identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B5B96CDB2CBC4DFF06FF67A019FA7E.text	03B5B96CDB2CBC4DFF06FF67A019FA7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gaeolaelaps Evans & Till 1966	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Gaeolaelaps Evans &amp; Till, 1966</p>
            <p> Hypoaspis (Gaeolaelaps) Evans &amp; Till, 1966: 159 . </p>
            <p> Type species:  Laelaps aculeifer G. Canestrini, 1884 , by original designation </p>
            <p>The short diagnosis below was modified from the genus diagnosis in Beaulieu (2009). The subsequent detailed diagnosis is similar to the genus description in Beaulieu (2009), but excludes several characters of lesser significance at the genus level.</p>
            <p> Short diagnosis. Hypoaspidine laelapid mites with the following combination of characters: dorsal shield usually bearing 39 pairs of simple, short to moderately elongate setae, sometimes with 0–3 additional unpaired median setae; adult female sternal shield longer than broad (rarely broader than long), presternal area weakly sclerotised, usually lineate and granulate; epigynal shield tongue-shaped or flask-shaped, not markedly broadened posteriorly, bearing a pair of setae, and usually well separated from, and never touching, the subtriangular or pearshaped anal shield; opisthogastric cuticle usually with 7–9 pairs of simple setae (rarely more); epistome margin rounded or subtriangular, denticulate; six (rarely five or seven) deutosternal rows with at least five denticles each (rarely 1–4); chelicerae strong, chelate-dentate, pilus dentilis setiform; leg setation normal for  Laelapidae , including nine setae on genu IV (pl2 absent); av on femur II, ventral setae on genu and tibia II-IV, and subapical setae of tarsi II–IV usually slightly thickened to spine-like. However, there are numerous exceptions within the genus (see detailed diagnosis below, and genus description in Beaulieu, 2009). </p>
            <p>Detailed diagnosis</p>
            <p>1. Dorsal shield partly or completely covering dorsal idiosoma, not extending ventrally, suboval to strongly tapered from level of setae r3–4, oval in several cockroach, carabid and scarab beetle associates (Strong &amp; Halliday, 1994; Faraji &amp; Halliday, 2009).</p>
            <p> 2. Dorsal shield usually bearing 39 pairs of simple, short to moderately elongate setae, including Px2–3 (occasionally absent, including in some arthropod associates), and 0–3 unpaired median setae (Jx); z3 occasionally absent, and rarely a few others; setae, especially J4–5, Z5, sometimes inconspicuously barbed, rarely other setae, e.g. all dorsal setae except j1 and z1 barbed in  G. jondishapouri . </p>
            <p> 3. Lateral soft cuticle with 1–8 pairs of marginal (r–R) and 0 to few UR setae, rarely more, with at least 18 pairs of setae in  G. millipedus Rosario (1981) and  G. angustiscutatus (Willmann, 1951) , and 32 – 37 in the carabid associate  G. carabidophilus Trach (2012) . </p>
            <p> 4. Dorsal shield with 16 pairs of poroids (five podonotal and 11 opisthonotal) and 4–6 gland pores (2–3 podonotal, gd2 sometimes absent, gd4 usually absent; 2–3 opisthonotal, gd6 sometimes absent; see discussion); gd4 present in some specimens of  G. oreithyiae (Walter &amp; Oliver, 1989) (Kazemi, personal observation). </p>
            <p> 5. Presternal region weakly sclerotised and granulate and/or lineate, rarely with a pair of well sclerotised platelets, e.g. in  G. orbiculatus Nemati &amp; Mohseni (2013) . </p>
            <p> 6. Sternal shield longer than wide; rarely broader than long, mostly in arthropod associates, sternal shield length 0.9 x width in free-living  G. jondishapouri . </p>
            <p> 7. Posterior margin of sternal shield straight, or slightly convex or concave; rarely deeply indented, e.g. in the cockroach associate  G. concavus (Faraji &amp; Halliday, 2009) . </p>
            <p> 8. Sternal shield bearing three pairs of simple setae and two pairs of poroids; rarely setae st1 off shield, e.g.  G. aculeiferoides (Teng, 1982) ,  G. debilis (Ma, 1996) ,  G. krantzi (Arutunian, 1993) , or borne on paired anterior extensions of shield in  G. jondishapouri ; rarely poroids iv3 captured by sternal shield e.g. in  G. carabidophilus . </p>
            <p> 9. Setae st4 on soft cuticle, rarely on separate platelets; in some cases, e.g.  G. minor (Costa, 1968) , st 4 may wrongly appear to be inserted on the endopodal platelet, probably due to the soft cuticle bearing st4 being folded over the platelet (Kazemi, personal observation). </p>
            <p> 10. Epigynal shield tongue- or flask-shaped, not markedly broadened posteriorly, bearing one pair of simple setae, and not touching anal shield; ornamented with two slightly curved diagonal lines that typically join medially (as such forming an inversed V, or joined by a transverse line) and enclose posteriorly a reticulated area comprising several cells, posterior area without typical reticulation in the following species: smooth in  G. minor and  G. negevi (Costa, 1969) (considered to be a junior synonym of  G. gracilis (Meledzhaeva, 1963) by Bregetova, 1977); with eight long narrow cells in  G. schusteri (Hirschmann, 1966 sensu Costa 1974), and G. </p>
            <p> theodori (Costa, 1974) ; with a series of diagonal lines oriented posteromedially (forming a series of Vs) in the ant associate  G. glabrosimilis (Hirschmann et al., 1969) , and  G. franzi (Van Aswegen &amp; Loots, 1970); with a few similar diagonal to semi-circular lines in  G. ruggi (Strong &amp; Halliday, 1994) and  G. etiopicus (Berlese, 1918 sensu Van Aswegen &amp; Loots, 1970); with slightly curved, almost transverse lines in  G. rosei (Strong &amp; Halliday, 1994) ; with somewhat longitudinal lines in  G. circularis (Hyatt, 1964) . </p>
            <p> 11. Anal shield small, inversely pear-shaped or subtriangular, more or less rounded anteriorly, narrowed posteriorly, rarely oval or pentagonal; nearly rounded in  G. orbiculatus , oval in  G. millipedus and  G. rosei (Strong &amp; Halliday, 1994) and pentagonal in  G. brevior (Faraji &amp; Halliday, 2009) and  G. segregatus (Faraji &amp; Halliday, 2009) ; with two ‘shoulders’ or bulging laterally at position of pore gv3 (and level of para-anal setae) in  G. farajii Nemati &amp; Mohseni, 2013 . </p>
            <p> 12. Postanal seta usually slightly longer than para-anal setae, exceptionally absent in the fire-ant associate  G. invictianus Walter &amp; Moser, 2010 . </p>
