taxonID	type	description	language	source
03B287D4095AFFFCFED1FE1CFE5158EA.taxon	type_taxon	Type species. Henicops maculatus Newport, 1844; by original designation.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095AFFFCFED1FE1CFE5158EA.taxon	diagnosis	Diagnosis. Member of Lamyctes-Henicops Group with 29 – 51 antennal articles; maxillipede coxosternum subtrapezoidal, dental margin moderately wide, with 3 + 3 or 4 + 4 teeth (exceptionally up to 6 + 6), lacking pseudoporodont; mandibular aciculae abundant, arranged in two (inner and outer) rows; several laciniate or plumose setae amidst simple setae on coxal process of first maxilla; projections on (at least) tergites 9, 11 and 13; last distal spinose projection of tibia on leg 14; subdivision of basitarsus of legs 1 – 12 indicated by paired larger setae; tarsi of legs 13 and 14 divided into three or four tarsomeres; distitarsus of leg 15 divided into at least two tarsomeres; first genital sternite of male divided longitudinally into two sclerites; male and female gonopods abundantly setose, with numerous distally curved setae on male gonopod. Assigned species. Lamyctes brevilabiatus Ribaut, 1923; Henicops dentatus Pocock, 1901 a (= H. oligotarsus Attems, 1911, n. syn.); H. milledgei n. sp.; H. tropicanus n. sp.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095AFFFCFED1FE1CFE5158EA.taxon	discussion	Discussion. The genus Henicops was erected by Newport (1844), with his concept of Henicops corresponding to the modern concept of the family Henicopidae. Two species of Henicops were originally named, Henicops maculatus from Tasmania and H. emarginatus Newport, 1844, from New Zealand. The latter was subsequently transferred to Lamyctes Meinert, 1868, and is a senior subjective synonym of the type species, L. fulvicornis Meinert, 1868 (Eason, 1996). In erecting a new genus, Paralamyctes, Pocock (1901 b) rediagnosed Henicops with emphasis on the division of the tarsi into tarsomeres. Henicops was again rediagnosed by Archey (1917), who included in that diagnosis the presence of plumose setae on the coxal process of the first maxilla, a character previously used to distinguish Lithobiidae from Henicopidae (Chamberlin, 1912). The “ plumose ” setae on the coxal process of the first maxilla in Henicops vary from laciniate (Fig. 15 J; Edgecombe et al., 2002: fig. 8 H for H. maculatus) to plumose, though the latter differ in detail from the complex plumose setae of lithobiids (Edgecombe & Hollington, 2002: fig. 9 C, E). The present study frames the generic diagnosis around synapomorphies that identify Henicops as a monophyletic group within a clade that also includes Lamyctes (= Lamyctinus Silvestri, 1909 a; see Edgecombe & Giribet, 2003 a), Lamyctopristus Attems, 1928, Analamyctes Chamberlin, 1955, Easonobius Edgecombe, 2003 a, and apparently Pleotarsobius Attems, 1909. The monophyly of this Lamyctes-Henicops Group is strongly supported by sequence data from multiple molecular markers (Edgecombe et al., 2002; Edgecombe & Giribet, 2003 b), and notably by insertions in the 18 S rRNA. Henicops is united with the South African Lamyctopristus Attems, 1928 (Edgecombe, 2004 a: fig. 34 D) by the mandibular aciculae being divided into inner and outer rows (Figs. 5 F, 10 C). This two-row arrangement is associated with a larger number of aciculae (about 20) in Henicops than in allied henicopines. Previous diagnoses (Archey, 1917, 1937; Attems, 1928) cited three joints in legs 1 – 13 as diagnostic of Henicops, but this is now observed to pertain to most but not all species (at least two undescribed species have only two tarsomeres on legs 1 – 12). A more general characterization of the Henicops tarsal condition is that subdivision of the basitarsus of legs 1 – 12 is indicated by paired larger setae. Where the tarsus has only two tarsomeres, a pair of strong, divergent setae is situated on the basitarsus in the same position as those larger setae immediately proximal to the most proximal tarsomere articulation in species with three tarsomeres, i. e., the tripartite arrangement on the tarsi of legs 1 – 12 in H. maculatus and the three species described herein occurs by subdivision of the basitarsus. Concerning the distitarsus, all species of Henicops uniquely share a division of the distitarsus of leg 15 into tarsomeres, a state not otherwise developed in the Henicopini (see Table 1 for a comparison of tarsal segmentation between species). The bipartite first genital sternite in males (Figs. 6 I, 11 B, 16 F) is invariant in Henicops. Elsewhere in Henicopidae, the division of this sternite into two sclerites is observed only in some members of Lamyctes, in those mostly African species split off by Verhoeff (1941) as Metalamyctes. The abundance of setae on the gonopods in both sexes is added to the generic diagnosis here, the number of setae in Henicops substantially exceeding that in most other Henicopinae, including other members of the Lamyctes-Henicops Group. This abundance could be explicitly defined in terms of the male gonopod having about 15 setae on the basal article and about 10 on the second and third. Among these are many distally curved setae (Figs. 4 E, 12 C). Additional distinction from Lamyctes is provided by the presence of tergal projections, subdivision of the tarsi, and a wider dental margin of the maxillipede coxosternum. A species that presents incongruence is the New Caledonian Lamyctes brevilabiatus Ribaut, 1923. Sequences from nuclear ribosomal as well as mitochondrial markers, separately as well as in combination, indicate that this species is more closely related to Henicops than to Lamyctes (Edgecombe & Giribet, 2003 a, b). The molecular data more precisely indicate an alliance with members of the Henicops dentatus Group as defined below. Apomorphic morphological characters shared with Henicops include the relatively large number of antennal articles (38 – 47), a relatively wide dental margin of the maxillipede coxosternum with 3 + 3 teeth and no pseudoporodont, laciniate setae on the coxal process of the first maxilla, and a bipartite first genital sternite in the male (AM KS 82626, Mt Humboldt). The spines beneath the posterior pretarsal accessory claw are short, as in Henicops (Fig. 14 L), rather than long and needle-like as in Lamyctes (Edgecombe & Giribet, 2003 a: figs. 34, 36). The New Caledonian species lacks, however, several characters shared by all other members of Henicops. It has transverse (rather than projected) posterior margins of TT 11 and 13, the last distal spinose projection is on the tibia of leg 13 (rather than 14), and the tarsi of legs 13 – 15 are bipartite, with no trace of a subdivision of either tarsomere. Legs 1 – 12 have indistinct tarsal articulations. The mandible differs from other members of Henicops in having relatively few (nine) aciculae in a single row (Fig. 2 B, C), rather than having about 20 aciculae in two rows. Cladistic analysis of morphological evidence resolves brevilabiatus as sister group of Henicops as diagnosed above in some minimal length cladograms, but as a member of Lamyctes in others, whereas combination of the morphological data with molecular data favours an ingroup position within Henicops (Edgecombe & Giribet, 2003 b). Based on strength of the molecular support together with the presence of several apomorphies of Henicops listed above, the species is reassigned to Henicops.