taxonID	type	description	language	source
03B287C3FF8BFF8CFF65C292FB40F164.taxon	description	Male genitalia. Pygofer short and broad, usually with attenuate apical lobe or long ventral process. Anal tube usually extended beyond pygofer posterior margin. Valve semicircular. Subgenital plate triangular. Style apophysis small, digitate, preapical lobe rounded, moderately developed. Connective Y-shaped. Aedeagal shaft gracile, long, gonopore apical or subapical. Female. 7 th sternite comparatively large.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF8BFF8CFF65C292FB40F164.taxon	discussion	Remarks. Linnavuori (1978) reviewed the African species of the genus, including 2 new combinations and 3 new species. Viraktamath (1981) proposed 2 new combinations and described 1 new species from the Indian subcontinent. Later, 1 new species was added to the genus by Viraktamath & Anantha Murthy (1999) from India. Subsequently, Dai et al. (2006) described a new species from China. Khatri & Webb (2010) reported 3 known species whereas Naveed & Zhang (2018) documented a total of 5 known species from Pakistan.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF88FF8FFF65C0A4FCBBF1D8.taxon	description	(Fig. 1)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF88FF8FFF65C0A4FCBBF1D8.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF88FF8FFF65C0A4FCBBF1D8.taxon	discussion	Remarks. This species is very similar to G. pugio but the crown is more acute and the frontoclypeus upper part is more strongly carinate than in any other species in the genus.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C2EAFAE9F3FE.taxon	description	(Fig. 2)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C2EAFAE9F3FE.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C2EAFAE9F3FE.taxon	discussion	Remarks. This species is similar to G. rahmani externally but can be differentiated by the bifurcate pygofer process and aedeagus with a pair of dorsally curved appendages. This species is also similar to G. pallidus in aedagal shape but differs in having the pygofer process branched and the aedeagus with short distal processes.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C053FAE4F100.taxon	description	(Fig. 3)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C053FAE4F100.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C053FAE4F100.taxon	discussion	Remarks. Khatri & Webb (2010) reported this species, originally described from Iran, from Pakistan. Males differ from others of the genus in lacking a pygofer process and having the anal tube relatively short.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C1F1FE3FF689.taxon	description	(Fig. 4 D, E)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C1F1FE3FF689.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C1F1FE3FF689.taxon	discussion	Remarks. This species also resembles G. pugio in external features and in having a retrorse dorsal spine on the pygofer process, but is much smaller in size and can readily be differentiated by the subapical position of the spine on the male pygofer process.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C77AFD0DF42B.taxon	description	(Fig. 5)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C77AFD0DF42B.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C77AFD0DF42B.taxon	discussion	Remarks. This species varies in the color pattern of the crown and is close to G. pallidus in external characters but can be easily differentiated by the male genitalia.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C41FFB17F5D5.taxon	description	(Fig. 6)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C41FFB17F5D5.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF89FF8EFF65C41FFB17F5D5.taxon	discussion	Remarks. G. kempi can be readily distinguished from other members of the genus by the longer crown with some partial longitudinal stripes and a larger pale irregular area preapically, the serrate male pygofer processs, the apically upturned subgenital plate, the denticulate style apex, the footlike apex of the aedeagus and the female 7 th sternum with lateral lobes surpassing the 8 th sternite. This species has no close relative in the genus.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF8FFF88FF65C4A7FB48F5D4.taxon	description	(Fig. 7)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF8FFF88FF65C4A7FB48F5D4.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF8FFF88FF65C4A7FB48F5D4.taxon	discussion	Remarks. This species is readily recognizable by the dorsal coloration and shape of female 7 th sternite. This species most closely resembles G. pugio in size but differs in having a less acute crown apex.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF83FF84FF65C4B2FD05F5FD.taxon	description	(Fig. 4 F, G)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF83FF84FF65C4B2FD05F5FD.taxon	materials_examined	Material examined. No specimen examined. Remarks. The male of this species is unknown.