            <p> 13. Cribrum typically with 3–4 rows of spicules; anteriormost row sometimes extending anteriorly to level or near level of para-anal setae, e.g. in  G. khajooii and  G. jondishapouri . </p>
            <p>14. Peritrematal shields narrow, connected to dorsal shield anteriorly, not extending beyond coxae IV posteriorly; nearly always free from exopodals. Parapodal and metapodal elements small, inconspicuously developed.</p>
            <p> 15. Peritremes narrow, usually reaching anteriorly to the level of coxae I, sometimes shorter, ending near middle of coxae II, exceptionally short in  G. carabidophilus , reaching only mid-level of coxa III. </p>
            <p> 16. Soft opisthogastric cuticle usually with 7–9 pairs of simple setae; hypertrichous in  G. carabidophilus , if considering ventrally inserted R–UR setae. </p>
            <p> 17. Male with holoventral shield, sometimes eroded laterally; rarely with a genitiventral shield eroded in opisthogastric region and a separate anal shield, e.g. in  G. invictianus . </p>
            <p> 18. Epistome always denticulate, with rounded or subtriangular anterior margin, sometimes more or less straight e.g. in  G. zhoumanshuae (Ma, 1997) and  G. dactylifera (Fouly &amp; Al-Rehiayani, 2011) ; rarely pronounced and pointed anteriorly e.g. in  G. brevior and  G. segregatus . </p>
            <p> 19. Deutosternal groove with six rows of denticles, rarely five or seven; seven rows in  G. dasypus (Menzies &amp; Strandtmann, 1952) , collected from an armadillo’s nest; each row bearing usually at least five denticles; rarely fewer, e.g. 1–5 denticles per row in  G. jondishapouri . </p>
            <p> 20. Palp tarsal claw usually two-tined, rarely three-tined; third tine long, or short and basal as in  G. jondishapouri . </p>
            <p> 21. Chelicerae well-developed, chelate-dentate; fixed digit often with a serrated row of small teeth proximal to the short, setiform pilus dentilis; movable digit with two teeth, exceptionally separated by a row of smaller teeth (in  G. angustiscutatus ). </p>
            <p> 22. Leg chaetotaxy normal for  Laelapidae ; nine setae on genu IV, including only one posterolateral; ventral and/or subapical setae of tarsi II-IV usually slightly thickened or modified into spines; av on femur II and ventral setae on genua and tibiae II-IV usually thickened or spine-like; femur IV occasionally with ad1 elongate, and often with ad2 and pd thickened; tarsus IV occasionally with 1–4 elongate setae (typically pd2–3, but also ad2–3). </p>
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	https://treatment.plazi.org/id/03B5B96CDB2CBC4DFF06FF67A019FA7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kazemi, Shahrooz;Rajaei, Asma;Beaulieu, Frédéric	Kazemi, Shahrooz, Rajaei, Asma, Beaulieu, Frédéric (2014): Two new species of Gaeolaelaps (Acari: Mesostigmata: Laelapidae) from Iran, with a revised generic concept and notes on significant morphological characters in the genus. Zootaxa 3861 (6): 501-530, DOI: 10.11646/zootaxa.3861.6.1
03B5B96CDB2BBC40FF06FA20A499FA0A.text	03B5B96CDB2BBC40FF06FA20A499FA0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gaeolaelaps jondishapouri Nemati & Kavianpour 2013	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gaeolaelaps jondishapouri Nemati &amp; Kavianpour, 2013</p>
            <p> Gaeolaelaps jondishapouri Nemati &amp; Kavianpour, 2013: 64 . </p>
            <p>Diagnosis (modified from Nemati &amp; Kavianpour, 2013). Female with dorsal shield abruptly tapering from level of S4 setae, bearing 39 pairs of moderately long setae, more or less reaching base of next seta in opisthosomal region, each seta with 1–4 small barbs; gland pores gd2 and gd6 present; seven marginal (r -R) setae, born on soft cuticle, with relatively thick sclerotised rings around alveoli. Sternal shield slightly wider than long, ratio of length/width ≈ 0.9; reticulate in its anterior third, smooth posteriorly, slightly concave posteriorly, with setae st1 on pointed edge of two anterior extensions of shield. Presternal region lightly sclerotised, lineate. Epigynal shield flask-shaped, slightly widened posteriorly, ratio of widths at level of broadest point/level of st5 ≈ 1.2, smooth except for an inversely Y-shaped line. Eight pairs of opisthogastric setae on relatively thick sclerotised rings around alveoli. Peritremes relatively short, anteriorly reaching posterior margin of acetabulum I. Epistome with anterior margin subtriangular. Deutosternum with six rows of 1–5 denticles. Fixed cheliceral digit with 5–7, rarely eight, teeth on its anterior half. Internal malae with long median projections and two additional pairs of lateral projections of similar length. Palp apotele with three tines, including a minute basal tine. Leg setae simple and mostly slender; ventral setae usually thicker than lateral and dorsal setae; av of femur II not spur-like; seta av on genu IV, setae av, pv and pl2 on tibia IV and setae md, pv2, pl2 and pd2 on tarsus IV thickened; setae pd2 and pv2 on tarsus IV inserted well proximal to ad2 and av2, respectively. Male with holoventral shield relatively narrow posteriorly, poorly developed behind coxae IV. Spermatodactyl short, curved and slightly extending beyond movable digit (male characters based on illustrations in Nemati &amp; Kavianpour, 2013).</p>
            <p>Redescription (Figs 1–12). Female (n=5, excluding holotype). Dorsal idiosoma (Figs 1, 11). Idiosoma 545–568 [573] long, 288–311 [291] wide. Dorsal shield 504–526 [562 (543–550)] long, 238–267 [276 (270–340)] wide, reticulate throughout except for smooth anteromedian region extending from behind j6 setae to anterior shield apex, covering most of dorsal idiosoma but leaving an exposed band of soft lateral cuticle; shield slightly tapering after humeral region from level of r3 to level of S4, where it then sharply tapers into a V-shape, and ends in a rounded posterior apex; shield with 39 pairs of setae of almost uniform length, relatively short and usually not reaching base of following setae. Dorsal shield setae j1 (39–42) [41] and z1 (16–18) [16 (17–19)] smooth; J5 (39–43) [39], Z4 (44–56) [58 (50–60)], S4 (48–52) [53], and S5 (53–59) [54] with 2–3 barbs; Z5 (57–61) [52] with 3–4 barbs; and all other setae (30–44) [33–47] usually with one small barb on distal half (Fig. 2); without unpaired setae. Shield with 16 pairs of discernible poroids (oval-shaped symbols) and six pairs of gland pores (circular symbols), including gd2 (posterolaterad of setae j4) and gd6 (posteromediad of z6). Setae r6 (24–27) [24 (34–36)], R1–6 (26–56) [28–57 (25–70] and one UR (33–34) [34] inserted on soft cuticle laterad of dorsal shield, with relatively thick sclerotised rings around alveoli (Figs 1, 11). Sigillae consistent across individuals, as illustrated in Fig. 1.</p>