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095BFFFAFC20FF10FEC15817.taxon	diagnosis	Diagnosis. Henicops with 29 – 40 (most commonly 36) antennal articles, with a relatively gradual change in length of articles along antenna; Tömösváry organ deep, outer edge at margin of cephalic pleurite; TT 7, 9, 11 and 13 with subquadrate emargination (posterior margin with transverse medial sector with independent curvature from lateral sectors); three moderate sized teeth on dental margin of maxillipede coxosternum; groove in accessory denticle field lacking on mandibular teeth; mandibular aciculae variable in structure, differentiated into some with pronounced serrations on both margins and others with a simple margin along most or all of length; ventral branching bristles of mandible lacking spine-like branches on basal part; three tarsomeres in legs 1 – 12; four tarsomeres in legs 13 and 14, basitarsus and distitarsus with two tarsomeres each; five tarsomeres in leg 15, basitarsus having two, distitarsus three. Assigned species. Henicops dentatus Pocock, 1901 a (= H. oligotarsus Attems, 1911); H. tropicanus n. sp.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095BFFFAFC20FF10FEC15817.taxon	discussion	Discussion. The diagnosis above lists characters that unite Henicops dentatus and H. tropicanus n. sp. that are either unique within Henicops or permit distinction from H. maculatus or other species. Several of these can be regarded as synapomorphies for H. dentatus and H. tropicanus. Character polarity is evaluated in the context of Henicops being sister to a clade that includes Lamyctes, Analamyctes, Easonobius and Lamyctopristus (Edgecombe et al., 2002; Edgecombe, 2003 a, 2004 a; Edgecombe & Giribet, 2003 b) (see synoptic classification below). Indistinct grooves in the accessory denticle field on the mandibles (Figs. 5 D, I, 10 A, F) are restricted to the H. dentatus Group within Henicops. Grooves or grooved ridges (see Fig. 14 D, F for their presence in H. milledgei) are observed in most Henicopini, including other members of the Lamyctes-Henicops Group (Fig. 2 A). The “ notched ” margin of tergite 7 in the H. dentatus Group (Figs. 3, 7 A) is distinctive for those two species within Henicops, though the same shape of this tergite is observed in some other Henicopinae, e. g., within Paralamyctes and in Zygethobiini. Tergite shapes were noted by Pocock (1901 a) as subquadrately emarginate TT 7, 9, 11 and 13, serving as a distinguishing character of H. dentatus and H. tropicanus relative to H. maculatus and H. milledgei. A differentiation of the mandibular aciculae into an outer row with serrated margins and an inner row with simple or weakly scalloped margins (Figs. 5 G, 10 C, D) is shared with Henicops milledgei (Fig. 14 I). This differentiation is basally (the entire basal half of the bristle lacks spines in H. dentatus: Fig. 5 H), but branch evenly along their distal part (Figs. 10 E, 14 H). This is also apparently apomorphic relative to the evenly spine-covered bristles in H. maculatus (Edgecombe et al., 2002: fig. 5 C), which have spines on their bases. The latter state is possessed by other taxa in the Lamyctes-Henicops Group (for Lamyctes, see Edgecombe et al., 2002: fig. 6 D, Edgecombe & Giribet, 2003 a: fig. 27; Fig. 2 B herein for H. brevilabiatus) and other Henicopini generally. The relatively even length of the antennal articles in the H. dentatus Group is potentially apomorphic relative to the groups of shortened articles occurring in pairs in other congeners because other genera of the Lamyctes-Henicops Group (e. g., Lamyctes, Analamyctes, Easonobius, Lamyctopristus) have the paired, shortened articles. The deep, submarginally positioned Tömösváry organ in the H. dentatus Group (Figs. 6 L, 9 D – F) is unique within Henicops, but is shared by most non- Henicops Henicopini. The deep, submarginal organ is thus apparently plesiomorphic relative to the shallow organ situated medially on the cephalic pleurite in H. maculatus (Edgecombe et al., 2002: fig. 1 G) and H. milledgei (Fig. 15 D, E). This character suggests that the H. dentatus Group is possibly sister to all other species of Henicops.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095DFFF6FF68FC1CFECF587F.taxon	description	Figs. 3 – 6	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095DFFF6FF68FC1CFECF587F.taxon	diagnosis	Diagnosis. Member of Henicops dentatus Group with female gonopod having two small, bullet-shaped or distally truncated spurs; outer spur up to 50 % wider than inner spur; branching bristles on mandibular gnathal edge narrow in dorsal part of fringe; ventral bristles in fringe lacking spines on their basal third or more.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095DFFF6FF68FC1CFECF587F.taxon	materials_examined	Type material. SYNTYPES: BM 1893.7.4.17, ♀ (Fig. 3 B), BM 1893.7.4.18, 3, Perth, WA, H. W. J. Turner. Other material. Types of Henicops oligotarsus Attems, 1911: LECTOTYPE: WAM 13 / 7172 (Fig. 3 A), ♀, Brunswick Junction, WA (station 139 of Michaelsen & Hartmeyer, 1907), 33 ° 15 ' S 115 ° 50 ' E, W. Michaelsen & R. Hartmeyer, 7 Oct 1905, designated here. A lectotype is selected to fix the name to a single type locality; the chosen specimen is the most intact large individual from a locality represented by both sexes in the WAM and NMV. PARALECTOTYPES: WAM, 233, from type locality. NMV: 3907, 1 ♀, Boyanup (station 146); 3908, 1 ♀, 233, Brunswick (station 139); 3909, 1 ♀, Albany (station 165). ZIUM: 6 ♀♀, 233, Lion Mill (station 99); 1 ♀, 13, Mundaring Weir (station 101); 13, Bridgetown (station 144); 1 ♀, Beverly (station 156). Attems (1911) listed additional syntypes from Wooroloo, East Fremantle, Jarrahdale, and Gooseberry Hill that we have not located. WESTERN AUSTRALIA: WAM: 25 / 822, 1 ♀, Dandaragan, 30 ° 40 ' S 115 ° 42 ' E, L. Glauert, Sep 1925; 28 / 740, 13, Serpentine, 32 ° 22 ' S 115 ° 58 ' E, L. Glauert, Aug 1928; 14 / 987, 1 ♀, Denmark, 34 ° 57 ' S 117 ° 21 ' E, W. B. Alexander, May 1914; T 42434, 4 ♀♀, 13, Hepburn Heights, 31 ° 49 ' 02 " S 115 ° 46 ' 13 " E, MSH & JMW, 13 Jul – 25 Sep 1995; T 42435, 3 ♀♀, Warwick Open Space, 31 ° 50 ' 34 " S 115 ° 48 ' 50 " E, MSH & JMW, 25 Sep – 28 Nov 1995; T 42436, 7 ♀♀ (Figs. 4 F, 5 F – I, 6 H), 2 33 (Fig. 4 E), Warwick Open Space, 31 ° 50 ' 33 " S 115 ° 49 ' 00 " E, MSH & JMW, 25 Sep – 28 Nov 1995; T 42437, 1 ♀, Balannup Lake, Gosnells, E. G. Cockett, 24 Dec 1968; T 42438, 1 ♀, Bald Head, Albany, D. D. Giuliani, 10 May 1969; T 42439, 13, Bentley, B. Anderson, 8 Sep 1976; T 42440, 1 ♀, Boranup, G. M. Riley, 10 Sep 1965; T 42441, 13, Peppermint Forest, 5 mi N Busselton, R. J. McKay & R. W. George, 22 Oct 1969; T 42442, 1 ♀, T 42443, 1 ♀, Cape Freycinet, 34 ° 06 ' S 114 ° 59 ' E, T. F. Houston, 15 – 18 Nov 1986; T 42444, 13, Cape Naturaliste NP, catchment of Yallingup Brook, 33 ° 38 ' 50 " S 115 ° 02 ' 10 " E, J. Mitchell et al., 1 – 10 Dec 2000; T 42445, 1 ♀, Fossil Deposit SF, 4 mi W cape, G. H. Riley & G. Kendrick, 8 Sep 1969; T 42446, 1 ♀, Cocanarup Timber Reserve, 33 ° 38 ' S 119 ° 54 ' E, G. Harold, Nov 1993; T 42447, 13, Worsley Alumina Project, Collie off Fletcher Rd, D. Walford & M. Sawle, Jul 1980; T 42448, 1 ♀, Crowea, S. J. Curry, 31 Oct 1980; T 42449, 13, Dog Pool on Shannon River, 34 ° 46 ' S 116 ° 22 ' E, MSH & JMW, 27 – 30 Apr 1990; T 42450, 1 ♀, 3 km N Dog Pool, 34 ° 45 ' S 116 ° 13 ' E, MSH & JMW, 30 Apr 1990; T 42451, 4 ♀♀, 333, Dwellingup, J. D. Majer, 7 – 25 Nov 1975; T 42452, 1 ♀, R. G. C. Mineral Sands, 10 km S Eneabba, 29 ° 50 ' S 115 ° 15 ' E, L. P. McMillan & P. West, 22 Jul 1997; T 