taxonID	type	description	language	source
03B787E5BA57C13DFDBA1BE671460734.taxon	diagnosis	Diagnosis Zyuzin defined this subfamily by the character “ The embolus enters the tegular apophysis, forming a single structural complex with it ” (Zyuzin 1985: 48). Roewer recognized this distinguishing feature for Evippinae (at the time, tribe Evippeae): Metatarsus IV shorter than patella IV + tibia IV (Roewer 1959). Roewer's character is invalid for identification, as Evippa amitaii sp. nov. deviates from these proportions.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA57C13DFDBA1BE671460734.taxon	description	Description Small- to medium-sized lycosids (Figs 1 – 5), superficially similar to Pardosinae and Lycosinae in somatic appearance, but distinct in genital structure (Figs 6 – 13). Anterior eye row narrower than second eye row (PME). Tibia I with 2 – 6 pairs of ventral spines (apart from apical pair). Spinnerets not elongated. Body densely covered with short (often specialized) setae (Alderweireldt 1992) (Figs 1, 14). Sexual dimorphism usually weak (Figs 4 – 5) (but see section about Evippomma (Figs 1 – 2, 4 – 5 )). Zyuzin (1985) described the Evippinae as having an embolus encased in a sheath of transparent tissue (Figs 6 – 8); embolus base in mesoapical position, partly flattened, lying against cymbium; bent sharply near tip, inserted into tegular apophysis, forming single structure. Epigyne atria very shallow, pale, conspicuous; septum with narrow base, often covered with setae; spermathecae usually large and conspicuous (Figs 10 – 11, 13). Coloration variable; carapace often with median band (Figs 1 – 3). Certain genera (Evippa, Zenonina) have a derived habitus, clearly discernible from other lycosid genera (Roewer 1959; Alderweireldt 1991, 1992; Marusik Kovblyuk & Koponen 2011).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA57C13DFDBA1BE671460734.taxon	biology_ecology	Natural history Most species inhabit deserts, steppes and savannas, but species of open patches in and along forests (Xerolycosa spp.) are known. Zyuzin (1985) described Xerolycosa as found in dry conifer forests, steppes and arable land, while Evippa inhabits flood plains, semideserts and deserts, often as a sole representative of Lycosidae. They are largely presumed vagrant (Alderweireldt & Jocqué 2017), although burrow construction behavior is known from species of Xerolycosa (Marusik et al. 2011) and Evippomma (Bayer et al. 2017). Both diurnal and nocturnal activity has been observed (Zyuzin 1985; * Direct submission to Gen Bank by Astrin J., Hoefer H., Spelda J., Holstein J., Bayer S., Hendrich L., Huber B. A., Kielhorn K. - H., Krammer H. - J., Lemke M., Monje J. C., Moriniere J., Rulik B., Petersen M., Janssen H. and Muster C. I. Armiach Steinpress, pers. obs.). As in all lycosids, the female carries the egg sac attached to her spinnerets and carries the young on her body for some time after hatching (I. Armiach Steinpress, pers. obs.).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA57C13DFDBA1BE671460734.taxon	distribution	Distribution The subfamily is restricted to the Old World where it is widely distributed, except in the polar region, wet tropics and wet subtropics. Xerolycosa is found across the Palearctic, from Europe to Japan (with a doubtful species from Zanzibar). Proevippa, Pseudevippa and Zenonina are restricted to sub-Saharan Africa. Evippomma is found in Africa and India (Indian species might belong to a different genus). Evippa is found across arid and semiarid climates throughout Africa and Eurasia, from Tanzania to Siberia (World Spider Catalog 2020).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA57C13DFDBA1BE671460734.taxon	discussion	Relationships Molecular studies that included Xerolycosa tend to place it in a rather basally branching position in Lycosidae, away from the Lycosinae (Park et al. 2007; Zehethofer & Sturmbauer 1998). Murphy et al. (2006) have placed Xerolycosa as a sister taxon of the venoniin genera Aulonia (C. L. Koch, 1847) and Hygrolycosa (Dahl, 1908) (Venoniinae was not recovered in the study). A recent molecular phylogeny included for the first time three representatives of Evippinae: Proevippa, Xerolycosa, and Evippomma, and recovered Evippinae as monophyletic (Piacentini 2019). Composition Evippinae consists of 67 species in six genera: 38 species in Evippa Simon, 1882; seven species in Evippomma Roewer, 1959; eleven species in Proevippa Purcell, 1903; one species in Pseudevippa Simon, 1910; four species in Xerolycosa Dahl, 1908; and six species in Zenonina Simon, 1898.