            <p>Ventral idiosoma (Fig. 5). Tritosternum with narrow columnar base, 24–34 [34 (44–49)] long, 14–17 [17] wide at base, 7–9 [7] at apex, and two sparsely pilose laciniae 109–114 [111 (119–139)] long, and fused basally for 5–7 [6] µm. Presternal area weakly sclerotised, lineate and very slightly granulate. Soft integument behind coxae I with three pairs of gland openings flanked by two minute valves. Sternal shield 108–114 [115 (120–138)] long, 117–121 [122 (130–145)] wide, finely reticulated anterolaterally, otherwise smooth; anterior and posterior margin of shield moderately concave, anterolateral corners narrowly extending between coxae I–II, distally bearing gland pores gvb; endopodal elements between coxae II and III fused with shield; shield bearing three pairs of smooth setae st1 (37–41) [41], st2 (38–42) [40] and st3 (35–38) [38], and two pairs of slit-like poroids, iv1 and iv2, their axes oriented transversely; vestige of gland pores gv1 apparently present on posterior margin of sternal shield (Fig. 12). Metasternal setae (33–35) [34 (30–34)] and poroids iv3 on soft cuticle flanked by narrow endopodal elements between coxae III/IV. Epigynal shield flask-shaped, 196–209 [216 (180–213)] long, 99–108 [109] from setae st5 level to posterior margin, 77–93 [94] wide, slightly broadened past st5; anterior hyaline margin of shield irregularly convex and usually slightly covering posterior margin of sternal shield, with an inverse Y–shaped ornamentation, otherwise smooth; epigynal shield separated from anal shield by almost length of anal shield; setae st5 (31–33) [33 (30–33)] inserted on lateral margins of shield, approximately at level of posterior edge of coxae IV. Poroids iv5 inserted on soft cuticle, at level of tip of st5. A strip-like postgenital sclerite (sometimes divided into 2–4 narrow strips) closely bordering posterior margin of epigynal shield. Anal shield subtriangular, 92–98 [107 (99–103)] long, 87–95 [94 (94–96)] wide, with lineate-reticulate ornamentation, anterior margin of shield slightly convex, circumanal setae smooth, postanal seta (33–39) [38 (33–36)] longer than para-anal setae (26–30) [28 (27–30)]; cribrum well developed, with two single files of denticles each extending from cribrum to near base of para-anal setae; anal opening located at mid-level of shield; pair of glands gv3 inserted on shield lateral margins, at level between para-anal setae and posterior edge of anus. Peritrematal shields weakly developed posteriorly beyond stigmata, almost reaching level of coxae IV posterior margin, extending anteriorly and fused to dorsal shield at base of seta z1 level; bearing three gland pores (gp) and three poroids (ip): two of these poroids and one of these pores located on short, narrow post-stigmatic plate, one pore located near anterior extremity of peritreme (Fig. 6). Peritremes somewhat short (218–228 [221]), extending from stigmata to posterior margin of coxae I (Fig. 6). Exopodal and parapodal platelets narrow, divided into sclerotised strips along coxae II-IV; gland pore gv2 present anteromediad of parapodal platelet. One pair of minute paragenital platelets anterior to setae ZV1. Primary (most lateral) metapodal platelets suboval, with irregular margin, and secondary (more median) pair tiny, oriented more or less transversely. Opisthogaster with five pairs of poroids (four ivo; ivp) and three pairs of smooth ventral setae JV1 (30–34) [34 (30–32)], JV2 (32–36) [32 (30–32)] and ZV1 (25–29) [28 (30–33)], and five pairs of setae with 1–2 barbs: JV3 (35–37) [38 (30–32)], JV4 (40–43) [44 (39–41)], ZV2 (32–37) [38 (30–33)], ZV3 (25–29) [31 (30–33)] and ZV4 (39–42) [42 (30–33)]; JV5 (58–60) [60 (57–60)] with 3–4 barbs.</p>
            <p> Gnathosoma (Figs 3–4, 7–8). Anterior margin of epistome subtriangular and more or less rounded apically, denticulate, with about 50 small denticles (Fig. 3). Corniculi horn-like, 42–47 [46] long. Salivary stylets narrow and apically pointed. Internal malae fringed, with a pair of thick, contiguous median projections, and two pairs of thinner lateral projections, the inner pair shortest. Labrum considerably longer than corniculi. Hypostomal setae smooth, h1 (43–47 [(31–37)]&gt; h3 (37–39 [(27–30)])&gt; h2 (27–30 [28 (24–30)]), capitular (pc) setae smooth (39–43 [(36–41)]). Deutosternal groove with six rows of denticles, each row with 2–4 and more rarely 1–5 denticles (holotype has most deutosternal rows with two denticles close to each of lateral margins of deutosternum); lateral margins of deutosternum subparallel (Fig. 7). First (basal) segment of chelicerae 64–76 [67] long, second segment 175–191 [186 (240–245)] long, including fixed digit, 54–56 wide; fixed digit of chelicera 57–63 [63] long, bearing 5–7, rarely eight, teeth on its anterior half, including subapical offset tooth (gabelzhan); pilus dentilis short and setiform (Fig. 8). Movable digit of chelicera 72–77 [79 (75-84)] long, bidentate (Fig. 8). Palp chaetotaxy normal for  Laelapidae ; all setae smooth, al1 and al2 of palpgenu and al of palpfemur slightly thickened, palptarsus apotele with two main tines, and an additional, very short basal tine (Fig. 4). </p>
            <p> Legs (Figs 9–10). Leg chaetotaxy normal for  Laelapidae (sensu Evans and Till, 1965). All legs with ambulacrum, lengths of legs I-IV 549–562 [572 (539–549)], 416–428 [429 (444–452)], 386–405 [424 (418–422)] and 608–634 [621 (623–628)], respectively. Lengths of femora I 94 –106 [113 (123–126)], II 78 –87 [(80–84)], III 72 –76 [76 (71–77)], IV 97 –109 [124 (130–135)]; genua I 81 –85 [80 (87–95)], II 61 –66 [63 (65–70)], III 56 –60 [59 (58–62)], IV 78 –83 [78 (80–85)]; tibiae I 88 –95 [85 (36–39)], II 64 –70 [66 (73–77)], III 59 –62 [63 (60–65)], IV 90 –96 [89 (95–100)]; tarsi I 158–164 [163 (155–161)], II 118–130 [127 (129–136)], III 124–135 [123 (129–136)], IV 194–199 [200 (200–208)]. Leg setae mostly simple, needle-like; tarsi II-IV with al1, pl1, av1, and pv1 relatively thick; leg I without conspicuously thickened seta; trochanter II with al short and thickened; trochanter IV with ad slightly thickened; femur IV with ad2 short and slightly thickened; leg IV with the following setae considerably thickened or spine-like: av on genu; av, pv and pl2 on tibia; md, av2, pv2, pl2 and pd2 on tarsus. Coxa I each bearing two gland pores (gc) (Fig. 5). </p>