42456, 5 ♀♀, 13, Fitzgerald River NP, West Mt Barren, NE slope, 34 ° 13 ' S 119 ° 26 ' E, MSH & JMW, 28 May 1994; T 42457, 1 ♀, Fitzgerald River NP, Twertup, 34 ° 01 ' S 119 ° 20 ' E, MSH & JMW, 29 May 1994; T 42458, 13, “ Glenbourne ”, Old Ellens Brook Rd, 33 ° 53 ' S 115 ° 00 ' E, L. Marsh et al., 27 – 28 Oct 1996; T 42459, 13, “ Glenbourne ”, Spring Rd S Gracetown, 33 ° 53 ' S 115 ° 00 ' E, 28 – 30 Jun 1997, L. M. Marsh et al.; T 42460, 13, “ Glenbourne ”, S Gracetown, 33 ° 54 ' 32 " S 115 ° 00 ' 24 " E, L. Marsh et al., 30 Oct – 1 Nov 1999; T 42461, 13, Gleneagle, 32 ° 15 ' S 116 ° 10 ' E, J. A. Springett, 22 Oct 1971; T 42463, 1 ♀, 12.5 mi NE Katanning, E. J. Car and G. M. Riley, 28 Oct 1964; T 42464, 13, Leeman, R. P. McMillan, 31 Aug 1981; T 42465, 1 ♀, T 42466, 1 ♀, Ludlow Tuart Forest, 33 ° 34 ' S 115 ° 29 ' E, J. A. Springett, 31 Aug 1971; T 42467, 4 ♀♀, Margaret River, L. E. Koch, 22 – 24 Aug 1973; T 42468, 1 ♀, T 42469, 13, 6.5 km and 5.5 km NW Meelup, 33 ° 33 ' 00 " S 115 ° 01 ' 45 " E, G. A. Harold, 22 – 27 Oct 1985; T 42470, 1 3, Mt Cooke, near summit, 32 ° 25 ' S 116 ° 18 ' E, MSH, JMW & M. Peterson, 7 Aug 1990; T 42471, 13, same locality, MSH & JMW, 1 Oct 1990; T 42472, 1 ♀, same locality and collectors, 16 Dec 1990; T 42473, 2 ♀♀, same locality, 31 Jul 1991; T 42474, 1 ♀, same locality, 19 Sep 1991; T 42475, 1 ♀, 13, base of Mt Dale, W side, 32 ° 08 ' S 116 ° 18 ' E, JMW et al., 27 Sep 1998; T 42477, 13, T 42478, 1 ♀, T 42479, 13, T 42485, 13, c. 4 km NNE Mt Lesueur, 30 ° 08 ' S 115 ° 12 ' E, K. Gaull et al., 9 – 12 Jul 1989; T 42481, 1 3, 1.6 km N Mt Lesueur, 30 ° 10 ' S 115 ° 12 ' E, K. Gaull et al., 11 Jul 1989; T 42482, 1 ♀, c. 5 km NE Mt Lesueur, 30 ° 09 ' S 115 ° 15 ' E, K. Gaull et al., 9 Jul 1989; T 42486, 233, 7 km NE Mt Lesueur, 30 ° 07 ' S 115 ° 15 ' E, K. Gaull et al., 1989; T 42487, 1 ♀, 4.5 km E Mt Peron, 30 ° 06 ' S 115 ° 12 ' E, K. Gaull et al., 7 Jul 1989; T 42488, 13, North Tarin Rock Reserve, D. Kitchener et al., 17 – 27 May 1971; T 42489, 1 ♀, Pemberton, Allis Rd, 34 ° 30 ' S 116 ° 00 ' E, J. A. Springett, 9 Nov 1971; T 42490, 13, Bluff Knoll, Stirling Ranges, D. D. Giuliani, 13 May 1969; T 42491, 233, Stirling Range NP, Ellen Peak, 34 ° 21 ' 32 " S 118 ° 19 ' 45 " E, 1000 m, S. Barrett, 4 Apr 1995; T 42492, 433, Swan River Dist, 15 – 18 Jul 1924; T 42493, 1 ♀, The Cascades, 8 km SSW Pemberton, 34 ° 30 ' S, 116 ° 00 ' E, MSH & JMW, 3 May 1990; T 42494, 13, Torbay hill, boundary track, 35 ° 05 ' S 117 ° 37 ' E, B. Y. Main, 20 Feb – 6 Mar 1983; T 42495, 1 3, Torbay Head, 35 ° 05 ' S 117 ° 38 ' E, B. Y. Main, 9 Aug – 7 Sept 1985; T 42496, 1 ♀, Torbay Head, B. Y. Main, 20 Jun 1986; T 42497, 13, Torndirrup NP, Sharp Point Rd, 35 ° 06 ' 42 " S 117 ° 52 ' 55 " E, JMW, 14 Nov 1998; T 42498, 1 ♀, Two People Bay, G. Smith, 15 Sep 1976; T 42499, 1 ♀, Tutanning Reserve, Pingelly, E. Mercer, 28 Aug 1980; T 42500, 13, Walyunga NP, M. Archer & E. Jeffreys, 4 May 1969; T 42501, 1 ♀, Wanneroo, 33 ° 54 ' 32 " S 115 ° 00 ' 24 " E, R. P. McMillan, 18 Aug 1952; T 42502, 1 ♀, West Cape Howe NP, Lake William, 35 ° 04 ' S 117 ° 37 ' E, B. Y. Main, 26 Mar – 5 Apr 1985; T 42503, 1 ♀, West Cape Howe NP, West Cape Howe, 35 ° 08 ' S 117 ° 36 ' E, B. Y. Main, 15 – 24 Jan 1988; WAM, 1 ♀, 13, Rottnest Island, Mt Herschell, 31 ° 59 ' S 115 ° 31 ' E, L. Glauert, Sep 1927. AUSTRALIAN MUSEUM: KS 15269, 3 ♀♀, 233, Dombalup SF, Marri Rd, 34 ° 30 ' S 116 ° 00 ' E, M. R. Gray, 15 Jan 1979; KS 15361 (Fig. 4 A – D), c. 200 individuals, Treen Brook SF, 8 km W Pemberton, 34 ° 26 ' S 116 ° 04 ' E, M. R. Gray, 13 Feb. 1979; KS 35584, 1 ♀, 333, Walpole-Nornalup NP, 34 ° 57 ' S 116 ° 16 ' E, M. R. Gray, 15 Feb 1979; KS 35644, 1 ♀, 233, Pine Ck, 0.5 km from Quarty Rd, Nannup-Pemberton area, 34 ° 15 ' S 115 ° 50 ' E, M. R. Gray, 14 Feb 1979; KS 77589, 13, Pemberton Forest Park, Karri Bush Walk, 34 ° 27 ' S 116 ° 01 ' 30 " S E, G. D. Edgecombe & Z. Johanson, 23 Feb 2001; KS 83629, 1 ♀ (Fig. 6 G, M), KS 83630, 13 (Figs. 5 A – E, J – O, 6 A – F, I – L), Gloucester NP, crossing of Bibbulmun Track and Burma Rd, 34 ° 27 ' S 116 ° 03 ' E, G. D. Edgecombe & Z. Johanson, 23 Feb 2001. CALIFORNIA ACADEMY OF SCIENCES (leg. E. S. Ross & D. Cavagnaro): 2 ♀♀, 333, Darlington, 450 ft, 5 Sep 1962; 1 ♀, 20 mi S Borden, 200 m, 23 Sep 1962; 1 ♀, 25 mi NW Walpole, 100 m, 25 Sep 1962; 4 ♀♀, 233, Cape Naturaliste, 5 m, 27 Sep 1962; 13, 7 mi NE Busselton, 50 m, 28 Sep 1962.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095DFFF6FF68FC1CFECF587F.taxon	distribution	Distribution. Southwest Western Australia, north to Eneabba (29 ° 50 ' S 115 ° 15 ' E), east to Cocanarup (33 ° 38 ' S 119 ° 54 ' E) (Fig. 1), sclerophyll forest.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D4095DFFF6FF68FC1CFECF587F.taxon	discussion	Discussion. Henicops dentatus is identical to or within the range of variation of H. tropicanus, described in full below, in most respects except for the female gonopods and details of the mandibular gnathal edge. Distinction between the two is discussed under the latter species. The present study of a large sample of southwest Western Australian specimens of Henicops, including the syntypes of both H. dentatus and H. oligotarsus, corroborates Archey’s (1937) suspicion that the two named Western Australian species are synonymous. When naming H. oligotarsus, Attems made no mention of the previously named H. dentatus. Attems (1928: 63) later acknowledged H. dentatus in a key to Henicops species. He distinguished the two Western Australian species on the basis of tergites 7, 9, 11 and 13 of H. oligotarsus being “ toothed ” and the corresponding tergites of H. dentatus and H. maculatus being “ rounded ”, in addition to leg 15 of H. oligotarsus having five tarsomeres as opposed to six tarsomeres in H. dentatus and H. maculatus. Both of these alleged differences are unfounded. Description of a “ toothed ” condition of tergites in H. oligotarsus refers to the triangular projections of the posterior angles of TT 7, 9, 11 and 13. This is a more or less accurate description of the tergites of Attems’ specimens (Fig. 3 A), but it is identical to the condition of these tergites in the types of H. dentatus (Fig. 3 B). Indeed, no Henicops specimens from anywhere in the range of H. dentatus (= H. oligotarsus) have tergites 7, 9, 11 and 13 that could be described as “ rounded ”. Leg 15 of all Western Australian specimens has five tarsomeres (Fig. 4 D) so no specific distinction can be made based on some specimens having five and others having six. The types of H. dentatus lack legs 14 and 15 (and seemingly lacked these legs when Pocock described the species, for no mention was made of them), so Attems’ claim of a difference from H. oligotarsus has no basis. In the absence of any morphological characters to support the presence of multiple taxa in Western Australia, all specimens are recognized as a single species, using the older name, H. dentatus. The description of H. oligotarsus (Attems, 1911: 150) stated that the species has 33 – 35 antennal articles but among the syntypes are large specimens with 29 – 36 articles. Other collections extend the range to 38 articles, though as in H. tropicanus the most common number is 36.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	description	Figs. 7 – 11	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	etymology	Etymology. For the Wet Tropics of northeastern Queensland, where the species occurs.