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	description	Figs 3 – 13	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	type_taxon	Type species Evippa arenaria (Audouin, 1826)	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	diagnosis	Diagnosis Evippa is distinguished easily from all other lycosids by a combination of characters: elongated tarsal claws, 5 – 6 pairs of ventral spines on tibia I (apart from apical pair), transverse depression in carapace posterior to ocular area (Fig. 3) (Alderweireldt 1991).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	description	Description Small- to medium-sized lycosids. Cephalic region elevated, separated from thoracic region by transverse depression. Ocular area almost as wide as cephalic region, adorned with long, forward-pointing setae. Clypeus vertical. Anterior row of eyes procurved, narrower than PME. AME larger than ALE. Labium wider than long. Most species with long, slender legs (see E. onager; Fig. 3). Claws usually long, toothed near base. Tarsi of many species with pseudoarticulation. Tibia I with 5 – 6 pairs of spines (apart from apical pair). Coloration cryptic, usually mottled yellow or brown (Fig. 3). Carapace usually with pale median band. Body covered with short setae throughout (Fig. 14 A) (Alderweireldt 1992). Male and female of similar appearance. Embolus large; base in meso-apical position (Figs 6 – 9, 12). Epigyne usually with well-developed, pale atria (Figs 10, 13). Spermathecae large, sperm ducts twisted (Figs 11, 13) (Tikader & Malhotra 1980; Alderweireldt 1991).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	biology_ecology	Natural history Evippa species inhabit deserts, steppes and savannas, often on sandy, clay or salt-covered plains (Barrientos et al. 2015; Ponomarev & Tsvetkov 2004) (Fig. 15), although mountain-dwelling species are known (Tikader & Malhotra 1980). All species are presumed vagrant (Alderweireldt & Jocqué 2017). Diurnal activity has been observed (I. Armiach Steinpress, pers. obs.) but most activity is crepuscular and nocturnal (Alderweireldt & Jocqué 2017). Adults in warm climates are observed active during most of the year (Barrientos et al. 2015).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	distribution	Distribution The species of Evippa are distributed in arid and semiarid environments across Eurasia and Africa. Eight species are reported from Africa, four species are reported from Europe, and 34 are reported from Asia (13 from Central Asia, nine from India, nine from the Middle East, seven from China and one from Siberia) (World Spider Catalog 2020). In Israel four species are found.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA59C135FD5C1952712A0119.taxon	discussion	Relationships The genus appears to be closely related to Pseudevippa Simon, 1910, from which it is distinguished by the number of ventral spines on tibia I (Alderweireldt 1991).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	description	urn: lsid: zoobank. org: act: CB 6 C 5814 - B 74 C- 4 FEC-AD 41 - 27 D 18232 D 3 FC Figs 4 A, 5 A, 6 A, 7 A, 8 A, 9, 10 A, 11 A, 12, 13 A, 13 C	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	diagnosis	Diagnosis Recognized by a combination of genital characters (compare with Evippa aequalis Alderweireldt, 1991). Male palp: tegular apophysis tip flat, curved ventrad. Basal process of tegular apophysis long, with sharp tip. Bulb differs from the bulbs of other Evippinae species found in Israel, and from that of E. aequalis, by the terminal apophysis (process between tegular apophysis and cymbium) being at least at 40 ° angle to tegular apophysis and oriented ventrad (in the others the tip of the tegular apophysis is oriented distad) (Figs 6 A, 7 A, 8 A, 12). Epigyne: Atrium borders indistinct. Septum wide, longer than atrium, narrowing slightly anteriorly and mesally, creating a small protrusion (Figs 10 A, 13 A). Spermathecae club-shaped, long, reaching over frontal edge of atria (Figs 11 A, 13 C). In contrast, the similar E. praelongipes lacks the protrusion in the septum (Fig. 10 D). The similar E. aequalis has a protrusion in the septum, but has short spermathecae, not reaching the frontal edge of the atria (Alderweireldt 1991).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	etymology	Etymology Named after Pinchas Amitai, an Israeli entomologist, writer and educator, who coined the Hebrew name for Evippa: Ş וחצר (Pronounced: rats’hol, meaning “ sand-runner ”).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	materials_examined	Material examined Holotype ISRAEL – Dead Sea Area • ♂; Ne’ot HaKikkar; [30.94 ° N, 35.37 ° E]; Nov. 2014; I. Renan leg.; HUJ INV-Ar 16051. Paratype ISRAEL – Dead Sea Area • 1 ♀; Ne’ot HaKikkar; [30.94 ° N, 35.37 ° E]; 3 – 27 Aug. 2014; I. Renan leg.; pitfall; HUJ INV-Ar 16048. Other material ISRAEL – Dead Sea Area • 1 ♂; Ne’ot HaKikkar; [30.94 ° N, 35.37 ° E]; 3 – 27 Aug. 2014; pitfall; I. Renan leg.; HUJ INV-Ar 16047 • 1 ♀; same collection data as for preceding; HUJ INV-Ar 16052 • 1 ♀; Sedom; [31.01 ° N, 35.35 ° E]; 21 Apr. 1954; A. Shulov leg.; HUJ INV-Ar 16053.