            <p> Remarks.  Gaeolaelaps jondishapouri can be distinguished from any other  Gaeolaelaps species by two distinctive, if not exceptional, characters among known members of the genus: (1) a dorsal shield abruptly tapering from the level of setae S4; (2) setae st1 inserted relatively far anteriorly from poroids iv1, on the apex of two pointed extensions of the sternal shield (making the anterior margin appear bilobed, see Figs 2, 12). Whereas the original description shows a sternal shield with a straight anterior margin, without extensions, leaving st1 completely off the shield, the re-illustration of the sternal shield by Kavianpour &amp; Nemati (2014) is similar to ours. Other notable discrepancies that we noticed between the original description and the specimens we examined (including the holotype) are: (1) denticulate rows of deutosternal groove usually with 2–4 denticles [rows appear smooth in the original illustration, and there is no mention of denticles in the text]; (2) palp apotele with a small basal tine, in addition to the two main tines; (3) 16 poroids and six gland pores on the dorsal shield [only 18 porelike structures in the original description, missing gland pore behind j4, and poroids near s3, behind s6, and laterad of Z4; Nemati &amp; Kavianpour had acknowledged in the text that they could have overlooked some pores or poroids]; (4) peritrematal shields anteriorly fused to dorsal shield, each bearing one gland pore [in the original description, peritremes appear without shielding anteriorly and therefore free from the dorsal shield]; (5) coxae I and surrounding soft integument with five gland openings; (6) cribrum with two files of denticles extending anteriorly to near the base of para-anal setae [not illustrated nor mentioned in the original description]; (7) setae h1 (43–47) and h3 (37–39) moderately longer [31–37 and 27–30, respectively, in the original description]. A few other measurements do not overlap between the original description and ours (e.g. dorsal shield; see notes in the redescription above), but these differences may at least in part be due to differences among populations; (8) most leg setae are needle-like, i.e. rather straight and with a relatively constant thickness through most of their length (not finely tapering as many setae in Nemati &amp; Kavianpour, 2013); setae of leg I and of coxa-tibia II are thin and needle-like, except thickened al1 on coxa and ventral setae on tibia (in contrast to the original illustrations showing most setae of legs I-II thickened or spine-like). Although this is a relatively inconspicuous feature,  G. jondishapouri also has all dorsal setae with one or more barbs, with the exception of j1 and z1. </p>
            <p> The specimens of  G. jondishapouri that we examined also differ from Nemati &amp; Kavianpour’s (2013) description by the shape of the posterior region of the dorsal shield. We would rather describe this region as Vshaped, or even subtriangular, with a rounded apex, instead of bell-shaped, as indicated in their species key (examination of the holotype shows a dorsal shield that may appear as ‘bell-shaped’ because the shield is bent posteriorly on that specimen). With such a distinctive tapering of the dorsal shield, we believe that  G. jondishapouri is closely related to  G. changlingensis (Ma, 2000) , with the  minor difference that the shield of  G. changlingensis abruptly tapers from the level of S3 instead of S4 as in  G. jondishapouri . Based on its description,  G. changlingensis also differs from  G. jondishapouri by its narrower sternal shield (Li-Ming Ma pers. comm.), st2 inserted more posteriorly, and possibly longer idiosomal setae (however, some of these apparent differences may be due to inaccurate illustrations). </p>
            <p> Nemati &amp; Kavianpour (2013) placed  G. jondishapouri in the  G. angusta species group (sensu Karg, 1979) based on the tapering posterior region of the dorsal shield. However, we do not concur that  G. jondishapouri can be placed in the  G. angusta group, because the four species originally placed in this species group have dorsal shields that taper much more anteriorly than that of  G. jondishapouri , and consequently are quite narrower posteriorly. Based on Karg’s hypothesis (1979), the dorsal shield of species in the  angusta group is wedge-shaped, characterised by ‘shoulders’ (typically near the level of r3–4), from which the shield gradually tapers to a relatively narrow apex (see, for example, the redescription of  G. queenslandicus (Womersley, 1956) by Costa, 1966). The dorsal shield of  G. jondishapouri is therefore quite distinct from that of members of the  angusta species group, and the same applies to the putatively close relative  G. changlingensis . Such shape of the dorsal shield is strongly reminiscent of that of  Stratiolaelaps scimitus (Womersley) , which differs from congenerics by a shield abruptly tapering at level of S1–2 (Walter &amp; Campbell, 2003). </p>
            <p> Very few species of  Gaeolaelaps have a deutosternum with fewer than five denticles per row.  Gaeolaelaps jondishapouri is among the exceptions, as is  G. spiniseta , which has a deutosternum with 3–4 denticles per row (Barilo, 1991). However,  G. jondishapouri can easily be distinguished from  G. spiniseta by several characters, including its posteriorly tapered dorsal shield (rounded in  G. spiniseta ), which bears 39 pairs of moderately long setae (38 in  G. spiniseta , with setae longer, usually reaching the base of next setae), flanked by seven pairs of marginal setae (only one pair in  G. spiniseta ); its sternal shield bearing st1 on anterior extensions of the sternal shield (st1 inserted closer to iv1 and well behind the anterior margin of the shield in  G. spiniseta ); and by its rounded epistome (subtriangular or pointed in  G. spiniseta , according to the illustration in Barilo, 1991). </p>
            <p>Material examined. Holotype female: Southwestern Iran, Ahwaz, Khuzestan Province, 2010, coll. M. Kavianpour. Paratypes: 10 females: southeastern Iran, Kerman Province, Jiroft County (28˚ 51' 29" N; 57˚ 71' 92" E), altitude 589 m above sea level, from cow manure in a citrus orchard, 4 Jan 2011, coll. A. Rajaei, deposited in ACISTE. One female with same data, deposited in ACJAZUT. One female with same data, deposited in CNC. Five females: collected from soil and litter, in the same orchard as above, same date and collector, deposited in ACISTE.</p>
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	https://treatment.plazi.org/id/03B5B96CDB2BBC40FF06FA20A499FA0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kazemi, Shahrooz;Rajaei, Asma;Beaulieu, Frédéric	Kazemi, Shahrooz, Rajaei, Asma, Beaulieu, Frédéric (2014): Two new species of Gaeolaelaps (Acari: Mesostigmata: Laelapidae) from Iran, with a revised generic concept and notes on significant morphological characters in the genus. Zootaxa 3861 (6): 501-530, DOI: 10.11646/zootaxa.3861.6.1