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	diagnosis	Diagnosis. Member of Henicops dentatus Group with female gonopod having two (exceptionally three) large, elongate, bullet-shaped spurs; legs 14 and 15 relatively sparsely setose, with few setae on tibia and basitarsus of leg 15.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	materials_examined	Type material. HOLOTYPE: ♀ QM S 39941 (Fig. 7 A), from head of Roots Creek, 12 km WNW Mossman, Queensland, 16 ° 24 ' 46 " S 145 ° 16 ' 03 " E, 1200 m, 28 Dec 1989 – 11 Jan 1990, ANZSES, pitfall. PARATYPES: 3 QM S 39941, from type locality, collection data as for holotype; S 39869, 3 ♀♀, S 39873, 13, S 39898, 2 ♀♀ (Figs. 8 A – D, 10 D – F, K, 11 C, H, J), 13 (Figs. 9 B – D, F – J, 10 A – C, G – H, J, M, 11 D, E, I), S 39934, 2 ♀♀, Mossman Bluff Track, 5 – 10 km W Mossman, GBM, G. Thompson & ANZSES, 20 Dec 1989 – 15 Jan 1990. Other material. QUEENSLAND: QM: S 22632, 1 ♀, 14.4 km N Wudjl Wudjl, 15 ° 52 ' S 145 ° 19 ' E, R. & S. Raven, P. & E. Lawless, 28 Nov 1992 – 18 Apr 1993; S 39861, 2 ♀♀, 13, Mt Dalrymple, 21 ° 03 ' S 148 ° 38 ' E, 1200 m, ANZSES, 21 Dec 1992 – 10 Jan 1993; S 39863, 1 3, Eungella, Pease’s Lkt, 21 ° 07 ' S 148 ° 31 ' E, 900 m, D. Cook & GBM, 17 Nov 1992 – Apr 1993; S 39866, 2 ♀♀, S 39899, Maalan Rd, 2 km S Palmerston Hwy, 17 ° 36 ' S 145 ° 42 ' E, 750 m, GBM & J. Hasenpusch, 10 Jan – 7 Mar 1995; S 39870, 1 ♀, Wongabel SF, 17 ° 19 ' 30 " S 145 ° 29 ' 45 " E, R. Raven, P. & E. Lawless & M. Shaw, 23 Jul – 26 Nov 1992; S 39872, 2 ♀♀, Longlands Gap SF, 17 ° 27 ' 30 " S 145 ° 28 ' 45 " E, R. Raven, P. & E. Lawless & M. Shaw, 23 Jul – 26 Nov 1992; S 39874, 1 ♀, 13, S 39875, Karnak-Devils Thumb, 8 – 12 km NW Mossman, 1100 m, ANZSES, 26 Dec 1989 – 15 Jan 1990; S 39884, 2 ♀♀, 433, Hughes Rd, Topaz, 17 ° 26 ' S 145 ° 42 ' E, 650 m, GBM & Breeden, Sep – Dec 1993; S 39930, 1 3, same locality, GBM, D. Cook & H. Jantezki, 6 Dec 1993 – 25 Feb 1994; S 39892, 4 ♀♀, Kjelberg Rd turnoff, 17 ° 32 ' S 145 ° 36 ' E, 850 m, GBM & J. Hasenpusch, 7 Mar – 15 May 1995; S 39961, 2 ♀♀, same locality, 25 Nov 1994, 10 Jan 1995; S 39896, 4 ♀♀, Kjelberg Rd, Mt Fisher, 17 ° 32 ' S 145 ° 33 ' E, 1000 m, J. Hasenpusch, 1 Dec 1993 – 25 Feb 1994; S 39902, 1 ♀, S 39962, 1 ♀, Big Tableland, 15 ° 43 ' S 145 ° 17 ' E, 740 m, ANZSES, 20 Dec 1990 – 8 Jan 1991; S 39906, 1 ♀, Eungella NP, Mt William, 21 ° 02 ' S 148 ° 36 ' E, 1200 m, GBM, 19 Apr 1979, rf; S 39924, 1 ♀, 13, Pandanus Ck, Cathu SF, 20 ° 48 ' S 148 ° 33 ' E, 80 m, GBM, 22 Apr 1979; S 39925, 433 (Fig. 7 C), 2.5 km S Mt Hartley, 15 ° 47 ' S 145 ° 19 ' E, L. Roberts, 8 Dec 1993 – 2 Feb 1994; S 39927, 2 ♀♀, 3 juveniles, Mt Fisher, 7 km SW Millaa Millaa, 17 ° 33 ' S 145 ° 33 ' E, 1000 m, GBM & D. K. Yeates, 3 May 1983, rf; S 39928, 3 ♀♀, 333, Westcott Rd, Topaz, 17 ° 25 ' S 145 ° 42 ' E, 680 m, GBM, Jul – Dec 1993; S 39940, 1 ♀, 2 km SE Mt Spurgeon via Mt Carbine, 16 ° 27 ' S 145 ° 12 ' E, 1100 m, GBM & G. Thompson, 20 Dec 1988, rf; S 39946, 13, 2 juveniles, Mt Bartle-Frere, W base, 17 ° 23 ' S 145 ° 46 ' E, 700 m, GBM & J. Hasenpusch, 25 Nov 1994 – 10 Jan 1995; S 39947, 2 ♀♀, 13, Lamins Hill, 17 ° 22 ' S 145 ° 42 ' E, 880 m, GBM & J. Hasenpusch, 1 Dec 1993 – 25 Feb 1994; S 39954, 1 ♀, Eungella NP, Broken R, 21 ° 10 ' S 148 ° 30 ' 30 " E, P. Lawless, R. Raven & M. Shaw, 10 Nov 1991 – 29 Jul 1992; S 39955, 1 ♀, same locality, 29 Jul – 4 Dec 1992; S 39956, 1 ♀, Malaan SF, 17 ° 35 ' 30 " S 145 ° 36 ' 45 " E, R. Raven, P. & E. Lawless & M. Shaw, 25 Jul – 26 Nov 1992; S 39967, 1 ♀, 1.5 km SE Mt William, 21 ° 02 ' S 148 ° 37 ' E, 1060 m, ANZSES, 21 Dec 1992 – 10 Jan 1993; S 39973, 1 ♀, Malaan SF, V. E. Davies & R. Raven, 20 – 24 Apr 1978. AUSTRALIAN MUSEUM: 13, Eungella NP, Broken River campsite, G. D. Edgecombe, 18 Apr 1998, rf; 1 ♀, KS 83631, 233 (Figs. 9 A, E, K – M, 10 I, L, 11 A, B, F, G, K), Eungella NP, Dalrymple Rd, 1.7 km and 2.9 km NE Snake Rd, G. D. Edgecombe, 21 Apr 1998, rf. AUSTRALIAN NATIONAL INSECT COLLECTION: 2 ♀♀, 9 km ENE Mt Tozer (ex. Berl. 1059, 1062), 12 ° 43 ' S 143 ° 17 ' E, T. Weir, 10 – 16 Jul 1986; 2 ♀♀, juveniles, larval stadia LII – IV, 3 km NE Mt Webb (ex. Berl. 692, 721), 15 ° 03 ' S 145 ° 09 ' E, T. Weir, 1 – 3 Oct 1980, 3 May 1981; 1 ♀, 14 km W by N Hope Vale Mission (ex. Berl. 729), 15 ° 16 ' S 144 ° 59 ' E, A. Calder & J. Feehan, 7 – 10 May 1981; 2 ♀♀, 333, Mt Windsor Tableland (ex. Berl. 490), 16 ° 18 ' S 145 ° 05 ' E, 850 m, RWT, 20 Mar 1975, rf; 1 ♀, Mt Tiptree (ex. Berl. 1006), 17 ° 03 ' S 145 ° 37 ' E, B. Halliday, 13 Jul 1984, rf; 1 ♀, 3.2 km N Atherton (ex. Berl. 275), 17 ° 14 ' S 145 ° 29 ' E, J. G. Brooks, 5 May 1970, rf; 2 ♀♀, Mulgrave R (ex. Berl. 316), 17 ° 15 ' S 145 ° 46 ' E, 75 m, RWT & J. Feehan, 19 Jun 1971, rf; 1 ♀, Barrine NP (ex. Berl. 486), 17 ° 16 ' S 145 ° 38 ' E, 760 m, RWT, 21 Mar 1975, rf; 1 ♀, Eacham NP (ex. Berl. 484), 17 ° 18 ' S 145 ° 37 ' E, RWT, 23 Mar 1975, rf; 1 ♀, 20 km S Ravenshoe (ex. Berl. 358), 17 ° 45 ' S 145 ° 32 ' E, 800 m, RWT & J. Feehan, 3 Jul 1971, rf; 1 ♀, Paluma (ex. Berl. 207), 19 ° 00 ' S 146 ° 12 ' E, 810 m, E. Britton, 18 Jan 1970, rf; 3 ♀♀, 433, c. 3 km S Eungella (ex. Berl. 487, 488), 21 ° 09 ' S 148 ° 29 ' E, 780 m, RWT, 26 Mar 1975, rf; 1 ♀, 16 km N Eungella (ex. Berl. 493), 21 ° 03 ' S 148 ° 35 ' E, RWT, 13 Sep 1975.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	distribution	Distribution. Queensland: throughout Wet Tropics, south to Eungella, north to Mt Tozer, Cape York Peninsula (Fig. 1), mostly in rainforest, 75 – 1200 m, mostly above 650 m.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	description	Description. Length up to 24 mm; width of head shield up to 2.85 mm. Colour (based on specimens in absolute ethanol): antenna yellow to pale orange with several dark pigment spots occasionally present on dorsal side of articles in proximal third of antenna; head shield yellow with brown mottled network; tergites pale yellow to pale orange with dark (sometimes tinged with purple) mottling concentrated in longitudinal median band and near margins; dark pigment spots often present near tergal margins; maxillipedes pale orange; sternites pale yellow to pale orange with some faint purple mottling around margins; prefemur to tibia pale to moderate yellow with purple mottling, tarsi a slightly deeper orange; genital sternite and gonopods yellow. Head shield smooth. Frontal margin with strong median notch; posterior margin transverse or weakly concave; border slightly wider posterolaterally than medially. Median furrow incised for about 1 / 6 length to transverse suture, shallow throughout. Antennal length 4.1 – 5.7 times width of head shield, usually extending back to tergite 5 (Fig. 7 A); 33 – 40 articles, majority of specimens examined with 36; basal two articles much larger than succeeding two; relatively gradual change in length of articles along rest of antenna (Fig. 10 G, H), articles 3 to 8 – 10 typically wider than long, gradually elongating along antenna with more distal articles up to twice as long as wide (Fig. 10 I); gradational increase in density of setae to about tenth article, then more or less constant. Ocellus moderately large, overhanging lateral margin of head (Fig. 9 A), usually translucent, sometimes whitish to dark purple, domed. Tömösváry organ small, deep, longitudinally elliptical, outer edge at anterolateral margin of cephalic pleurite (Fig. 9 D – F). Small transverse seta projects medially from pit in labral sidepiece (Fig. 9 C). Labral margin with rounded shoulder beside midpiece, with pronounced break in curvature where branching bristles