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	description	Description Male MALE HOLOTYPE MEASUREMENTS. AME diameter: 0.15; PME diameter: 0.36; carapace length: 2.37; carapace width: 2.2; abdomen length: 2.34; leg I (Fe, Pa, Ti, Mt, Tr): 3.16, 1.21, 2.92, 3.07, 1.48; leg II: 2.95, 1.2, 2.8, 3.07, 1.54; leg III: 2.98, 1.05, 2.87, 3.67, 1.55; leg IV: 3.0, 1.04, 2.88, 3.77, 1.71. VARIATION IN MALES (n = 2). AME diameter: 0.15 – 0.16; PME diameter: 0.36 – 0.39; carapace length: 2.37 – 2.61; carapace width: 2.2 – 2.25; abdomen length: 2.1 – 2.34; legs I, III, IV of male HUJ INV-Ar 16047 are missing. Leg II (Fe, Pa, Ti, Mt, Tr): 2.99, 1.07, 2.9, 3.9, 1.6. CARAPACE. Transverse depression posterior to ocular area. CHELICERAL TEETH. 3 promarginal, 2 retromarginal. COLOR. Carapace yellow, lateral bands faint, brown-grey; margins spotted. Clypeus yellow to brown. Chelicerae yellow, darker retrolaterally. Legs yellow. Femora III – IV with grey dorsal bands. Palps yellow. Sternum yellow. Abdomen dorsum yellow, with dark spots on margins, cardiac mark dark. Abdomen venter yellow to dark yellow, sometimes with longitudinal lines. Spinnerets yellow to orange (Fig. 4 A). GENITALIA. Palpal organs strongly sclerotized. Part of sperm duct visible through tegulum as sinuous line. Tegulum bulging. Tegular apophysis large, curved ventrad at about 90 °, tip flat (Figs 6 A, 7 A, 8 A, 9, 12), best examined in distal view (Figs 9, 12 C). Terminal apophysis long, sharp, slightly curved, oriented distad. LEGS. Pseudoarticulation of tarsi present. Metatarsus I ventral spination: 3 pairs + apical triplet; tibia I ventral spination: 6 pairs + single weak retrolateral spine + apical pair. Female FEMALE PARATYPE MEASUREMENTS. AME diameter: 0.15; PME diameter: 0.37; carapace length: 3.18; carapace width: 2.46; abdomen length: 2.85; leg I (Fe, Pa, Ti, Mt, Tr): 2.6, 1.2, 2.8, 2.77, 1.4; leg II: 2.9, 1.0, 2.85, 2.69, 1.38; leg III: 3.0, 1.2, 2.7, 3.2, 1.5; leg IV: 4.2, 1.2, 3.98, 5.27, 2.0 VARIATION IN FEMALES (n = 3). AME diameter: 0.15 – 0.19; PME diameter: 0.35 – 0.46; carapace length: 3.14 – 3. 2; carapace width: 2.46 – 2.8; abdomen length: 2.85 – 4.37; leg I (Fe, Pa, Ti, Mt, Tr): 2.6 – 3.1, 1.2, 2.8 – 3.09, 2.77 – 2.8, 1.4 – 1.5; leg II: 2.9 – 3.19, 1.0 – 1.18, 2.85 – 2.9, 2.69 – 2.79, 1.35 – 1.38; leg III: 3.0 – 3.7, 1.2 – 1.25, 2.7 – 2.88, 3.2 – 3.5, 1.5 – 1.8; leg IV: 4.2 – 4.6, 1.2 – 1.37, 3.98 – 4.1, 5.6 – 5.27, 2.0. COLOR. Carapace yellow, lateral bands distinct, orange to brown, sometimes radiated. Clypeus yellow to brown. Chelicerae orange. Legs yellow. Legs I – II with faint grey spots. Femora III – IV with three dorsal bands. Palps yellow, darker towards tip, femora with dark stain. Sternum yellow. Abdomen dorsum yellow, with four brown, unfused chevrons (sometimes indistinct); dark cardiac mark. Abdomen venter whitish. Spinnerets whitish yellow with brown setae (Fig. 5 A). CARAPACE. Transverse depression posterior to ocular area. CHELICERAL TEETH. 3 promarginal, 2 retromarginal. GENITALIA. Epigyne as wide as long, septum approximately straight, mildly widening on proximal half, atria indistinct laterally. Spermathecae club-shaped, slightly longer than atria (Figs 10 A, 11 A, 13 C). LEGS. Pseudoarticulation of tarsi present. Metatarsus I ventral spination: 3 pairs + apical triplet; tibia I ventral spination: 6 pairs + apical pair. Natural history No live specimens observed. Adults collected in April, August and September (Table 3), from salt pans and oases (Fig. 16).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	distribution	Distribution Israel, southern Dead Sea area (Ne’ot HaKikkar, Sedom). (Fig. 16).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA45C12EFD6C1952773902D1.taxon	discussion	Remarks The species closely resembles the African species Evippa aequalis Alderweireldt, 1991. The Dead Sea area is known to be a refugium for tropical species (Yom-Tov & Tchernov 1988), mainly of African origin. We suggest that populations of E. amitaii sp. nov., or a closely related species, exist in suitable habitats south of Israel. This species does not conform to Roewer’s diagnostic character for Evippinae (metatarsus IV shorter than patella IV + tibia IV), as the females’ metatarsus IV (5.27 mm) is slightly longer than its patella and tibia IV (1.2 + 3.98 = 5.18 mm).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	description	Figs 3 A, 4 B, 5 B, 6 B, 7 B, 8 B, 10 B, 11 B	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	diagnosis	Diagnosis Recognized by a combination of genital characters. Male palp: Tegular apophysis parallel to cymbium, tip sharp, distally oriented. Subapical process blunt, located near base of TA. Terminal apophysis between tegular apophysis and cymbium (Figs 6 B, 7 B, 8 B). Epigyne: atria distinct, kidney shaped; septum wide, constricted proximally (Figs 10 B, 11 B). It is the only Evippinae in Israel to have kidney-shaped atria.