species and comments on the relationship between Cancricepon and Trapezicepon Bonnier, 1900. Zootaxa 4559 (1): 136-150, DOI: 10.11646/zootaxa.4559.1.5