overhang margin; fringe composed of up to 30 densely grouped bristles with many short, spinelike projections along their length, a few longer spines at distal end (Fig. 9 G). Maxillipede coxosternum trapezoidal to subsemicircular, dental margin broad, each half weakly convex (Fig. 9 H – K); median notch lacking; teeth moderately large, triangular projections, invariably 3 + 3, usually with a slightly wider space between the outer tooth and middle tooth (Fig. 9 K) but ranging from equidistant to space between outer and middle tooth nearly twice that between inner and middle tooth. Coxosternum bearing moderate number of large, scattered setae, usually with distinctly denser setation behind dental margin and anterolaterally (Fig. 9 I, K). Tarsungulum with long, slender pretarsal section, as much as twice length of tarsal section (Fig. 9 H, L). Setae on telopodite somewhat more abundant on inner part of femur, tibia and tarsal section of tarsungulum than on outer part, these setae moderately long. Mandible: Four paired teeth on right mandible, left with smaller flattened fifth tooth adjacent to furry pad (Fig. 10 A, B). Approximately 20 aciculae differentiated into two rows (Fig. 10 C, D); aciculae in dorsal part of outer row with pronounced serrations on both margins along distal half, those in ventral part of outer row with a simple margin or with weak scalloping distally; aciculae of inner row with simple margin (Fig. 10 C, D). Fringe of branching bristles skirts aciculae, with transition to rows of scales with multifurcating spine fringes adjacent to midpoint of the second most ventral tooth; scales gradually shortening dorsally (Fig. 10 F) to terminate before reaching furry pad (Fig. 10 K); ventralmost bristles in fringe narrow-based, lacking spine-like projections on basal sixth to eighth of bristle, spine-like projections abundant and even along remainder of bristle (Fig. 10 E); bristles branching into a few spines distally. Accessory denticle field without grooves between rows of denticles or at margin of denticle field (Fig. 10 A, F); largest accessory denticles bluntly conical, grading into smaller triangular denticles then small elongate rods near fringe of scale-like bristles; band of unsculpted cuticle on dorsalmost tooth separates accessory denticles from furry pad; furry pad with many elongate, simple bristles (Fig. 10 K), some bristles with bifid terminations. First maxilla: Minute wedge-shaped sternite (Fig. 10 L). Coxal process trapezoidal, with a few plumose setae and up to 10 simple setae on anteromedial edge (Fig. 10 J), this cluster slightly separated from a few simple setae on inner edge of coxal process; plumose setae with numerous short, spine-like branches along their distal third. Cluster of about six barb-like sensilla microtrichoidea on posterolateral part of coxal process (Fig. 10 J). Distal article of telopodite with two rows of about 15 plumose setae along inner margin; branches developed along distal half of these setae (Fig. 10 M); shorter simple setae on membranous strip alongside inner margin; main field of distal article of telopodite bearing a few setae anteriorly. Second maxilla: Sternite small, fused with coxae. Band of 8 – 11 short setae along anterior part of coxa. Tarsus bearing numerous simple setae on outer surface, cluster of many plumose setae on membranous patch on inner surface (Fig. 11 D, E); branches on plumose setae mostly confined to distal half. Claw small, composed of five digits, median digit long, thick, with shorter, slender digit between median and each outer digit (Fig. 11 E). Tergites weakly wrinkled. T 1 generally trapeziform, smaller than T 3, slightly narrower than head shield (Fig. 7 A), about 70 % width of widest tergite (T 8), posterior angles rounded, posterior margin transverse; lateral margins subparallel in T 3, slightly convex in T 5, posterior angles rounded in both, posterior margin faintly concave in TT 3 and 5; TT 1 and 3 bordered posteriorly; borders of TT 5 and 7 incomplete posteriorly; posterior margin of T 7 with transverse or convex medial sector, having independent curvature from lateral sectors, posterior angles gently triangular, rounded; lateral margins of TT 4 – 14 convex, all bordered laterally; posterior margin of T 8 weakly concave, posterior angle blunt, TT 10 and 12 posterior margins gently concave, posterior angles with obtuse, blunt corners; TT 6, 9, 11 and 13 embayed with transverse posteromedial third, posterior angle of T 6 rounded, T 9 blunt, TT 11 and 13 with sharp triangular projections; posterior margin of T 14 concave, with blunt posterior angles. Short to moderate, slender setae along lateral borders of all tergites, often with 3 – 5 longer, thicker setae concentrated on anterolateral border; often cluster of several short setae on posterior angles of tergites; few short, slender setae scattered sparsely over surface of most tergites. A few moderate to long setae on margins of sternites, 1 – 3 setae at anterolateral corner most prominent anteriorly in trunk; setae along posterolateral and posterior margins more prominent in posterior segments; posterior margin fringed by 6 – 10 setae on sternites 13 – 15; consistently strong pair of setae on anteromedial part of sternites. Legs: Strong, pointed distal projections with sclerotized tips on tibiae of legs 1 – 14. Prefemur of legs 1 – 13 with short to moderate setae scattered on anterior and posterior faces, some longer, thicker setae on ventral edge of prefemur, femur and tibia; longest setae on femur and tibia equal to (Fig. 8 B) or shorter than (Fig. 8 A) longest on prefemur; a few thickened setae encircling distal margin of femur; strong, pigmented seta at ventrodistal end of tibia on anterior face of legs 1 – 13; tarsus with combination of slender setae of fairly even length, and four pairs of thicker, divergent, pigmented setae on ventral side of legs 1 – 13 (Fig. 11 K), two of these just proximal to articulations between tarsomeres, one near midlength of distal tarsomere. Setae on legs 14 and 15 more uniformly slender and sparsely distributed than on preceding legs, especially sparse on tibia and basitarsus of leg 15. Three tarsomeres in legs 1 – 12 (Fig. 8 A), with divided basitarsus. Condyle at joint between basitarsus and distitarsus on legs 13 – 15 only. Four tarsomeres in legs 13 and 14, basitarsus and distitarsus with two parts each (Figs. 8 B, C, 11 I); proximal tarsomere nearly 70 % length of basitarsus and 55 – 60 % length of distitarsus on leg 14. Leg 15 with five tarsomeres, basitarsus having two, distitarsus three (Figs. 8 D, 11 H). Distitarsus 70 – 80 % length of basitarsus on leg 15; basitarsus 12 – 14 times longer than broad; proximal tarsomere 55 – 65 % length of basitarsus; three tarsomeres comprise about 45 – 50, 15 – 20 and 30 – 40 % (proximal to distal) length of distitarsus. Pretarsal claws (Fig. 11 F, G) as described below for H. milledgei. Coxal pores round or ovate (Fig. 7 B, C), commonly 5,6,6,6 / 5,6,6,6 in large males and females, males maximum 5,6,6,6 / 5,6,6,6, minimum 3,4,4,4 / 3,4,4,4; females maximum 7,7,7,7 / 7,7,7,7, minimum 3,4,5,5 / 3,4,5,5. Coxal pore field set in shallow groove, distal pore (s) partly concealed by anteroventral face of coxa. Anal pores large in both sexes. Female (Fig. 7 B): Sternite of segment 15 transverse or convex posteromedially. Tergites of first genital segment and telson usually well sclerotized. Sternite of first genital segment large, posterior margin concave between