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	materials_examined	Material examined ISRAEL – Coastal Plain • 1 ♀; Ben Zakkay; [31.86 ° N, 34.71 ° E]; 23 May 1973; faunistics course leg.; HUJ INV-Ar 16502 • 1 ♀; Holon; [31.999 ° N, 34.788 ° E]; 20 Jul. 2017; I. Armiach Steinpress leg.; HUJ INV-Ar 16509 • 1 ♀; Miqwe Yisra’el; [32.02 ° N, 34.77 ° E]; 11 Apr. 1938; A. Shulov leg.; HUJ INV-Ar 16076 • 1 ♀; Nahal Soreq (” Nahr Rubin ”); [31.94 ° N, 34.72 ° E]; 10 Apr. 1962; P. Amitai leg.; HUJ INV-Ar 16075 • 1 ♀ with eggs; Nizzanim; [31.75 ° N, 34.62 ° E]; 29 Sep. 1946; A. Shulov leg.; HUJ INV-Ar 16084 • 2 ♀♀; Nizzanim; [31.75 ° N, 34.62 ° E]; 29 Sep. 1946; A. Shulov leg.; HUJ INV-Ar 16085 to 16086 • 1 ♂; Nizzanim; 31.7432 ° N, 34.6249 ° E; 14 Jul. 2015; B. Shacham leg.; HUJ INV-Ar 16056 • 1 ♀; Nizzanim sands; 31.7265 ° N, 34.6059 ° E; 6 Jul. 2017; B. Shacham leg.; HUJ INV-Ar 16507 • 1 ♂; Nizzanim sands [31.7259 ° N, 34.6064 ° E]; 6 Jul. 2017; B. Shacham leg.; HUJ INV-Ar 16548 • 1 juv.; Palmahim; [31.91 ° N, 34.72 ° E]; 17 Oct. 2014; B. Shacham leg.; HUJ INV-Ar 16061 • 1 ♀; Ziqim sands; [31.61 ° N, 34.51 ° E]; 20 Jul. 2019; B. Shacham leg.; HUJ INV- Ar 20313. – Negev • 1 ♀; Ashalim; 30.9856 ° N, 34.6865 ° E; 6 Aug. 2015; HUJ INV-Ar 16058 • 1 ♂; Be’er Mash’abbim; [31.01 ° N, 34.76 ° E]; 15 Jul. 1990; Y. Ayal leg.; HUJ INV-Ar 16093 • 1 ♀; same locality as for preceding; 18 Sep. 1990; Y. Lubin leg.; HUJ INV-Ar 16095 • 1 ♂; Be’er Mash’abbim; [31.01 ° N, 34.76 ° E]; 30 Apr. 1993; Y. Lubin leg.; HUJ INV-Ar 16094 • 1 ♀; Be’er Sheva road to Mizpe Ramon; [31.09 ° N, 34.82 ° E]; 24 Mar. 1954; A. Shulov leg.; HUJ INV-Ar 16067 • 1 ♂; Bor Mashash; [31.07 ° N, 34.84 ° E]; 5 Feb. 1970; faunistics course leg.; HUJ INV-Ar 16064 • 1 ♂; same locality as for preceding; 5 Feb. 1970; faunistics course leg.; sands; HUJ INV-Ar 16069 • 1 ♀ with eggs; same locality as for preceding; 5 Feb. 1970; faunistics course leg.; sands; HUJ INV-Ar 16073 • 1 ♂; Hatira Ridge; [30.93 ° N, 35.04 ° E]; 20 Jun. 1991; Y. Lubin leg.; HUJ INV-Ar 16099 • 1 ♀; same locality as for preceding; 23 Oct. 1991; Y. Lubin leg.; HUJ INV-Ar 16098 • 1 ♀; Holot Agur; [30.97 ° N, 34.40 ° E]; 26 Feb. 2013; I. Renan leg.; HUJ INV-Ar 16057 • 1 ♀ with egg; Makhtesh Ramon; [30.59 ° N, 34.83 ° E]; 25 Apr. 1954; A. Shulov leg.; HUJ INV-Ar 16079 • 2 ♀♀; Mamshit; [31.02 ° N, 35.06 ° E]; 24 Mar. 1954; A. Shulov leg.; HUJ INV-Ar 16065, 16077 • 1 ♀; Mamshit; [31.02 ° N, 35.06 ° E]; 15 Jun. 1973; Gershoni leg.; HUJ 16066 • 1 ♂; same collection data as for preceding; 16070 • 1 ♀; Mamshit; [31.02 ° N, 35.06 ° E]; 5 Jul. 2015; A. Uzan leg.; HUJ INV-Ar 16506 • 1 ♀; Mishor Yamin; [31.00 ° N, 35.10 ° E]; 11 May 2014; Y. Zvik leg.; HUJ INV-Ar 16060 • 1 ♂; Mizpe Ramon; [30.60 ° N, 34.80 ° E]; 9 Apr. 2015; HUJ INV-Ar 16505 • ♀; Nahal Ashan; [31.28 ° N, 34.74 ° E]; 10 May 2016; B. Shacham leg.; HUJ INV-Ar 16062 • 1 ♀, 1 ♂; same locality as for preceding; 30 Apr. 2018; B. Shacham leg.; HUJ INV-Ar 16549 to 16550 • 1 ♀; Nahal Hatira; [30.93 ° N, 35.03 ° E]; 21 Aug. 1990; Y. Ayal leg.; HUJ INV- Ar 16097 • 1 ♀; Nahal Sekher; [31.09 ° N, 34.82 ° E]; 22 May 1958; A. Shulov leg.; sands; HUJ INV-Ar 16087 • 1 ♂; same locality as for preceding; 15 Apr. 1969; faunistics course leg.; sands; HUJ INV-Ar 16068 • 1 ♀ with eggs; same locality as for preceding; 23 May 1972; M. Pener leg.; sands; HUJ INV-Ar 16072 • 1 ♀; same locality as for preceding; 29 May 1980; M. Pener leg.; sands; HUJ INV-Ar 16092 • 1 ♀; same locality as for preceding; 3 Jun. 1980; faunistics course leg.; sands; HUJ INV-Ar 16096 • 1 ♂; same locality as for preceding; 11 Apr. 1991; Y. Lubin leg.; sands; HUJ INV-Ar 16501 • 1 ♀; Nahal Sekher; 31.091 ° N, 34.8121 ° E; 18 Apr. 2015; B. Shacham leg.; Subadult; sands; HUJ INV-Ar 16063 • 1 ♀ with eggs; Nahal Sekher; 31.107 ° N, 34.8193 ° E; 6 Apr. 2017; B. Shacham leg.; HUJ INV-Ar 16503 • 1 ♀; Nahal Sekher; 31.1046 ° N, 34.8228 ° E; 6 Apr. 2017; B. Shacham leg.; HUJ INV-Ar 16504 • 1 ♀; Nahal Sekher; [31.09 ° N, 34.82 ° E]; 24 Mar. 2018; B. Shacham leg.; sands; HUJ INV-Ar 16552 • 1 ♀; Retamim; [31.05 ° N, 34.68 ° E]; 22 Mar. 2018; S. Aharon leg.; HUJ INV-Ar 16553 • 1 subadult ♀; Sede Boqer; [30.85 ° N, 34.78 ° E]; 14 Mar. 2016; T. Mei – Dan leg.; molted to maturity in lab; HUJ INV-Ar 16089 • 1 ♀, 1 ♂; Sede Zin; [30.85 ° N, 34.77 ° E]; 22 Jul. 1993; Y. Lubin leg.; HUJ INV-Ar 16090 to 16091 • 1 ♀; Yeroham; [30.98 ° N, 34.90 ° E]; 5 Apr. 1954; A. Shulov leg.; HUJ INV-Ar 16078 • 1 ♀; same locality as for preceding; 30 May 1957; M. Pener leg.; HUJ INV-Ar 16088 • 1 ♂; same