03B5B96CDB26BC44FF06F9D2A239FA0A.text	03B5B96CDB26BC44FF06F9D2A239FA0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gaeolaelaps khajooii Kazemi, Rajaei & Beaulieu	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gaeolaelaps khajooii Kazemi, Rajaei &amp; Beaulieu sp. nov.</p>
            <p>(Figs 13–28)</p>
            <p>Diagnosis (based on adult female). Dorsal shield clearly broadest at level of setae r3 (ratio of widths at r3 / S3 level≈1.5), progressively tapering until s6, subparallel from s6 to level of S4, rounded posteriorly; with 39 pairs of mostly smooth and moderately long setae; an additional, median unpaired seta may be present between J4–5; gland pores gd2 and gd6 absent; three pairs of marginal setae present on lateral soft cuticle. Sternal shield longer than wide (ratio of length/width≈1.3), reticulate throughout, its anterior margin with or without a small median invagination; posterior margin slightly convex, with two minute projections. Epigynal shield tongue-shaped, very slightly broadened posteriorly. Anal shield subtriangular, with two single files of denticles reaching level of paraanal setae; anal shield separated from epigynal by about length of anal shield. Peritremes relatively short, anteriorly reaching mid-level of coxae II. Post-stigmatic area of peritrematal shield narrowly extended to posterior edge of coxae IV. Opisthogastric and dorsolateral soft integument bearing seven and three pairs of setae, respectively.</p>
            <p>Deutosternal groove with six rows of 10–17 denticles. Internal malae with a pair of long median projections and a pair of shorter lateral projections. Palp apotele 2-tined. Fixed digit of chelicera with 10–13 teeth, including a serrated row of 6–9 small teeth posterior to pilus dentilis. Leg setae simple and slender, except following setae thickened and/or spine-like: ad3 (only slightly thickened) on femur I; av on femur II; pd on femur IV; all ventral setae on genua and tibiae II-IV (some only slightly thickened), and pl1 on tibia IV; md, av1–2, mv and pv1–pv2 on tarsus II; al1, av1–2, pv1–2, pl1 (slightly thickened), md and mv on tarsus III; pl2 (slightly thickened), av2, pv2 and mv on tarsus IV; pd2–3 on tarsus IV slender and elongate.</p>
            <p>Description. Female (n=3). Dorsal idiosoma (Fig. 13). Idiosoma length 541–552, width 301–307. Dorsal shield 489–522 long, 203–213 wide at j6 level, covering most of dorsal idiosoma, 248–253 wide at level of r3 and 163–170 at level of S3; reticulate throughout except anteromedially; with 39 pairs of needle-like setae of which some of the most posterior pairs (e.g. J4–5, Z5, Jx) have a few minute barbs; 0–1 unpaired median seta Jx between setae J4-J5. Dorsal shield setae j 1–5 32–36, j 6 28–31, z 1 23–28, z 2 31–32, z3 33–34, z4 34–38, z5 34–36, z 6 28–30, s 1 22 –23, s 2 24 –27, s3 35 –40, s 4 29 –35, s 5 28 –31, s 6 29 –36, r 2 31–35, r3 38–45, r 4 31–34, r 5 29–34, J 1–5 27–31, Jx 28–30, Z 1 28–31, Z 2–3 24–29, Z 4 28–30, Z5 38–44, S 1–4 29–34, S5 33 –39 long. Setae R1 (29–31), R4 (30–33) and R5 (34–38), and a pair of poroids (idR3) on lateral soft integument. Dorsal shield with 16 pairs of poroids and only four pairs of gland pores (gd2 and gd6 pores absent).</p>
            <p>Ventral idiosoma (Figs 14, 16–17). Tritosternum with a narrow columnar base, 32–36 long, 14–17 wide at base, 7–9 wide at apex, and two sparsely pilose laciniae, length free for 90–95 and fused basally for 2–4 µm. Presternal area lightly sclerotised and punctate, with a few transverse lines. Sternal shield 154–160 long, 114–118 wide, reticulate throughout; anterolateral corners narrowly extending between coxae I-II; anterior margin irregularly straight, with or without a small median invagination; posterior margin slightly convex, with two minute projections near medial axis, which may represent vestiges of gland pore gv1 (Figs 16–17); shield bearing three pairs of smooth setae st1 (38–39), st2 (37–38) and st3 (34–36), and two pairs of poroids, lyrifissures iv1 slitlike and oriented obliquely, iv2 suboval. Metasternal setae (27–30) and poroids iv3 on soft cuticle. Endopodal elements between coxae III–IV narrow, anteriorly free from sternal shield. Epigynal shield 185–190 long, 92–95 from st5 level to posterior margin, 69–71 wide, 58–62 wide at narrowest point (near coxae IV level), and 81–83 at broadest point past setae st5; anterior hyaline margin irregular, not covering posterior area of sternal shield; surface reticulated with an inverse Y–shaped pattern, with the Y posteriorly embracing eight large cells; setae st5 (27–29) inserted on lateral margins of shield at level of posterior margin of coxae IV, and poroids iv5 inserted laterad of st5. Anal shield subtriangular, anteriorly rounded, 74–80 long, 74–80 wide, anterior margin gently rounded, and surface lineate-reticulate; circumanal setae smooth, postanal seta (35–38) longer than para-anal setae (29–31), cribrum well developed, narrowly extending laterally to level of adanal setae; anus located slightly posterior to mid-level of shield; pair of glands gv3 inserted on lateral margins of shield at level of anterior margin of anus. Peritrematal shields well developed anteriorly and fused narrowly to dorsal shield at level of z1, with a narrow strip of granular cuticle parallel to anterior edge of shield; between stigmata and coxa II, shield almost limited to a relatively narrow band of cuticle at level between coxae II and III, bearing a poroids and a gland pore; post-stigmatic region of shield narrowly extending to posterior level of coxae IV, bearing two and one gland pore; an additional pore located on peritrematal shield at level of seta s1. Peritremes relatively short (178–182) and narrow, extending from stigmata to mid-level of coxae II. Three exopodal platelets between coxae I-II, II-III and III-IV present, anterior two platelets small and subtriangular, those between coxae III-IV slightly extending posteriorly; parapodal platelets strip-like, bearing gland pore gv2. Opisthogastric integument with two pairs of small, narrow paragenital platelets between st5 and ZV1, and two pairs of narrow metapodal platelets, the smallest at or above level of ZV1 and the largest one near level of JV1; seven pairs of ventral opisthogastric setae JV1 (19–23), JV2 (18–22), JV3 (24–26), ZV1 (23–28) and ZV2 (27–31) smooth; JV4 (28–32), JV5 (44–49) with 1–2 minute barbs; and five pairs of poroids.</p>
