identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B287A1C136FFB938F24561FB5EFE5A.text	03B287A1C136FFB938F24561FB5EFE5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rovnodryinini Olmi & Guglielmino & Vasilenko & Perkovsky 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tribe  Rovnodryinini trib. nov.</p>
            <p> Type genus:  Rovnodryinus gen. nov.</p>
            <p>Diagnosis. Female (Figs 1A, B): apterous (Figs 1A, B); antennomeres 3–10 with ADOs; head hypognathous (Fig. 1D), with ocelli situated in anterior half of face not very far from clypeus (Figs 1C, D); occipital carina complete (Figs 1A, C, D); mesosoma almost completely composed of fused segments (Figs 1A, 2C); in mesosoma, only propleuron mobile and articulated with rest of mesosoma (Fig. 2C); protarsus chelate (Fig. 3A); chela with rudimentary claw (Figs 3B, 4A); tibial spurs 1/0/2. Male: unknown.</p>
            <p>Distribution. Only known from Rovno amber.</p>
            <p>Hosts. Unknown.</p>
            <p>World species. One species is known.</p>
            <p>World genera. One.</p>
            <p> Remarks. The tribe  Rovnodryinini is known only as fossil in amber. It is characterized by two lateral ocelli (Figs 1C, D) located just posteriorly to the antennal toruli, especially not far from the clypeus. The ocelli are located on the anterior half of the face, on the border of the frons and vertex: just the vertex is very long, longer than that of a hornet. This position of the ocelli is absolutely unusual in the  Hymenoptera . Usually, in fact, the ocelli are located in the posterior half of the face. Another anomaly is the number of the ocelli (two), whereas in  Dryinidae there are no ocelli (as in females of  Apodryininae ) or three ocelli situated in the posterior half of the frons, as in all males and remaining subfamilies. Really, there is a small doubt about a third ocellus (median ocellus), because photos of the head in frontal view do not show clearly the anterior third of the frons between the antennal toruli (because an optical distortion caused by the amber). However, at the stereomicroscope, this area is more visible and does not show the median ocellus, in addition, a third ocellus located close to the clypeus appears improbable. Furthermore, the authors discussed a lot about the possibility that the supposed ocelli were really air bubbles, because they are covered by a milk-like substance (common on inclusions in succinite ambers), that conceals the ocelli. However, we excluded this option, because the ocelli are perfectly symmetrical and compatible and no other similar structures are present on the face. Unluckily, it is not possible to show the ocelli symmetry by a photo, because of an optical distortion caused by the amber. However, at the stereomicroscope the symmetry is distinctly visible. </p>
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	https://treatment.plazi.org/id/03B287A1C136FFB938F24561FB5EFE5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Olmi, Massimo;Guglielmino, Adalgisa;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C135FFB938F24654FD67F8EC.text	03B287A1C135FFB938F24654FD67F8EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rovnodryinus Olmi & Guglielmino & Vasilenko & Perkovsky 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus †  Rovnodryinus gen. nov.</p>
            <p> Type species:  Rovnodryinus khomychi sp. nov.</p>
            <p>Diagnosis. Female (Figs 1A, B): see above tribe diagnosis.</p>
            <p>Distribution. Only known from Rovno amber.</p>
            <p>Hosts. Unknown.</p>
            <p>World species. One species is known.</p>
            <p> Etymology. The term “  Rovnodryinus ” derives from Rovno- (the Ukrainian region where the amber piece has been found) +  Dryinus , type genus of  Dryinidae ). </p>
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	https://treatment.plazi.org/id/03B287A1C135FFB938F24654FD67F8EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Olmi, Massimo;Guglielmino, Adalgisa;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C134FFBE38F240CDFB2CFCE0.text	03B287A1C134FFBE38F240CDFB2CFCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rovnodryinus khomychi Olmi & Guglielmino & Vasilenko & Perkovsky 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> †  Rovnodryinus khomychi sp. nov.</p>
            <p>(Figs 1, 2, 3A, 3B, 4A)</p>
            <p> Material examined:  Female holotype number L-905 (deposited in SIZK). Specimen in Upper Eocene Rovno amber, collected in Varash district, Rovno region. Horizon: Priabonian (35–38 Ma).</p>
            <p>Diagnosis. See above tribe diagnosis.</p>
            <p>Etymology: the new species is named after the owner of the amber piece, Mr. Nikolai R. Khomych.</p>