locality as for preceding; 24 Apr. 2014; I. Armiach Steinpress leg.; HUJ INV-Ar 16059 • 1 ♀; same locality as for preceding; 10 Apr. 2019; Y. Zvik leg.; HUJ INV-Ar INVAr 20319 • 1 ♀; Ze’elim; [31.17 ° N, 34.55 ° E]; 9 Apr. 1967; P. Amitai leg.; HUJ INV-Ar 16071 • 2 ♀♀ with egg-sac; same locality as for preceding; 9 Apr. 1967; P. Amitai leg.; HUJ INV-Ar 16080 to 16081 • 2 ♀♀; same locality as for preceding; 9 Apr. 1967; P. Amitai leg.; HUJ INV-Ar 16082 to 16083 • 2 ♀♀; Ze’elim; [31.176 ° N, 34.556 ° E]; 21 Jul. 2015; I. Armiach Steinpress leg.; HUJ INV-Ar 16054 to 16055 • 1 ♂, 1 ♀; same locality as for preceding; 23 Jul. 2015; I. Armiach Steinpress leg.; HUJ INV-Ar 16045 to 16046. PALESTINE – Gaza strip • ♀; Jabaliya; [31.52 ° N, 34.48 ° E]; 10 Jul. 1972; M. Timtpulson leg.; HUJ INV-Ar 16074.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	biology_ecology	Natural history Adults of Evippa arenaria were collected from February through October, and egg-sacs are recorded from February to September (Table 3). Both nocturnal and diurnal activity were observed. Evippa arenaria is the most mesophilic species of the four Evippa species found in Israel. It is found in sand and loess substrate throughout the semi-arid and arid parts of the Negev (Figs 15 B, 16). It is also found in the Mediterranean climate zone, along the coastal dune strip (Figs 15 D, 16). In the northern edge of its distribution in Israel and Palestine, E. arenaria is a strict psammophile, found only on exposed sand, but in the Negev desert it appears to be more generalist in its edaphic preferences, and is even found on somewhat rocky hillsides.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	distribution	Distribution Algeria, Tunisia, Lybia, Chad and Egypt (Alderweireldt 1991); Israel, Palestine; the report from Syria (Reimoser, 1919) is doubtful.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	description	Records Israel: Negev (Ashalim, Be’er Mash’abbim, Bor Mashash Sands, Hatira Ridge, Holot Agur, Makhtesh Ramon, Mamshit, Mishor Yamin, Nahal Ashan, Nahal Sekher (Sands), Sede Boqer, Sede Zin, Yeroham, Ze’elim), Coastal Plain (Ben Zakkay, Miqwe Yisra’el, Nizzanim, Nahal Soreq (Wadi Rubin), Palmahim). Palestine: Gaza (Jabalyia) (Fig. 16).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA42C122FDAF19537099044E.taxon	discussion	Remarks Despite extensive sampling in the Negev in all seasons, no adult E. arenaria were collected from November to January. This may be due to seasonal changes in activity (see Discussion).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	description	Figs 3 C – D, 4 C, 5 C, 6 C, 7 C, 8 C, 10 C, 11 C	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	diagnosis	Diagnosis Recognized by short, robust legs (carapace longer than ¼ of leg I), coloration and its genital morphology. Prosoma of preserved specimens dark brown (other Evippinae in Israel are yellow after preservation). Male palp: tegular apophysis parallel to cymbium, with single, small, subapical process oriented ventrad (Figs 6 C, 7 C, 8 C). It is the only Evippa in Israel with a subapical process directed ventrad at 90 ° to the tegular apophysis. In the similar Evippa caucasica Marusik, Guseinov & Koponen, 2003, the base of the subapical process reaches the tip of the tegular apophysis, whereas in E. onager, the tip and the process are distinct. Epigyne: atria distinct, widest in middle. Septum narrow, slightly constricted in middle, broadened distally (Figs 10 C, 11 C).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	materials_examined	Material examined ISRAEL – Dead Sea Area • 1 ♂; Qalya; [31.74 ° N, 35.46 ° E]; 15 Feb. 1941; A. Shulov leg.; HUJ INV- Ar 16510. – Negev • 1 subadult ♀; Nahal Hiyyon; [30.191 ° N, 35.008 ° E]; 30 Jul. 2019; HUJ INV-Ar 20315 • 1 ♂; same collection data as for preceding; 30 Jul. 2019; HUJ INV-Ar 20316 • 1 ♀; Biq’at ‘ Uvda (‘ Uvda valley); [29.998 ° N, 34.973 ° E]; 18 Mar. 2018; E. Gavish-Regev leg.; HUJ INV-Ar 20314.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	description	Description Male MEASUREMENTS (n = 2, specimen HUJ INV-Ar 20316 (the smaller of the two) was raised in the laboratory and might not be representative of sizes found in wild populations). AME diameter: 0.24 – 0.2; PME diameter: 0.51 – 0.4; carapace length: 3.97 – 3.58; carapace width: 2.94 – 2.67; abdomen length: 2.81 – 3.4; leg I (Fe, Pa, Ti, Mt, Tr): 2.99 – 2.74 (rest missing in specimen HUJ INV-Ar 16510), 1.26, 2.38, 2.3, 1.27; leg II: 2.91 – 2.44, 1.27 – 1.17, 3.04 – 2.3, 2.91 – 2.4 (rest missing in specimen HUJ INV-Ar 16510), 1.19; leg III: 2.29 – 2.6, 1.38 – 1.22, 2.59 – 2.22, 3.51 – 2.86, 1.51 – 1.34; leg IV: 3.83 – 3.34, 1.64 – 1.37, 3.43 – 2.9, 5.13 – 4.13, 1.99 – 1.61. COLOR. Carapace brown, darker on margins and in ocular area, sparsely radiated. Clypeus center dark, margins white. Chelicerae dark yellow, striated brown. Legs yellow, dorsum with wide brown annulations. Palps yellow, tarsus darkest. Sternum brown to black. Abdomen dorsum brown to black with white setae. Abdomen venter black to brown, with sparse white setae. Spinnerets yellow (Fig. 4 C). Live specimens dark yellow; legs with faint annulations. CARAPACE. Transverse depression posterior to ocular area. CHELICERAL TEETH. 