taxonID	type	description	language	source
03B287A1C136FFB938F24561FB5EFE5A.taxon	type_taxon	Type genus: Rovnodryinus gen. nov.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C136FFB938F24561FB5EFE5A.taxon	diagnosis	Diagnosis. Female (Figs 1 A, B): apterous (Figs 1 A, B); antennomeres 3 – 10 with ADOs; head hypognathous (Fig. 1 D), with ocelli situated in anterior half of face not very far from clypeus (Figs 1 C, D); occipital carina complete (Figs 1 A, C, D); mesosoma almost completely composed of fused segments (Figs 1 A, 2 C); in mesosoma, only propleuron mobile and articulated with rest of mesosoma (Fig. 2 C); protarsus chelate (Fig. 3 A); chela with rudimentary claw (Figs 3 B, 4 A); tibial spurs 1 / 0 / 2. Male: unknown.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C136FFB938F24561FB5EFE5A.taxon	distribution	Distribution. Only known from Rovno amber.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C136FFB938F24561FB5EFE5A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C136FFB938F24561FB5EFE5A.taxon	discussion	Remarks. The tribe Rovnodryinini is known only as fossil in amber. It is characterized by two lateral ocelli (Figs 1 C, D) located just posteriorly to the antennal toruli, especially not far from the clypeus. The ocelli are located on the anterior half of the face, on the border of the frons and vertex: just the vertex is very long, longer than that of a hornet. This position of the ocelli is absolutely unusual in the Hymenoptera. Usually, in fact, the ocelli are located in the posterior half of the face. Another anomaly is the number of the ocelli (two), whereas in Dryinidae there are no ocelli (as in females of Apodryininae) or three ocelli situated in the posterior half of the frons, as in all males and remaining subfamilies. Really, there is a small doubt about a third ocellus (median ocellus), because photos of the head in frontal view do not show clearly the anterior third of the frons between the antennal toruli (because an optical distortion caused by the amber). However, at the stereomicroscope, this area is more visible and does not show the median ocellus, in addition, a third ocellus located close to the clypeus appears improbable. Furthermore, the authors discussed a lot about the possibility that the supposed ocelli were really air bubbles, because they are covered by a milk-like substance (common on inclusions in succinite ambers), that conceals the ocelli. However, we excluded this option, because the ocelli are perfectly symmetrical and compatible and no other similar structures are present on the face. Unluckily, it is not possible to show the ocelli symmetry by a photo, because of an optical distortion caused by the amber. However, at the stereomicroscope the symmetry is distinctly visible.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.taxon	type_taxon	Type species: Rovnodryinus khomychi sp. nov.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.taxon	diagnosis	Diagnosis. Female (Figs 1 A, B): see above tribe diagnosis.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.taxon	distribution	Distribution. Only known from Rovno amber.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.taxon	etymology	Etymology. The term “ Rovnodryinus ” derives from Rovno- (the Ukrainian region where the amber piece has been found) + Dryinus, type genus of Dryinidae).	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	description	(Figs 1, 2, 3 A, 3 B, 4 A)	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	materials_examined	Material examined: Female holotype number L- 905 (deposited in SIZK). Specimen in Upper Eocene Rovno amber, collected in Varash district, Rovno region. Horizon: Priabonian (35 – 38 Ma).	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	diagnosis	Diagnosis. See above tribe diagnosis.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	etymology	Etymology: the new species is named after the owner of the amber piece, Mr. Nikolai R. Khomych.