            <p> Gnathosoma (Figs 20–24). Anterior margin of epistome convex and finely denticulate, with about 30 denticles (Fig. 20). Corniculi horn-like (58–64). Internal malae fringed, slightly longer than corniculi, with a pair of long median projections and a pair of shorter lateral projections. Labrum acuminate, considerably longer than internal malae. Hypostomal and capitular setae smooth, h1 (50–51)&gt; h3 (35–38)&gt; h2 (34–35)≈ pc (32–35). Deutosternal groove with six rows of denticles, progressively broader from posterior to anterior, each with 9–17 denticles; anteriormost row angled medially. First cheliceral segment 58–67 long, second segment 190–196 long and 54–56 wide; fixed digit 73–75 long, with 10–13 teeth, including (from distal to proximal) a subapical large offset tooth (gabelzhan) followed by two other teeth, one small and one large, anterior to the short, setiform pilus dentilis, and 6–9 smaller, tightly aligned teeth followed posteriorly by a larger tooth; movable digit of chelicera 85–87 long, bidentate; dorsal cheliceral seta short and setiform (Fig. 22). Palp chaetotaxy normal for  Laelapidae , all setae smooth; palpgenu setae al1 and al2 slightly thick and spatulate, palpfemur al slightly thick and spine-like; palp apotele 2-tined (Figs 23–24). </p>
            <p> Legs (Figs 25–28). All legs with ambulacrum, that of leg I longer (35–36) than those of legs II-IV (24–26) (Figs 18–19), lengths of legs I-IV as follows: 591–597, 425–451, 384–392, 582–605, respectively. Lengths of femora I 104–111, II 85 –96, III 75 –77, IV 117–123; genua I 83 –87, II 69 –70, III 50 –53, IV 89 –92; tibiae I 95 –97, II 66 –67, III 51 –53, IV 96 –98; tarsi I 156–163, II 108–115, III 105–110, IV 156–160. Leg chaetotaxy normal for  Laelapidae (sensu Evans &amp; Till, 1965) and  Gaeolaelaps (see Beaulieu, 2009). Most setae simple, needle-like and of similar length, except a few shortened, elongate or thickened setae: trochanter I with al short and slightly thickened; trochanter II with al1 thickened; trochanter IV with pv1 slightly thickened; femur I with ad3 slightly thickened, femur II with av somewhat spine-like and al2 very short, slightly thickened; femur IV with pd spinelike, ad2 slightly thickened, and pl short and thin; genu II with av short and spine-like, and genua III-IV with ventrals slightly thickened; tibia II with av1 spine-like, pv1 thickened; tibia III and IV with pv spine-like, av slightly thickened; pl also spine-like on tibia IV; tarsus II with av1–2, md, mv, and pv1–2 spine-like; tarsus III with md and mv spine-like, and av1–2, pv1–2 slightly thickened; tarsus IV with pl2 slightly thickened, mv, av2 and pv2 thickened, and pd2 and pd3 slender and elongate with pd2 longer than half the length of tarsus. Coxa I bearing two gland pores (gc) (Fig. 14). </p>
            <p>Material examined. Holotype: female southern Iran, Kerman Province, Baft County (28˚39' 46" N; 56˚45' 37" E), altitude 1044 m above sea level, from soil and litter at an alfalfa farm, 2 April 2012, coll. A. Rajaei, deposited in ACISTE. One paratype female with same collection data, deposited in ACISTE; another paratype female with same collection data, deposited in CNC.</p>
            <p>Etymology. The species is named in honour of the famous Persian poet, Khajooi-e Kermani.</p>
            <p> Remarks. The new species can be placed in the  similisetae species group (sensu Karg, 1979) based on its short peritremes reaching only the mid-level of coxae II, and differs from other species of the group by the shape of the dorsal shield, which is clearly widest at the level of setae r3 and tapers posteriorly. This shape is reminiscent of the  G. angusta species group, but differs from most members of that group by its dorsal shield with almost parallel sides in the opisthonotal region (from S1 to S4). The species most similar to  G. khajooii is  G. zhoumanshuae (Ma, 1997) , which, justifiably, was classified in the  angusta species group by Nemati &amp; Kavianpour (2013). In addition to the very similar shape of their dorsal shield,  G. zhoumanshuae and  G. khajooi both have peritremes shortened anteriorly, ending before the anterior margin of coxae II, seven opisthogastric setae (six in  G. zhoumanshuae , but JV 5 may have been illustrated dorsally), three dorsomarginal (R) setae (four in G.  zhoumanshuae , perhaps including JV5 dorsally), and have a cheliceral fixed digit with a serrated row of small teeth proximally to the pilus dentilis (based on figures in Ma, 1997).  Gaeolaelaps khajooii differs from it at least by having 39 pairs of setae on the dorsal shield, including Px2–3, which are absent in  G. zhoumanshuae . The new species also resembles  Hypoaspis (sensu lato)  atomarius Berlese, 1917 (sensu Van Aswegen &amp; Loots, 1970), which has a similar dorsal shield albeit posteriorly wider, but it can be easily distinguished from  H. atomarius by having one ventral setae on genu IV (two ventrals in  G. atomarius ) and 39 setae on dorsal shield (38 in  G. atomarius , with z3 absent). </p>
            <p> The two minute projections on the posterior margin of the sternal shield are unusual, but also occur in other  Gaeolaelaps species, including  G. jondishapouri ,  G. nolli (Karg, 1962) ,  G. praesternalis (Willmann, 1949) and  G. farajii , and an undescribed species that have dorsal shield shaped similarly to  G. khajooii . It is possible that these minute projections represent the vestiges of gland pores gv1. </p>
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	https://treatment.plazi.org/id/03B5B96CDB26BC44FF06F9D2A239FA0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kazemi, Shahrooz;Rajaei, Asma;Beaulieu, Frédéric	Kazemi, Shahrooz, Rajaei, Asma, Beaulieu, Frédéric (2014): Two new species of Gaeolaelaps (Acari: Mesostigmata: Laelapidae) from Iran, with a revised generic concept and notes on significant morphological characters in the genus. Zootaxa 3861 (6): 501-530, DOI: 10.11646/zootaxa.3861.6.1
03B5B96CDB22BC46FF06F9C8A3CBF855.text	03B5B96CDB22BC46FF06F9C8A3CBF855.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gaeolaelaps ahangarani Kazemi & Beaulieu	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gaeolaelaps ahangarani Kazemi &amp; Beaulieu sp. nov.</p>
            <p>(Figs 29–40)</p>