2 promarginal, 2 retromarginal. GENITALIA. Palpal organs weakly sclerotized. Tegulum flat. Part of sperm duct visible through tegulum makes two sinoid curves. Tegular apophysis large, somewhat transparent, oriented distad, with subapical process oriented ventrad. Subapical process sharp, distinct from sharp tip of tegular apophysis (Figs 6 C, 7 C, 8 C). LEGS. Pseudoarticulation of tarsi not evident, but tarsi curved. Metatarsus I ventral spination: 3 pairs + apical triplet; tibia I ventral spination: 5 pairs + apical pair. Female MEASUREMENTS. AME diameter: 0.2; PME diameter: 0.56; carapace length: 3.88; carapace width: 2.8; abdomen length: 4.44; leg I (Fe, Pa, Ti, Mt, Tr): 2.54, 1.37, 2.35, 1.94, 1.03; leg II: 2.55, 1.36, 2.26, 1.96, 0.96; leg III: 2.58, 1.31, 2.22, 2.39, 1.1; leg IV: 3.31, 1.47, 2.93, 3.9, 1.41. COLOR. Carapace blackish, with some orange setae and orange fringe of setae. Clypeus blackish. Chelicerae reddish brown. Legs yellow, dorsum annulated grey. Palps yellow. Sternum black. Abdomen dorsum dark brown. Abdomen venter grey, spotted yellow. Spinnerets yellow (Fig. 4 C). Live specimens yellow, legs mildly annulated (Figs 3 C – D). CARAPACE. Transverse depression posterior to ocular area. CHELICERAL TEETH. 3 promarginal, 2 retromarginal. GENITALIA. Epigyne septum wider posteriorly, margins with small tooth or terrace in middle. Atria distinct (Fig. 10 C), width variable (compare Šternbergs 1979). Spermathecae round, unbent, distinct from sperm ducts (Fig. 11 C). LEGS. No clear pseudoarticulation on tarsi. Metatarsus I ventral spination: 3 pairs + apical triplet; tibia I ventral spination: 6 pairs + apical pair.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	biology_ecology	Natural history Outside Israel inhabits steppes. In Israel inhabits hyperarid deserts. Specimens were found at night, on and in the cracks of fine-grained floodplain deposits (Fig. 15 C). Adult female collected in March, adult male collected in February, juveniles collected in July (Table 3). The short legs, not typical of Evippinae, may be an adaptation for a somewhat fossorial lifestyle.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	distribution	Distribution Turkmenistan (Šternbergs 1979), Iran (Zamani et al. 2016), Israel.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	description	Records Israel: Dead Sea area (Qalya), Negev (Biq’at ‘ Uvda, Nahal Hiyyon) (Fig. 16).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4CC120FE1619C776B604EF.taxon	discussion	Remarks The female of Evippa onager was originally described from China by Simon (1895), and redescribed from both sexes by Šternbergs (1979) from Turkmenistan. It is very unlikely that these later specimens are conspecific with the holotype (fide Marusik et al. 2003). As we have not examined Simon’s type specimen, we avoided describing a new species and instead have chosen to treat the species as Evippa onager sensu Šternbergs, following Marusik et al. (2003). The species’ population in Israel is by far the southernmost and westernmost of the three known localities, and the only one not in a steppe habitat. Nevertheless, cool steppe habitats are found in Israel and it would be helpfull to search them for E. onager. See Phylogenetic relationships of Evippinae based on COI and NADH for a partial molecular phylogeny and a discussion of the placement of E. onager.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	description	Figs 3 B, 4 D, 5 D, 6 D, 7 D, 8 D, 10 D, 11 D, 14 A	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	diagnosis	Diagnosis Recognized by a combination of genital characters. Male palp: tegular apophysis parallel to cymbium, with small, prolaterally oriented subapical process. Short terminal apophysis positioned ventrad to tegular apophysis (Figs 6 D, 7 D, 8 D). Epigyne: straight atria with indistinct retrolateral margins. Septum with smooth margins, without protrusions (compare with similar E. amitaii sp. nov. palp, with a small protrusion) (Fig. 10 D, 11 D) It is the only Evippa in Israel to have a septum without protrusions.