            <p>Description of female. Apterous (Figs 1A, B); length 3.7 mm. Head black; antenna brown; mesosoma black; metasoma black-brown; legs brown. Dorsal surface of head, body and legs covered by white milk-like substance (Fig. 1B). Antenna composed of 10 antennomeres (Fig. 2A), filiform, much shorter than head + mesosoma + metasoma (44:62), posteriorly reaching second segment of metasoma; antennomeres apparently covered with short setae; ADOs (Figs 2B, D) present in antennomeres 3–10 (= flagellomeres 1–8); antennomeres in following proportions: 20:8:15:14:14:11:10:9:9:11; antenna long, more than twice as long as head: 44:27. Head hypognathous (Figs 1D, 2C), with sculpture of face and vertex not distinct, because covered with milk-like substance; mandible and clypeus not distinct; occipital carina complete (Figs 1A, C, D); temple prominent, very long, much longer than eye (10:8) (Figs 1D, 2C); eye normally bulging, apparently unsetose; anterior half of face with two lateral ocelli situated between eyes (Figs 1B, C, D) and very far from the occipital carina; median ocellus absent; OPL much longer than TL (19:10). Palpal formula 6/3. Mesosoma much longer than head (50:27), much shorter than metasoma (50:80), almost completely composed of fused segments (Figs 1A, 2C); in mesosoma, only propleuron mobile and articulated with rest of mesosoma (Figs 1A, 1D, 2C). Dorsal surface of mesosoma not visible, because covered with milklike substance. Apparent pronotum (Fig. 1B) distinct, though covered with milk-like substance, because separated from rest of mesosoma by transverse furrow dorsally easily visible among milk-like substance (Fig. 1B), laterally distinct, because not covered with milk-like substance (Figs 1A, 2C). Lateral surface of mesosoma (mesopleuron + metapleuron) covered with dense setae, with sculpture not distinct. Mesoscutum, mesoscutellum, metanotum, disc of metapectal-propodeal complex and propodeal declivity not visible. Apparent metapectal-propodeal complex densely setose, visible only laterally. Proleg podomeres in following proportions: 19 (coxa): 15 (trochanter): 38 (femur): 41 (tibia): 22 (first tarsomere): 4 (second tarsomere): 5 (third tarsomere): 13 (fourth tarsomere): 20 (fifth tarsomere). Protarsus chelate (Fig. 3A). Rudimentary claw present (Figs 3B, 4A). Chela (Figs 3B, 4A) composed of normal enlarged claw + protarsomere 5. Enlarged claw (Figs 3B, 4A) slightly shorter than protarsomere 5 (17:20), with long row of setae on outer side and long row of lamellae on inner side, number of lamellae indistinct. Protarsomere 5 (Figs 3B, 4A) apparently with two rows of lamellae, shorter row composed of about six long lamellae; longer row composed of indistinct number of lamellae. Protrochanter much longer than maximum width (15:5), with slender proximal stalk, broadened distally (Fig. 2C). Protarsomere 3 produced into hook. Arolium about as long as rudimentary claw (Figs 3B, 4A). Mesoleg podomeres in following proportions: 13 (coxa): 7 (trochanter): 38 (femur): 42 (tibia): 24 (first tarsomere): 8 (second tarsomere): 7 (third tarsomere): 4 (fourth tarsomere): 7 (fifth tarsomere). Metaleg podomeres in following proportions: 15 (coxa): 10 (trochanter): 58 (femur): 64 (tibia): 30 (first tarsomere): 13 (second tarsomere): 10 (third tarsomere): 6 (fourth tarsomere): 8 (fifth tarsomere). Petiole not visible. Tibial spurs 1/0/2.</p>
            <p>Male. Unknown.</p>
            <p> Remarks. Because of the peculiar morphologic characters indicated in the above diagnosis, the tribe  Rovnodryinini trib. nov. can be included in the subfamily  Apodryininae . The keys to the subfamilies and tribes of  Dryinidae females presented by Olmi 1984, 1987, Olmi &amp; Guglielmino 2010 and Olmi et al. 2020 can be modified by replacing the first six couplets as follows: </p>
            <p>1 Protarsus not chelate (Fig. 13 in Olmi 1984)................................................................ 2</p>
            <p>- Protarsus chelate (Fig. 3A; Fig. 16 in Olmi 1984)............................................................ 4</p>
            <p>2 Pterostigma less broad, more than three times as long as broad (Fig. 3A in Olmi et al. 2020); forewing with three basal cells (costal, radial and first cubital) enclosed by pigmented veins (Fig. 3A in Olmi et al. 2020).................................................... 17. Archaeodryininae Olmi, Perkovsky, Martynova, Contarini, Bückle &amp; Guglielmino (extinct)</p>
            <p>- Pterostigma broader, less than three times as long as broad (Figs 3 B-D in Olmi et al. 2020); forewing with one (costal) or two (costal and radial) basal cells enclosed by pigmented veins (Figs 3 B-D in Olmi et al. 2020)........................... 3</p>