            <p>Diagnosis (based on adult female). Dorsal shield suboval, reticulated throughout, covering almost all dorsal idiosoma, leaving exposed a narrow lateral band of soft cuticle which bears four pairs of marginal setae (R1–R2, R4–5); shield with 37 pairs of short, needle-like dorsal setae, setae Px2–3 and unpaired setae Jx absent, with 21 pairs of pore-like structures, including six pairs of gland pores (gd2 and gd6 present). Sternal shield longer than wide (length/width ratio ≈1.35), reticulate throughout except for a narrow smooth area posteriorly; anterolateral arms of shield broadly fused to endopodals between coxae I–II, and posterior margin of shield slightly convex. Epigynal shield tongue-shaped, only slightly widened posteriorly, with eight cells surrounded by a Λ-shaped line, with cell borders sinuous. Post-stigmatic region of peritrematal shield short, extending slightly beyond level of mid-coxa IV. Soft opisthogastric cuticle with ten pairs of short setae, with alveoli surrounded by relatively thick sclerotised rings. Anal shield somewhat pear-shaped, broadly rounded anteriorly; anal opening almost entirely on anterior half of shield; cribrum spicules not extending significantly anteriorly. Peritremes long, reaching anterior margin of coxae I. Epistome margin rounded, with variously sized denticles. Deutosternum with six rows of 17–20 denticles. Fixed digit of chelicera with seven teeth. Internal malae with two pairs of lateral fringed projections in addition to the pair of long median projections. Palp apotele 2-tined. Femur II with av seta thickened, tarsi II-IV with most subapical setae thickened or spine-like; setae av1, pv1 on tarsus II, pl2–3 on tarsus IV, and pl1 on tibia IV apically blunt, spur-like; tarsus IV with pd2 slightly longer than other setae, otherwise without significantly elongate setae, ad3 and pd3 very short; most dorsal setae of genu IV and tibia IV short.</p>
            <p>Description. Female (n=2). Dorsal idiosoma (Fig. 29). Idiosoma 486–494 long, 266–271 wide. Dorsal shield 486–494 long, 247–252 wide, suboval, reticulate throughout, more distinctly on opisthonotal region, and with cells more elongate between setae j5 and j6; shield broadest at level of setae r3, ratio of widths at the level of r3 / S3 =1.2; shield covering most of dorsal idiosoma, leaving exposed a narrow marginal band of soft cuticle. Dorsal shield with 37 pairs of short setae, setae Px2–3 absent, setae j2–4, z4, s3, r2–3, Z5 longest (20–25), z1 shortest (6–8), other setae 12–17 long; Z5 usually with 1–2 minute barbs. Setae R1–R2 (12–14), R4 (14–15), R5 (11–12) and one pair of poroids (idR3) on soft lateral cuticle. Dorsal shield with 16 pairs of discernible poroids (oval-shaped symbols) and six pairs of gland pores (circular symbols).</p>
            <p>Ventral idiosoma (Fig. 30). Tritosternum with a narrow columnar base, 32–33 long, 10–11 wide at base, 7–8 wide at apex, and two free pilose laciniae (68–71). Presternal area weakly sclerotised, granulate-lineate. Sternal shield 146–150 long, 109–111 wide, distinctly reticulate throughout except for a narrow smooth area posteriorly; anterior margin of shield straight, posterior margin slightly convex with a small median depression; anterolateral corners broadly fused to endopodal platelets between coxae I-II, bearing gland pores gvb; shield bearing three pairs of smooth setae st1 (24–25), st2 (26–27) and st3 (26–28), and two pairs of poroids, iv1 slit-like and iv2 sub-oval. Metasternal setae (26–28) and poroids iv3 on soft cuticle (iv3 asymmetrically captured by sternal shield in one specimen). Epigynal shield tongue-shape, very slightly broadened posteriorly, 166–169 long, and 81–83 from st5 to posterior margin, 70–72 wide; anterior hyaline margin slightly convex, irregular, covering posterior smooth area of sternal shield; epigynal shield with a Λ–shape line delimiting posteriorly eight cells with sinuous borders; setae st5 (20–21) on lateral margins of shield, almost at level of posterior edge of coxae IV. Paragenital poroids iv5 on soft cuticle, near level of coxae IV posterior edges. Anal shield somewhat pear-shaped, anterior margin of shield broadly rounded, 67–70 long, 56–58 wide, lineate-reticulate anteriorly and laterally; circumanal setae smooth, postanal seta (20–22) longer than para-anal setae (15–16); cribrum well developed, with three rows of spicules (four medially), the two anteriormost rows only slightly extending anteriorly of postanal seta; anal opening located almost entirely on anterior half of shield; para-anal gland pores gv3 on lateral shield margins at level of the posterior edge of anal opening. Peritrematal shields relatively well developed throughout, anteriorly fused to dorsal shield behind setae z1, bearing one pair of gland pores near external margin of shield at level of anterior edge of coxae III and one pair of poroids at level of posterior edge of coxae II; poststigmatic region with a longitudinal line from stigmata to shield apex, and with two pairs of poroids and one pair of gland pores. Peritremes long (236–243), reaching anterior margin of coxae I. Exopodal platelet between coxae II-III well developed, exopodal between coxae III-IV fused to parapodal. Opisthogastric soft integument with pair of primary metapodal platelets moderately small, narrow, laterad of ZV1–2; 2–3 additional pair(s) of minute metapodal platelets anterior to and inbetween ZV1 and primary metapodals; paragenital platelets minute, at level of ZV1; five pairs of poroids and 10 pairs of short, needle-like setae JV1–5, ZV1–5, 14–20 long.</p>
            <p> Gnathosoma (Figs 31–34). Anterior margin of epistome somewhat rounded, with about 17 variously sized teeth (Fig. 31). Corniculi horn-like, 45–47 long. Salivary stylets narrow and apically pointed, aligned beneath corniculi. Internal malae fringed, with a pair of adjacent median pilose projections, flanked by two pairs of shorter and thinner lateral projections. Labrum acuminate, pilose, considerably longer than corniculi. Hypostomal and capitular setae smooth, h3 (46–49)&gt; h1 (34–37)&gt; pc (26–27)&gt; h2 (19–21). Deutosternal groove with six rows of 17–20 denticles each; lateral margins of deutosternum subparallel (Fig. 32). Basal segment of chelicerae 50–52 long, second segment 140–143 long, 46–48 wide; fixed digit of chelicera 40–42 long, with seven teeth, including subapical offset tooth (gabelzhan), followed by two small teeth, and four larger teeth proximal to the short and setiform pilus dentilis; dorsal seta short and setiform. Movable digit of chelicera 48–50 long, bidentate (Fig. 33). Palp 146–149 long, palp chaetotaxy normal for  Laelapidae ; all setae smooth, al1 and al2 on palpgenu and al on palpfemur thick and subspatulate, ad on palpfemur slightly thickened, somewhat spine-like; palptarsus apotele 2- tined, anterior tine with spatulate hyaline margin (Fig. 34). </p>