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	materials_examined	Material examined ISRAEL – Dead Sea Area • 1 ♀; Nahal ’ Ammi’az; 31.073 ° N, 35.342 ° E; 12 Apr. 2018; I. Armiach Steinpress leg.; HUJ INV-Ar 16554 • 1 ♀ with juv.; same collection data as for preceding; HUJ INV-Ar 16555 • 3 ♂♂; same collection data as for preceding; HUJ INV-Ar 16556 to 16558 • 2 ♀♀; Nahal ’ Azgad; 31.067 ° N, 35.335 ° E; 12 Apr. 2018; I. Armiach Steinpress leg.; HUJ INV-Ar 16559 – 16560 • 1 ♀; Sedom; [31.01 ° N, 35.35 ° E]; 21 Apr. 1954; A. Shulov leg.; HUJ INV-Ar 16518. – ’ Arava Valley • 1 ♂; ’ Avrona; [29.67 ° N, 35.00 ° E]; 2 – 8 Sep. 2016; I. Armiach Steinpress leg.; HUJ INV-Ar 16522 • 1 ♀; same collection data as for preceding; 7 Sep. 2016; I. Armiach Steinpress leg.; HUJ INV-Ar 16521 • 1 ♂, 1 ♀; same collection data as for preceding; 11 Sep. 2017; I. Armiach Steinpress leg.; HUJ INV-Ar 16534 to 16535 • 1 ♀; Elat; [29.57 ° N, 34.96 ° E]; 15 Apr. 1987; V. and B. Roth leg.; HUJ INV-Ar 16530 • 3 ♀♀; same collection data as for preceding; 18 Aug. 2016; A. Weinstein leg.; HUJ INV-Ar 16519 to 16520, 16044 • 1 ♀; ’ En Yotvata; [29.879 ° N, 35.044 ° E]; 25 Jan. 1958; A. Shulov leg.; HUJ INV-Ar 16517 • 1 ♂; Hazeva; 30.7139 ° N, 35.1964 ° E; 14 Jun. 2009; HUJ INV-Ar 16531 • 1 ♂; Hazeva; 30.8121 ° N, 35.2804 ° E; 2 Jul. 2010; HUJ INV-Ar 16532 • 1 ♂; Hazeva; 30.7129 ° N, 35.1964 ° E; 20 ul. 2010; HUJ INV-Ar 16533 • 1 ♂; Nahal Amazyahu N of ’ Iddan; [30.85 ° N, 35.29 ° E]; 15 Sep. 1988; B. Shalmon leg.; HUJ INV-Ar 16043 • 1 ♀; same collection data as for preceding; 16 Sep. 1988; B. Shalmon leg.; HUJ INV-Ar 16023 • 1 ♂; Nahal Amazyahu N of ’ Iddan; [30.85 ° N, 35.29 ° E]; 17 Sep. 1988; J. Koach leg.; HUJ INV-Ar 16524 • 1 ♀; Nahal Amazyahu; [30.85 ° N, 35.29 ° E]; 24 Sep. 1988; A. Weinstein leg.; HUJ INV-Ar 16529. – Negev • 1 ♀; Nahal Zevira; [30.64 ° N, 35.20 ° E]; 24 Feb. 2017; E. Gavish-Regev leg.; HUJ INV-Ar 16560 • 1 ♀; Biq’at ’ Uvda (’ Uvda valley), Har Shahrur; [29.980 ° N, 34.953 ° E]; 13 Mar. 2018; E. Gavish-Regev leg.; HUJ INV-Ar 16562 • 1 ♂; Biq’at ’ Uvda (’ Uvda valley), Kasuy Sands; [29.964 ° N, 34.979 ° E]; 13 Mar. 2018; E. Gavish-Regev leg.; HUJ INV-Ar 16563 • 1 ♂, 1 ♀; Biq’at ’ Uvda (’ Uvda valley), Nahal Hiyyon; [29.998 ° N, 34.973 ° E]; 13 Mar. 2018; E. Gavish-Regev leg.; HUJ INV-Ar 16564 to 16565 • 1 ♀; Shizzafon; [30.105 ° N, 35.002 ° E]; 24 Jul. 2014; I. Armiach Steinpress leg.; HUJ INV-Ar 16511 • 1 ♂, 1 ♀; Makhtesh Ramon; [30.59 ° N, 34.83 ° E]; 28 Mar. 1993; Y. Lubin leg.; HUJ INV-Ar 16527 to 16528 • 3 ♂♂; same collection data as for preceding; 21 Feb. 1994; Y. Lubin leg.; HUJ INV-Ar 16041, 16566 to 16567 • 1 ♀; same collection data as for preceding; 21 Feb. 1994; Y. Lubin leg.; HUJ INV-Ar 16042 • 1 ♂, 1 ♀; same collection data as for preceding; 8 Nov. 1992; Y. Lubin leg.; HUJ INV-Ar 16525 to 16526 • 1 ♀; Nahal Shizzafon; [30.042 ° N, 35.021 ° E]; 30 Jul. 2019; I. Armiach Steinpress leg.; HUJ INV-Ar 20317. EGYPT – Sinai • ♀; Abu – sela; 13 Aug. 1968; G. Tsabar leg.; HUJ INV-Ar 16512 • ♀; Mt. Catherine; [28.50 ° N, 33.95 ° E]; 16 Aug. 1968; G. Tsabar leg.; HUJ INV-Ar 16515 • ♀; Saint Catherine; [28.50 ° N, 33.95 ° E]; 15 Jul. 1968; A. Shulov leg.; HUJ INV-Ar 16513 • 2 ♂♂; Wadi Nequra; 21 Jan. 1969; S. Reichenstein leg.; HUJ INV-Ar 16514, 16516.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	biology_ecology	Natural history Inhabits hyper-arid deserts. Adults were collected January through April, June through September and in November (Table 3). Egg - sacs were observed in April and September. We suggest that E. praelongipes is active and reproducing throughout the year. Specimens were collected at night, mostly in dry stream beds, under and near shrubs (Fig. 15 A).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	distribution	Distribution Lybia to Saudi Arabia (Bonnet 1956; Alderweireldt & Jocqué 2017), Israel. Reports from Turkmenistan (Schmidt 1895), Pakistan and India (Tikader & Malhotra 1980) might be due to misidentification.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4EC121FE1A1A5E70DB002C.taxon	description	Records Israel: (Fig. 16) Dead Sea area (Sedom); ‘ Arava Valley (Elat, ‘ En Yotvata, ‘ Avrona, Nahal Amazyahu); Negev (Makhtesh Ramon, Shizzafon). Egypt: Sinai (Mt. Catherine, Wadi Nequra).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	description	Figs 1 – 2, 4 E, 5 E, 6 E, 7 E, 8 E, 10 E, 11 E, 13 B, D, 14 B	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	type_taxon	type species: Evippomma squamulatum (Simon, 1898).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	diagnosis	Diagnosis This is the only known genus of Lycosidae in which the body (mainly, the cephalothorax) is densely covered in scale-like, leaf-shaped setae (Figs 1 D, 14 B).