            <p> 3 Head hypognathous (Fig. 7 in Olmi &amp; Guglielmino 2010); face and part of clypeus without strong median raised carina (Fig. 5 in Olmi &amp; Guglielmino 2010), with or without normal frontal line............................ 1.  Aphelopinae Perkins</p>
            <p> - Head prognathous (Fig. 2 in Olmi &amp; Guglielmino 2010); face and part of clypeus with one strong median raised carina (Fig. 1 in Olmi &amp; Guglielmino 2010)............................................... 2.  Erwiniinae Olmi &amp; Guglielmino </p>
            <p>4 Ocelli absent (Figs 3C, 5A; Fig. 15 in Olmi 1984); ADOs present in antennomeres 3–10 or 4–10...................... 5</p>
            <p>- Ocelli present (Figs 1B, C, D; Fig. 662 in Olmi 1984) ADOs absent or present in antennomeres 3–10 or 5–10, 6–10, 7–10, 8–10 or 9–10............................................................................................. 6</p>
            <p> 5 Mesosoma almost completely composed of fused segments (Figs 3C, 3D, 5A, 5B); prothorax fused with mesothorax; in mesosoma, only propleuron mobile and articulated with other segments of mesosoma...................................................................................... 11.  Apodryininae Olmi , Tribe  Apodryinini trib. nov. (extant) </p>
            <p>- Mesosoma partly composed of fused segments (Figs 3, 4 in Olmi 1987); entire prothorax mobile, articulated with rest of mesosoma........................................................................... 12. Plesiodryininae Olmi</p>
            <p> 6 Mesosoma almost completely composed of fused segments (Figs 1A, 2C); prothorax fused with mesothorax; in mesosoma, only propleuron mobile and articulated with other segments of mesosoma; head with ocelli situated in the anterior half of the face (Figs 1B, C, D); ADOs present in antennomeres 3–10..................................................................................................... 11.  Apodryininae Olmi , Tribe  Rovnodryinini trib. nov. (extinct) </p>
            <p>- Mesosoma not composed of fused segments; head with ocelli situated in the posterior half of the face (Fig. 5C); ADOs absent, or present in antennomeres 5–10, 6–10, 7–10, 8–10 or 9–10................................................... 7</p>
            <p>7 Chela without rudimentary claw (Fig. 314 in Olmi 1984)...................................................... 8</p>
            <p>- Chela with rudimentary claw (Fig. 1223 in Olmi 1984)...................................................... 10</p>
            <p> In the above key, the new tribe  Rovnodryinini is distinct from the nominal tribe  Apodryinini because of the presence of the ocelli (absent in  Apodryinini ). Following the description of  Rovnodryinini and the rules of the International Code of Zoological Nomenclature, the new tribe  Apodryinini can be described as follows: </p>
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	https://treatment.plazi.org/id/03B287A1C134FFBE38F240CDFB2CFCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Olmi, Massimo;Guglielmino, Adalgisa;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
03B287A1C131FFBD38F2457DFE61F8BF.text	03B287A1C131FFBD38F2457DFE61F8BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apodryinini Olmi & Guglielmino & Vasilenko & Perkovsky 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tribe  Apodryinini trib. nov.</p>
            <p> Type genus:  Apodryinus Olmi, 1984</p>
            <p>Diagnosis. Female: apterous; antennomeres 3–10 or 4–10 with ADOs; head without ocelli; occipital carina complete or incomplete; mesosoma almost completely composed of fused segments; in mesosoma, only propleuron mobile and articulated with rest of mesosoma; protarsus chelate; chela with rudimentary claw; tibial spurs 1/1/1 or 1/0/1. Male (see descriptions and figures in Olmi 1984, 2007 and Olmi et al. 2019): fully winged; fore wing with three basal cells (C2c, R2c, 1Cu2c) clearly enclosed by pigmented veins; maxillary palpi composed of five or six palpomeres; labial palpi composed of three palpomeres; epicnemium distinct; mesosternum distinct, not fused with mesopleura; mandible with 3–4 teeth (if quadridentate, with a rudimentary tooth); tibial spurs 1/1/2.</p>
            <p>Distribution. Known from Australia, South America and Africa.</p>
            <p>Hosts. Unknown.</p>
            <p>World species. Nineteen species are known.</p>
            <p>World genera. Seven.</p>
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	https://treatment.plazi.org/id/03B287A1C131FFBD38F2457DFE61F8BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Olmi, Massimo;Guglielmino, Adalgisa;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Olmi, Massimo, Guglielmino, Adalgisa, Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2022): Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa 5162 (1): 54-66, DOI: 10.11646/zootaxa.5162.1.3