condyles of gonopods except for small median projection, posterior two-thirds fairly evenly scattered with setae of varied length; short setae concentrated in a band along posteromedial margin. First article of gonopod bearing two (three in one specimen) large, bullet-shaped spurs, inner spur slightly smaller than outer; spurs gently curved such that tip points up (Fig. 11 C, J); first and second articles of gonopod with mix of many short, moderate and long setae, up to 56 on first article, up to 37 on second; third article with up to 10 setae, mostly moderate, one or two long ones (Fig. 11 J). Claw large, undivided. Male (Fig. 7 C): Sternite of segment 15 rounded posteriorly. Sternite of first genital segment small, divided, with transverse posterior margin, bearing numerous moderately long setae, short setae concentrated in a band near posterior margin (Fig. 11 B). Gonopod of three articles and tapering, seta-like terminal process that bears many short spine-like projections (Fig. 11 A), the three articles each bearing numerous short to moderately long setae, typically about 15, 10 – 12 and 10 for first to third articles, respectively; extended penis approximately equal in length to first article of gonopod. Larval stadia (identified by correspondence to leg and limb-bud pairs in Lamyctes emarginatus: Andersson, 1979, 1984) (ANIC Berl. 1062): Stadium LII with 8 pairs of legs and two pairs of limb-buds; 14 antennal articles. Stadium LIII with 10 pairs of legs and two pairs of limb-buds; 25 antennal articles. Stadium LIII with 12 pairs of legs and three pairs of limb-buds; 25 antennal articles. Stadia LII – IV all with 3 + 3 teeth on dental margin of maxillipede coxosternum.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40951FFF0FE85FCC5FB21587F.taxon	discussion	Discussion. As discussed above under the Henicops dentatus Group, H. dentatus and H. tropicanus n. sp. share numerous characters distinguishing them from other species of Henicops, and they appear to be sister taxa. In spite of their geographic separation, the two species are remarkably similar. They can consistently and most readily be distinguished by the morphology of the female gonopod, in particular the size and shape of the spurs on the basal article. Henicops dentatus has two, small bullet-shaped (Fig. 6 G) or distally-truncated (Figs. 4 F, 6 H) spurs, with the outer spur as much as 40 % longer and 50 % wider than the inner spur. In contrast, the gonopods of H. tropicanus (Figs. 7 B, 11 C) bear spurs that are larger and more elongate (with the outer spur of H. tropicanus being approximately 40 % the length of the inner margin of the first article of the gonopod, compared to approximately 20 % in H. dentatus) and both spurs are of more nearly equal size. Though the shape of the spurs varies within H. dentatus, they are consistently shorter and stouter than in H. tropicanus. In contrast to the distinction made possible by the female gonopods, the male gonopods of the three species of Henicops treated herein (Figs. 4 C, 7 C, 12 C) provide no obvious characters for specific distinction. Two features of the mandibular gnathal edge permit distinction between Henicops tropicanus and H. dentatus. The Western Australian H. dentatus has a narrower fringe of branching bristles on the dorsal part of the mandible (Figs. 5 D, I versus 10 F), this including the entire row of scale-like bristles. As well, the branching bristles on the ventral part of the fringe have a longer non-spinose (basal) extent (Fig. 5 H versus 10 E). In both of these characters, the state in H. tropicanus is plesiomorphic, being shared with H. maculatus (Edgecombe et al., 2002: figs. 4 A, 5 C), H. milledgei (Fig. 14 H), H. brevilabiatus (Fig. 2 A, B, D) and outgroups. Specimens of Henicops tropicanus often have a narrower medial sector to “ n otched ” tergites 7, 9 and 11 than does H. dentatus (compare Fig. 3 with Fig. 7 A) but this difference is inconsistent. Setation of legs 14 and 15 often also serves to distinguish between the two species: H. tropicanus typically has fewer setae on legs 14 and 15 (Fig. 8 C, D) than H. dentatus (Figs. 4 C, D), most notably on the tibia and basitarsus of leg 15 (Fig. 11 H), and has gracile setae on the prefemur of leg 15 versus sometimes spine-like setae in H. dentatus. Henicops tropicanus normally has two spurs on the female gonopod, only one specimen (QM S 38899) having three spurs. Three-spur variants are also exceptional in other members of the Henicopinae. For example, specimens of Lamyctes mauriesi Demange, 1981, are recorded as normally having two conical spurs on each gonopod, but single specimens from Tenerife and Gran Canaria have two such spurs on one gonopod and three on the other (Eason & Enghoff, 1992).	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	description	Figs. 12 – 16	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	etymology	Etymology. For Graham Milledge, who collected this species throughout most of its range.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	materials_examined	Type material. HOLOTYPE: MV NOH- 2207, ♀ (Fig. 12 A), Strzelecki Ra, Tarra-Bulga NP, VIC, 0.5 km NNE Tarra Valley Picnic Area, 38 ° 26 ' 40 " S 146 ° 32 ' 30 " E, Nothofagus cunninghamii forest, G. A. Milledge, 14 Nov 1995 – 10 Jan 1996. P ARATYPES: NOH- 2208, 1 ♀, 333, from type locality, 10 Jan – 5 Mar 1996; NOH- 2209, 1 ♀, 13, type locality, 5 Mar – 7 Mar 1995; NOH- 2211, 1 ♀, 13, Tarra-Bulga NP, 0.2 km W Tarra Valley Picnic Area, 38 ° 27 ' S 146 ° 32 ' E, 14 Sep – 14 Nov 1995; NOH- 2212, 13, same locality, 14 Nov 1995 – 10 Jan 1996; NOH- 2213, 1 3, same locality, 14 Nov 1995 – 10 Jan 1996; NOH- 2214, 1 ♀, 333, same locality, 10 Jan – 5 Mar 1996; NOH- 2215, 13, same locality, 7 May – 16 Jul 1996; NOH- 2220, 1 ♀, 233, same locality, 5 Mar – 7 May 1996; NOH- 2217, 1 ♀, Tarra-Bulga NP, Bulga Picnic Area, 38 ° 25 ' 30 " S 146 ° 34 ' 20 " E, 14 Nov 1995 – 10 Jan 1996; NOH- 2218, 1 3, same locality, 10 Jan – 5 Mar 1996; NOH- 2219, 2 ♀♀, 13, same locality, 5 Mar – 7 May 1996. Other material. VICTORIA: MV: OTWAY RA, leg. G. A. Milledge: NOH- 1045, 2 ♀♀, 13, Philips Track, 0.5 km N Triplet Falls, 38 ° 40 ' S 143 ° 29 ' E, 6 Sep – 15 Nov 1994; NOH- 1046, 1 ♀, 13, same locality, 15 Nov 1994 – 31 Jan 1995; NOH- 1047, 3 ♀♀, same locality, 31 Jan – 11 Apr 1995; NOH- 1048, 13, same locality, 11 Apr – 14 Jun 1995; NOH- 1049, 1 ♀, same locality, 14 Jun – 29 Aug 1995; NOH- 1050, 2 ♀♀, 233, Young Creek Rd, 0.4 km NW Triplet Falls, 38 ° 40 ' S 143 ° 29 ' E, 6 Sep – 15 Nov 1994; NOH- 1052, 7 ♀♀, 333, same locality, 15 Nov 1994 – 31 Jan 1995; NOH- 1053, 1 ♀, same locality, 31 Jan – 11 Apr 1995; NOH- 1054, 3 ♀♀, same locality, 11 Apr – 14 Jun 1995; NOH- 1055, 3 ♀♀, 13, Young Creek Rd, 0.2 km NE Ciancio Ck Crossing, 38 ° 40 ' S 143 ° 29 ' E, 6 Sep – 15 Nov 1994; NOH- 1056, 1 ♀, same locality, 15 Nov 1994 – 31 Jan 1995; NOH- 1058, 2 ♀♀, 13, 31 Jan – 11 Apr 1995; NOH- 1059, 1 ♀ (Figs. 14 K, L, 15 F, 16 G – J), 2 33, same locality, 11 Apr – 4 Jun 1995; NOH- 1060, 1 3, Aire Crossing Tk, 0.5 km N Aire R. Crossing, 38 ° 40 ' S 143 ° 29 ' E, 6 Sep – 15 Nov 1994; NOH- 1061, 233, Beauchamp Falls, 38 ° 39 ' S 143 ° 36 ' E, 6 Sep – 15 Nov 1994; NOH- 1062, 7 ♀♀, 13, same locality, 15 Nov 1994 – 31 Jan 1995; MV NOH- 1063, 8 ♀♀, 13, same locality, 31 Jan – 11 Apr 1995. CENTRAL HIGHLANDS, leg. G. A. Milledge: NOH- 1778, 1 larval stadium LII, 2 larval stadia LIII, Cement Creek Reserve, 2.2 km ESE Mt Donna Buang, 37 ° 43 ' S 144 ° 42 ' 15 " E, 