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	description	Description of female. Apterous (Figs 1 A, B); length 3.7 mm. Head black; antenna brown; mesosoma black; metasoma black-brown; legs brown. Dorsal surface of head, body and legs covered by white milk-like substance (Fig. 1 B). Antenna composed of 10 antennomeres (Fig. 2 A), filiform, much shorter than head + mesosoma + metasoma (44: 62), posteriorly reaching second segment of metasoma; antennomeres apparently covered with short setae; ADOs (Figs 2 B, D) present in antennomeres 3 – 10 (= flagellomeres 1 – 8); antennomeres in following proportions: 20: 8: 15: 14: 14: 11: 10: 9: 9: 11; antenna long, more than twice as long as head: 44: 27. Head hypognathous (Figs 1 D, 2 C), with sculpture of face and vertex not distinct, because covered with milk-like substance; mandible and clypeus not distinct; occipital carina complete (Figs 1 A, C, D); temple prominent, very long, much longer than eye (10: 8) (Figs 1 D, 2 C); eye normally bulging, apparently unsetose; anterior half of face with two lateral ocelli situated between eyes (Figs 1 B, C, D) and very far from the occipital carina; median ocellus absent; OPL much longer than TL (19: 10). Palpal formula 6 / 3. Mesosoma much longer than head (50: 27), much shorter than metasoma (50: 80), almost completely composed of fused segments (Figs 1 A, 2 C); in mesosoma, only propleuron mobile and articulated with rest of mesosoma (Figs 1 A, 1 D, 2 C). Dorsal surface of mesosoma not visible, because covered with milklike substance. Apparent pronotum (Fig. 1 B) distinct, though covered with milk-like substance, because separated from rest of mesosoma by transverse furrow dorsally easily visible among milk-like substance (Fig. 1 B), laterally distinct, because not covered with milk-like substance (Figs 1 A, 2 C). Lateral surface of mesosoma (mesopleuron + metapleuron) covered with dense setae, with sculpture not distinct. Mesoscutum, mesoscutellum, metanotum, disc of metapectal-propodeal complex and propodeal declivity not visible. Apparent metapectal-propodeal complex densely setose, visible only laterally. Proleg podomeres in following proportions: 19 (coxa): 15 (trochanter): 38 (femur): 41 (tibia): 22 (first tarsomere): 4 (second tarsomere): 5 (third tarsomere): 13 (fourth tarsomere): 20 (fifth tarsomere). Protarsus chelate (Fig. 3 A). Rudimentary claw present (Figs 3 B, 4 A). Chela (Figs 3 B, 4 A) composed of normal enlarged claw + protarsomere 5. Enlarged claw (Figs 3 B, 4 A) slightly shorter than protarsomere 5 (17: 20), with long row of setae on outer side and long row of lamellae on inner side, number of lamellae indistinct. Protarsomere 5 (Figs 3 B, 4 A) apparently with two rows of lamellae, shorter row composed of about six long lamellae; longer row composed of indistinct number of lamellae. Protrochanter much longer than maximum width (15: 5), with slender proximal stalk, broadened distally (Fig. 2 C). Protarsomere 3 produced into hook. Arolium about as long as rudimentary claw (Figs 3 B, 4 A). Mesoleg podomeres in following proportions: 13 (coxa): 7 (trochanter): 38 (femur): 42 (tibia): 24 (first tarsomere): 8 (second tarsomere): 7 (third tarsomere): 4 (fourth tarsomere): 7 (fifth tarsomere). Metaleg podomeres in following proportions: 15 (coxa): 10 (trochanter): 58 (femur): 64 (tibia): 30 (first tarsomere): 13 (second tarsomere): 10 (third tarsomere): 6 (fourth tarsomere): 8 (fifth tarsomere). Petiole not visible. Tibial spurs 1 / 0 / 2. Male. Unknown.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.taxon	discussion	Remarks. Because of the peculiar morphologic characters indicated in the above diagnosis, the tribe Rovnodryinini trib. nov. can be included in the subfamily Apodryininae. The keys to the subfamilies and tribes of Dryinidae females presented by Olmi 1984, 1987, Olmi & Guglielmino 2010 and Olmi et al. 2020 can be modified by replacing the first six couplets as follows:	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C131FFBD38F2457DFE61F8BF.taxon	type_taxon	Type genus: Apodryinus Olmi, 1984	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C131FFBD38F2457DFE61F8BF.taxon	diagnosis	Diagnosis. Female: apterous; antennomeres 3 – 10 or 4 – 10 with ADOs; head without ocelli; occipital carina complete or incomplete; mesosoma almost completely composed of fused segments; in mesosoma, only propleuron mobile and articulated with rest of mesosoma; protarsus chelate; chela with rudimentary claw; tibial spurs 1 / 1 / 1 or 1 / 0 / 1. Male (see descriptions and figures in Olmi 1984, 2007 and Olmi et al. 2019): fully winged; fore wing with three basal cells (C 2 c, R 2 c, 1 Cu 2 c) clearly enclosed by pigmented veins; maxillary palpi composed of five or six palpomeres; labial palpi composed of three palpomeres; epicnemium distinct; mesosternum distinct, not fused with mesopleura; mandible with 3 – 4 teeth (if quadridentate, with a rudimentary tooth); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C131FFBD38F2457DFE61F8BF.taxon	distribution	Distribution. Known from Australia, South America and Africa.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C131FFBD38F2457DFE61F8BF.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