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C2EAFED9F3DC.taxon	description	(Fig. 8)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C2EAFED9F3DC.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C2EAFED9F3DC.taxon	discussion	Remarks. This species was previously known from Sudan and India. Khatri & Webb (2010) reported it from Pakistan. This species resembles G. minabicus Dlabola and G. genoicus but can be distinguished from them by the male genitalia.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C455FA8FF5E1.taxon	description	(Fig. 4 A – C)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C455FA8FF5E1.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF80FF87FF65C455FA8FF5E1.taxon	discussion	Remarks. This species can be distinguished from other members of the genus by male genitalia characters. It is possible that G. pugio is a synonym of G. niveimarginatus but the male of the latter species is unknown so far.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C2EAFDD8F3DC.taxon	description	(Fig. 9)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C2EAFDD8F3DC.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C2EAFDD8F3DC.taxon	discussion	Remarks. This species was described based on syntypes from Punjab, Lyallpur, Oct., 1939 (A. Rahman). Later Mahmood (1979) also recorded this species from Pakistan. This species resembles G. pugio but differs in lacking a retrorse spine on the pygofer process.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C793FAEAF4C0.taxon	description	(Fig. 10)	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C793FAEAF4C0.taxon	materials_examined	Material examined. No specimen examined.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF86FF65C793FAEAF4C0.taxon	discussion	Remarks. Naveed & Zhang (2018) recorded this species from Pakistan. It was previously known only in India. It resembles G. rahmani in coloration but can be differentiated easily by the male genitalia characters.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF83FF65C530FA75F2E6.taxon	description	(Figs 11 – 12) Measurement. Male: 5.5 mm. Coloration and morphology. Yellowish brown (Fig. 11 A – C). Crown dark brown with median longitudinal white stripe reaching tip of anterior margin, also with pair of lateral ochraceous stripes, pair of posterior yellowish spots present on each side of median white stripe. Eyes color tinged with red (Fig. 11 A, C). Face whitish yellow with brownish narrow band along anterior margin (Fig. 11 D). Pronotum with pale irregular spots, as well as with bright median stripe. Mesonotum and scutellum yellowish brown with pale spots (Fig. 11 A, C). Femora and tibiae with dark brown marks (Fig. 11 B). Crown narrower than pronotum, subconical, longer than 2 x width between eyes (Fig. 11 A, C). Anteclypeus with submedian constriction, apex extending beyond normal curve of genae (Fig. 11 D). Pronotum shorter than crown. Mesonotum and scutellum nearly as long as pronotum (Fig. 11 A, C). Male genitalia. Pygofer lobe broad basally while narrowed towards posterior margin, with posterior short setae covering membranous area, tuft of long microsetae present near posterior margin. Pygofer process long, directed postero-dorsally, surpassing pygofer lobe, with subapical posteroventrally directed spine near apex (Fig. 12 A, 12 B). Subgenital plate triangular, broad basally, tapered towards apex with with lateral uniseriate macrosetae. Style elongated, with small apophysis (Fig. 12 D). Connective Y-shaped with arms well separated, stem short (Fig. 12 E, F). Aedeagus broad at base with well-developed dorsal apodeme, gonopore apical, aedeagal shaft slightly narrowed at mid-length, widened near apex in dorsal view, sinuate and evenly tapered distally in lateral view (Fig. 12 E – G).	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF83FF65C530FA75F2E6.taxon	materials_examined	Material examined. Holotype: male, Pakistan: Punjab: Murree Hills, 33 ° 54 ′ 23.508 N, 73 ° 23 ′ 37.284 E, 2000 m, Sweeping hand net, 23 July 2018, coll. Bismillah Shah.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF83FF65C530FA75F2E6.taxon	etymology	Etymology. This species is named for Punjab province in Pakistan where the type specimen has been collected.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
03B287C3FF81FF83FF65C530FA75F2E6.taxon	discussion	Remarks. This species resembles G. furcatus in external characters but can readily be distinguished by the subgenital plate with lateral margins not incurved at mid-length and the aedeagus without dorsal apical processes.	en	Shah, Bismillah, Naveed, Hassan, Duan, Yani (2019): Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan. Zootaxa 4688 (1): 71-85, DOI: 10.11646/zootaxa.4688.1.3