13 Dec 1994; NOH- 1779, 2 ♀♀, 13 (Figs. 14 B, C, H – J, 15 B, D, E, G, L, M, 16 A, D – F, K, L), same locality, 29 Nov 1994 – 20 Jan 1995; NOH- 1784, 3 ♀♀, same locality, 21 Jan – 7 Apr 1995; MV NOH- 1782, 4 ♀♀ (Fig. 12 B), 433, 1 larval stadium LII, Donna Buang Rd, 1 km SW Mt Donna Buang, 37 ° 43 ' S 144 ° 40 ' E, 29 Nov 1994 – 20 Jan 1995; NOH- 1783, 6 ♀♀, 13, same locality, 21 Jan – 7 Apr 1996; NOH- 1786, 2 ♀♀, 233, 1 larval stadium LII, Myrtle Gully Reserve, 3.4 km WSW Mt Donna Buang, 37 ° 43 ' S 144 ° 38 ' 30 " E, 29 Nov 1994 – 20 Jan 1995; NOH- 1787, 5 ♀♀, 333, same locality, 21 Jan – 7 Apr 1995; NOH- 1788, 1 ♀, Road 26, 0.2 km WNW Donna Buang Rd junct., 37 ° 43 ' S 144 ° 39 ' 30 " E, 16 Feb 1995; NOH- 1790, 1 ♀, Acheron Gap, 6 km NE Mt Donna Buang, 37 ° 40 ' 43 " S 144 ° 44 ' 20 " E, 20 Feb 1996; NOH- 1792, 233 (Fig. 12 C), same locality, 26 Oct – 28 Dec 1995; NOH- 1793, 2 ♀♀, same locality, 28 Dec 1995; NOH- 1794, 4 ♀♀, same locality, 21 Feb – 23 Apr 1996; NOH- 1795, 1 ♀, same locality, 23 Apr – 25 Jun 1996; NOH- 1796, 2 ♀♀, 233, 0.7 km N Acheron Gap, 7 km NE Mt Donna Buang, 37 ° 40 ' 17 " S 144 ° 44 ' 20 " E, 21 Feb – 23 Apr 1996; NOH- 1797, 4 33, same locality, 28 Dec 1995 – 21 Feb 1996; NOH- 1800, 1 larval stadium LIII, The Big Culvert, 2.5 km ENE Mt Observation, 37 ° 33 ' 36 " S 145 ° 52 ' 15 " E, 28 Dec 1995. STRZELECKI RA, leg. G. A. Milledge: NOH- 2222, 1 larval stadium LIII, Tarra-Bulga NP, Bulga Picnic Area, 38 ° 25 ' 30 " S 146 ° 34 ' 20 " E, 7 May – 16 Jul 1996; NOH- 2225, 1 ♀ (Fig. 13 A – D), 2 33, Gunyah-Toora Rd, 2 km SSW Gunyah Gunyah, 38 ° 32 ' 30 " S 146 ° 19 ' E, 14 Nov 1995 – 10 Jan 13, (C) leg 14, (D) leg 15. Scale 0.5 mm. 1996; NOH- 2227, 1 ♀, Jeeralang West Rd, 0.1 km N Binns Hill Junction, 38 ° 26 ' 30 " S 146 ° 29 ' E, 10 Jan – 5 Mar 1996. AUSTRALIAN MUSEUM: KS 35864, 1 ♀, Otway Ra, 38 ° 27 ' S 143 ° 58 ' E, A. Frazer, Aug 1979; KS 35869, 13, Otway Ra, Lavers Hill, 38 ° 41 ' S 143 ° 24 ' E, A. Frazer, 24 Feb 1979; KS 83632, 1 ♀, Otway NP, Sandy Ridge, Spur Road, 38 ° 46.12 ' S 143 ° 32.9 ' E, 278 m, G. Cassis et al., 9 Nov 2002; KS 83633, 2 ♀♀ (Figs. 14 A, D – G, 15 A, C, H, K, 16 B, C), Grampians NP, Wonderland Range, Silverband Rd, 1.2 km S Rosea Campground, 37 ° 10 ' 28 " S 142 ° 30 ' 29 " E, G. D. Edgecombe & Z. Johanson, 15 Feb 2000, fern gully.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	distribution	Distribution. Victoria: Otway Ranges, Strzelecki Ranges, Central Highlands, mostly from Nothofagus cunninghamii forest; Grampians, sclerophyll forest (Fig. 1).	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	diagnosis	Diagnosis. Henicops with 36 – 43 (most commonly 40 or 41) antennal articles. Uneven change in length of articles in proximal part of antenna, with short paired articles interspersed between longer articles; T 7 with rounded, concave margin; Tömösváry organ small and weak, centrally located on cephalic pleurite; groove in accessory denticle field pronounced on mandibular teeth; female gonopod with two moderately large, bullet-shaped spurs (shared with H. maculatus). Mandibular aciculae differentiated into outer row with serrated margins and inner row with mostly simple margins; branching bristles of mandible with sparse spinelike branches on basal part of bristle (shared with H. dentatus Group). Uniquely: 3 + 3 moderate sized teeth on maxillipede coxosternum, with space between outer and middle teeth invariably more than twice the space between inner two teeth; three tarsomeres in leg 13, four in leg 14, five in leg 15, basitarsus undivided in legs 13 – 15; proximal coxal pores elliptical, distal pores round.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	description	Description. Length up to 28 mm; width of head shield up to 3.3 mm. Colour (based on specimens in absolute ethanol): antenna dark orange, with dark pigment spots usually present, especially on dorsal side and proximally but also common on ventral side of antenna and more distally; head shield dark orange with chestnut mottled network; tergites dark orange with dark mottling covering most of tergite, concentrated in longitudinal median band and near margins, and dark pigment spots concentrated on margins; maxillipedes orange; sternites pale orange with purple mottling around margins, becoming more concentrated in the three posterior sternites; prefemur to tibia pale to moderate yellow with purple mottling; tarsi pale to moderate yellow; genital sternite and gonopods orange with faint purple mottling on genital sternite. Head shield smooth. Frontal margin with strong median notch and short median furrow; posterior margin transverse; border very slightly wider posterolaterally than posteromedially. Antennal length 3.1 – 4.7 times width of head shield, usually extending back to tergite 5 (Fig. 12 A); 36 – 43 articles, commonly 40 or 41; basal two articles much larger than succeeding two (Fig. 14 A); antenna with a markedly uneven change in the length of articles in proximal part, with short paired articles interspersed between longer articles (Fig. 14 B); articles mostly substantially longer than wide in distal half of antenna; antenna densely setose along most of length, with many short, curved sensilla interspersed with long trichoid sensilla (Fig. 14 C). Ocellus moderately large, overhanging lateral margin of head, usually translucent, whitish to dark purple, domed. Tömösváry organ small, fairly shallow, longitudinally elliptical, situated inward of anterolateral margin of cephalic pleurite (Fig. 15 D, E). Clypeus with transverse band of six setae medially just in front of labrum (Fig. 15 B), cluster of 10 setae at clypeal apex (Fig. 15 A). Labral margin with rounded shoulder beside midpiece, with gradual break in curvature where fringe of branching bristles overhangs margin (Fig. 15 B); most bristles with short, spine-like projections along their length, some branching into long, slender spines distally (Fig. 15 C). Maxillipede coxosternum trapezoidal to subsemicircular, dental margin broad, each half gently convex (Fig. 16 A, B); median notch at most narrow, shallow (Fig. 16 D); teeth small, triangular projections, invariably 3 + 3, with space between outer tooth and adjacent tooth more than twice space between inner two teeth (Fig. 16 C – E), inner tooth smaller than the others. Coxosternum bearing moderate number of small to moderate sized setae (Fig. 16 D), mostly confined to anterior half of coxosternum (Fig. 16 A, B). Tarsungulum with long, slender pretarsal section (Fig. 16 A, B). Setae on telopodite denser on inner part of femur, tibia and tarsal section of tarsungulum than on outer part, especially long on tarsungulum (Fig. 16 C). Mandible as described for H. tropicanus except for the following: ventral aciculae in outer row of aciculae with both margins serrated along their distal fifth to third (Fig. 14 G, I); grooved ridge on accessory denticle field of all but ventralmost paired tooth (Fig. 14 D, F); ventral bristles in fringe of branching bristles lacking spine-like projections on basal fifth of bristles (Fig. 14 H); fringe of scale-like branching bristles extends to furry pad (Fig. 14 J); bristles of furry pad curved towards dorsalmost tooth (Fig. 14 F, J). First maxilla (Fig. 15 H – J) as described for H. tropicanus except: coxal process with a few short simple setae, several laciniate and plumose setae with intermediates between these (some with a few terminal spines, others with numerous short, slender projections along their distal