            <p> Legs (Figs 35–40). Leg chaetotaxy normal for  Laelapidae (sensu Evans &amp; Till, 1965). Legs II–III with largest claws and pulvillus, and broadest ambulacral stalk; leg I with narrowest stalk, claws and pulvillus; lengths of legs I- IV 435–438, 335–341, 289–292 and 432–440, respectively. Lengths of femora I 70 –73, II 57 –59, III 56 –59, IV 89 –92; genua I 62 –65, II 62 –63, III 38, IV 62 –67; tibiae I 70 –72, II 51 –54, III 38, IV 62; tarsi I 116–119, II 73 –76, III 83 –86, IV 113–121. Leg setae mostly thin and moderately long, except: trochanter I with al and ad short and slightly thickened; trochanter II with al slightly thickened; trochanter IV with pv1 slightly thickened; femur II with av and pd1 slightly thickened, ad3 and al2 short; femur III with al and ad1 thickened, ad2, pd and pl short; femur IV with ad1 and av slightly thickened, ad2, pd and pl short; genu II with pd2 slightly thickened, and genua III-IV with ventrals slightly thickened; tibiae II-III with pv thickened; tibia IV with av and pv thickened, pl1 thick, somewhat spur-like, pd2–3 very short; tarsus II with al2–3 and pl2–3 slightly thickened, av1 and pv1 stout, spurlike (apically blunt), al1, pl1 and md stout and spine-like, mv, av2 and pv2 thick; tarsus III with pl2–3 slightly thickened, ad2, pd2, pv2 and al3 thickened, av1–2, pv1–2, al1–2, mv and pl1 thickened, somewhat spine-like, md stout, spine-like; tarsus IV with most setae thickened and of moderate length, and pl2–3 somewhat spur-like, pl3 half as long as pl2, pd2 slightly longer than other setae, ad3 and pd3 short and thin. Coxa I bearing two gland pores (gc) (Fig. 30). </p>
            <p>Material examined. Holotype: female, northern Iran, Mazandaran Province, Tonekabon County, Tirom Forest (40˚ 62' 69" N; 47˚ 11' 26" E), 1465 m above sea level, from decayed wood of Beech trees, 2 June 2010, coll. Y. Ahangaran, deposited in ACISTE. One paratype female with same collection data, deposited in ACISTE.</p>
            <p>Etymology. The species is named in honour of Yazdanfar Ahangaran (General Office of Natural Resources of Mazandaran Province, Nowshahr, Iran), for his help in collecting these mites.</p>
            <p> Remarks.  Gaeolaelaps ahangarani can most readily be distinguished from all other species in the genus by the following combination of characters: very short dorsal setae, particularly so in the opisthonotal region; the absence of Px2–3; only three pairs of marginal setae on soft lateral cuticle; epigynal shield with posterior cells with sinuous borders; an anal shield broadly rounded anteriorly, with the anus almost entirely in the anterior half of shield; and modified leg setae, such as pl2–3 of tarsus IV and pl1 of tibia IV spur-like, and ad3, pd3 very short. </p>
            <p> Many other  Gaeolaelaps have short dorsal setae, but the large majority of them possess Px2–3 setae, rarely have the anal opening positioned as anteriorly as in the new species (based on species descriptions), and have other distinctive characters. For instance,  G. tenuisetus Rosario (1981) has short setae, but seems to possess Px2–3 (at least on one side; the illustration is difficult to interpret), and has an epigynal shield considerably expanded posteriorly (nearly parallel-sided in  G. ahangarani ), and a subtriangular anal shield that is relatively flat anteriorly.  Gaeolaelaps analis (Karg, 1982) has a broader, more oval dorsal shield, Px2 present, and a broader anal shield with anus located on its posterior half.  Gaeolaelaps gleba Karg, 1979 (=  G. glabra Karg, 1978 ) has Px2–3 setae as well as shortened peritremes. </p>
            <p> The few  Gaeolaelaps species lacking Px2–3 setae that we are aware of are distinguished from  G. ahangarani by at least a few characters.  Gaeolaelaps queenslandicus and  G. angustus (Karg, 1965) have a posteriorly tapered dorsal shield, longer dorsal setae, a three-tined palp apotele, and a deutosternum narrowing posteriorly (based on Costa, 1966). Based on the illustrations,  Gaeolaelaps tengi (Gu &amp; Bai, 1991) may appear similar to the new species, but it has longer setae, a broader, subtriangular anal shield that is flatter anteriorly, a sternal shield truncate posteriorly, with st3 setae near the edge, a post-stigmatic region of peritrematal shield almost reaching the posterior margin of coxae IV (shorter in  G. ahangarani ), and a narrower deutosternal groove.  Gaeolaelaps wufengensis (Liu &amp; Ma, 2003) also has relatively short setae, but possesses an epigynal shield ornamented by circular lines posteriorly, seven rows of deutosternal denticles, and an acuminate epistome.  Gaeolaelaps barbarae (Strong 1995) , associated with funnel-web spiders, has a more oval dorsal shield, and much longer setae, and a subtriangular epistome. Other arthropod associates also lack Px2–3 (e.g.  G. disjuncta Hunter &amp; Yeh, 1969 ,  G. circularis ,  G. ruggi , and  G. rarosae Rosario, 1981 ), but have fewer setae on the dorsal shield (&lt;37), longer setae, and often a more oval dorsal shield, a shorter, more compact sternal shield, and/or an epigynal shield with distinct ornamentation. </p>
            <p> Gaeolaelaps ahangarani shares characters with  G. aculeifer and relatives (see Discussion), such as the spinelike setae on tarsi, especially of legs II-III, the deutosternal groove with numerous denticles per row, and similarly shaped and ornamented dorsal, sternal, and epigynal shields (although these shields are similar in several other  Gaeolaelaps species). Like  G. aculeifer , it also has internal malae with two pairs of short lateral projections. However, it differs from  G. aculeifer and relatives by lacking Px2–3 setae, and having only three marginal setae laterad of the dorsal shield, a fixed cheliceral digit with fewer teeth proximal to the pilus dentilis, and shortened leg setae (see Diagnosis). At first glance, the sinuous borders of the cells of the epigynal shield appear as a fairly unique character of  G. ahangarani . However, we suspect that such ornamentation occurs in other species, which we may have overlooked, or that are still undescribed. </p>
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	https://treatment.plazi.org/id/03B5B96CDB22BC46FF06F9C8A3CBF855	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kazemi, Shahrooz;Rajaei, Asma;Beaulieu, Frédéric	Kazemi, Shahrooz, Rajaei, Asma, Beaulieu, Frédéric (2014): Two new species of Gaeolaelaps (Acari: Mesostigmata: Laelapidae) from Iran, with a revised generic concept and notes on significant morphological characters in the genus. Zootaxa 3861 (6): 501-530, DOI: 10.11646/zootaxa.3861.6.1