halves) (Fig. 15 J); distal article of telopodite with several simple setae scattered over its main field (Fig. 15 I). Second maxilla (Fig. 15 K – M) as described for H. tropicanus. Tergites wrinkled, faintly so anteriorly but considerably so in posterior half of trunk. T 1 trapeziform, about as wide as head shield (Fig. 12 A), about 80 % width of widest tergite (T 8), posterior angles rounded and posterior margin gently concave; lateral borders subparallel in TT 3 and 5, posterior angles rounded, posterior margins gently concave; TT 1, 3 and 5 bordered posteriorly; border of T 7 incomplete posteriorly; posterior margin of T 7 evenly concave, posterior angle blunt; posterior margins of TT 8 moderately, fairly evenly concave, posterior angles obtuse, blunt; posterior margins of TT 6, 11 and 13 all evenly concave, increasingly so more posteriorly; T 9 variably with nearly transverse median sector; posterior angle of T 6 rounded, TT 9, 11 and 13 with triangular projections; posterior margin of T 14 moderately concave, with obtuse posterior angles; tergite of intermediate segment with weakly concave posterior margin in female, moderately concave margin in male. Tergal surface and margins scattered with short setae, in some specimens relatively numerous, e. g., posterior margins of TT 3 and 4 fringed by about 20 setae. Longitudinal median furrow confined to anterior part of sternites, not more than one-third length of sternite. Legs: Distal spinose projections on tibiae strong, pointed on legs 1 – 14. Setae relatively uniformly distributed along length of legs, with similar setal density on inner and outer faces of all podomeres; legs 14 and 15 (Fig. 13 C, D) with similar setal density as 1 – 13 (Fig. 13 A, B). Three tarsomeres in legs 1 – 12 (Fig. 13 A), with divided basitarsus; tarsal articulations discontinuous on dorsal side of leg in anterior part of trunk; three pairs of consistently strong, pigmented, divergent setae on tarsi of legs 1 – 12, one just proximal to each articulation between tarsomeres, one near midlength of distal tarsomere. Basitarsus-distitarsus joint with a condyle in legs 13 – 15 only, basitarsus undivided in these three legs; three tarsomeres in leg 13 (Figs. 13 B, 15 F); four tarsomeres in leg 14 (Fig. 13 C), distitarsus about 65 % length of basitarsus, three tarsomeres comprise about 45 – 55, 15 – 25 and 25 – 35 % (proximal to distal) length of distitarsus; five tarsomeres in leg 15, the proximal distitarsal podomere about 50 – 55 % length of distitarsus, second and third shorter than fourth (Fig. 13 D). Distitarsus 70 – 85 % length of basitarsus in leg 15; basitarsus 10 – 12 times longer than broad. Pretarsus with anterior and posterior accessory claws on all legs, about 30 % length of main claw, inserted on dorsolateral side of main claw, with gentle dorsoventral curvature (Fig. 14 K). Main claw gently curved, divided into many elongate, scales along most of its length, with short, polygonal scales ventrolaterally in region beneath proximal part of accessory claws, a few pores between these scales (Fig. 14 L), pore openings in the middle of small polygons; scales weakly defined proximally beneath bases of accessory claws and on proximoventral surface of main claw, well defined dorsoproximally. Posteroventral spine (' sensory spur' of Eason, 1964: fig. 486) present on legs 1 – 14 (Fig. 14 L), absent on leg 15 (Fig. 15 G), about 15 % length of main claw, directed distally, bearing slender subsidiary spine that parallels it on its dorsoproximal half (Fig. 14 L); accessory claws and posteroventral spine with surface ornament of linear grooves and ridges. Coxal pores round, transversely ovate, or elliptical, typically with round distal pores and elliptical proximal pores (Fig. 12 B); overwhelmingly 6,5,5,5 / 6,5,5,5 in large females (width head shield> 1.75 mm), 5,4,4,4 / 5,4,4,4 in large males; maximum in males 5,5,5,5 / 5,5,5,5. Coxal pore field set in shallow groove. Anal pores large in both sexes. Female (Fig. 12 B): Sternite of segment 15 transverse or convex posteromedially, with about a dozen setae along posterolateral and posteromedial margins. Tergites of first genital segment and telson usually well sclerotized; sternite of first genital segment large, posterior margin concave, with median bulge, between condyles of gonopods, surface scattered with setae, longest and densest posterolaterally (Fig. 16 G). First article of gonopod bearing two moderately large, bullet-shaped spurs, inner spur slightly smaller (Fig. 16 G – J); spurs gently curved such that tip points up (Fig. 16 I); first article usually bearing more than 30 setae in large specimens (up to 38), second article with 12 – 20 moderately long setae; third article with three to eight setae. Claw large, undivided, numerous pores with sensilla coeloconica on dorsodistal surface of claw (Fig. 16 J). Male (Fig. 12 C): Sternite of segment 15 with subtransverse (Fig. 12 C) to moderately convex (Fig. 16 F) posteromedial margin. Sternite of first genital segment small, divided, with transverse posterior margin, bearing about 20 moderate to long setae on each half with slight to moderate concentration near posterior margin. Gonopod of three articles and tapering, seta-like terminal process (Fig. 16 K, L), with many short spine-like projections along its inner margin; articles of gonopod each typically bear 10 – 12 setae; many setae on gonopod and first genital sternite curved, at least distally. Larval stadia: Stadium LII with 8 pairs of legs and two pairs of limb-buds; 14 antennal articles, all elongate; 3 + 3 teeth on dental margin of maxillipede coxosternum, outer tooth more distant than inner pair; tarsi unjointed. Stadium LIII with 10 pairs of legs and two pairs of limb-buds; 25 antennal articles; maxillipede dentition as in stadium LII; single tarsal articulation faintly defined on many legs.	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40957FFEAFC5AFCCEFB6E5F51.taxon	discussion	Discussion. All specimens of Henicops milledgei share a large space between the outer and middle tooth of the maxillipede coxosternum (more than twice the distance between the inner and middle tooth), exceeded only by H. brevilabiatus. No notable geographic variation has been detected between populations of H. milledgei from different regions of Nothofagus forest in Victoria. Segmentation of the larval antenna of Henicops milledgei and H. tropicanus presents a character that may be autapomorphic for Henicops or a group within the genus. Larval stadium LIII in both species has a uniquely high number of antennal articles (25), a count retained in larval stadium LIV in H. tropicanus at least. In comparison, Lamyctes emarginatus (= fulvicornis) and L. coeculus have only 14 articles in stadium LIII, and Lithobiidae have 14 – 17 articles in this stadium (Andersson, 1979: table III). Of Lithobiomorpha for which larval ontogeny has been described, only Cermatobius longitarsis approaches Henicops in having 20 – 24 articles in stadium LIII (Murakami, 1960; Andersson, 1979).	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
03B287D40940FFE7FDF1FE71FC0F5F87.taxon	description	Buethobius translucens Williams & Hefner, 1928 (USA: Ohio)	en	Hollington, L. M., Edgecombe, G. D. (2004): Two New Species of the Henicopid Centipede Henicops (Chilopoda: Lithobiomorpha) from Queensland and Victoria, With Revision of Species from Western Australia and a Synoptic Classification of Henicopidae. Records of the Australian Museum 56: 1-28