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	description	Description Small- to medium-sized wolf spiders. Tibia I with 4 – 5 pairs of ventral spines (apart from apical pair). Body covered in scale-like, leaf-shaped setae. These setae are reflective and give live specimens a pearly shine (Figs 1 – 2). Coloration of preserved specimens is brown or yellow. Cephalic region elevated. Ocular area with long macrosetae. Genitalia relatively uniform across genus. Embolus large but not conspicuous. Embolic base in meso-apical position. Tegular apophysis hook-shaped (Figs 6 E, 7 E, 8 E). Median septum of epigyne widened posteriorly. Atria narrow (Figs 10 E, 13 B). Spermathecae large and sperm ducts twisted (Figs 11 E, 13 D). Anterior row of eyes procurved, narrower than PME. ALE smaller than AME.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	biology_ecology	Natural history The species inhabit deserts, grasslands and savannas. Evippomma rechenbergi is known to construct silk-lined burrows in sand (Bayer, Foelix & Alderweireldt 2017), as we recorded for E. simoni as well (Fig. 2 D).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	distribution	Distribution Found across Africa, with a new record from Israel (but see the section on Composition).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA4FC126FD811D65716E013D.taxon	discussion	Relationships Evippomma is traditionally considered to be related to Evippa (Alderweireldt 1992). Our molecular phylogeny supports this placement (see Phylogenetic relationships of Evippinae based on COI and NADH). Composition Seven species are included: Evippomma albomarginatum Alderweireldt, 1992; E. evippiforme (Caporiacco, 1935); E. evippinum (Simon, 1897); E. plumipes (Lessert, 1936); E. rechenbergi Bayer et al., 2017; E. simoni Alderweireldt, 1992; and E. squamulatum (Simon, 1898). Two representatives of the genus (E. evippiforme and E. evippinum) that were described from India are not considered here as belonging to Evippomma, as they have three pairs of ventral spines on tibia I (Caporiacco 1935), whereas Evippomma is defined as having 4 – 5 pairs. They might form a separate genus. Moreover, E. evippiforme appears to have genitalia uncharacteristic of the other known Evippomma species.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	description	Figs 1 – 2, 4 E, 5 E, 6 E, 7 E, 8 E, 10 E, 11 E, 13 B, D, 14 B	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	diagnosis	Diagnosis Recognized by a combination of genital characters. Male palp: tegular apophysis small, translucent, oriented distad, parallel to plane of bulb, laterally barb-shaped (Figs 6 E, 7 E, 8 E). Tegulum presents single sinus shape curve of sperm duct (in Evippa: two) (Fig. 6 E). Epigyne: rounded or hat-shaped, wider than long, septum wide. Atria narrow, shallow, greatly constricted distally (in Evippa: width approximately equal throughout), similar to epigyne of E. squamulatum, but proximal portion of atrium wide (in E. squamulatum proximal portion of atrium slit-shaped) (Figs 10 E, 13 B, D). It is the only Evippinae in Israel covered in flat, leaf-shaped setae and without annulations on the legs.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	materials_examined	Material examined ISRAEL – Dead Sea Area • 1 ♂; Near Sedom; [31.01 ° N, 35.35 ° E]; 10 Apr. 1967; P. Amitai leg.; HUJ INV-Ar 16537. – Negev • 1 ♀; Be’er Mash’abbim; [31.01 ° N, 34.76 ° E]; 18 Sep. 1990; Y. Lubin leg.; HUJ INV-Ar 16544 • 3 ♂♂; same collection data as for preceding; 11 Apr. 1991; Y. Lubin leg.; HUJ INV-Ar 16541, 16568 to 16569 • 2 ♂♂; same collection data as for preceding; 27 May 1992; Y. Lubin leg.; HUJ INV-Ar 16540, 16570 • 3 ♀♀; same collection data as for preceding; 25 Jun. 1992; Y. Lubin leg.; HUJ INV-Ar 16050, 16571 to 16572 • 4 ♂♂; same collection data as for preceding; 30 Apr. 1993; Y. Lubin leg.; HUJ INV-Ar 16049, 16573 to 16575 • 1 ♀; same collection data as for preceding; 30 Apr. 1993; Y. Lubin leg.; HUJ INV-Ar 16042 • 1 ♂; Holot Agur; [30.97 ° N, 34.40 ° E]; 1 Apr. 2012; I. Renan leg.; HUJ INV-Ar 16536 • 1 subadult ♀; same collection data as for preceding; 26 Feb. 2013; I. Renan leg.; HUJ INV-Ar 16576 • 1 juv.; same collection data as for preceding; 14 Mar. 2013; I. Renan leg.; HUJ INV-Ar 16545 • 1 ♀; Mash’abbim sands; 31.002 ° N, 34.754 ° E; 16 Jun. 2020; S. Aharon leg.; HUJ INV-Ar 20421 • 1 ♀; Mash’abbim sands; [31.00 ° N, 34.75 ° E]; 28 Jun. 2020; S. Aharon leg.; HUJ INV- Ar 20422 • 4 ♂♂; Nahal Sekher; [31.09 ° N, 34.82 ° E]; 5 May 1967; P. Amitai leg.; HUJ INV-Ar 16538 to 16539, 16577 to 16578 • 1 ♀; Nizzana; [30.89 ° N, 34.41 ° E]; 29 Aug. 1991; J. Henschel leg.; HUJ INV-Ar 16543 • 1 ♂; Mamshit; [31.02 ° N, 35.06 ° E]; 10 Apr. 2018; A. Uzan leg.; HUJ INV-Ar 16579.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	description	Description Male MEASUREMENTS (n = 12). AME diameter: 0.13 – 0.2; PME diameter: 0.27 – 0.4; carapace length: 2.8 – 3.99; carapace width: 2.1 – 2.8; abdomen length: 2.7 – 4.17; leg I (Fe, Pa, Ti, Mt, Tr): 2.77 – 4.7, 0.98 – 1.6, 2.39 – 5.96, 2.8 – 4.86, 1.7 – 2.3; leg II: 3.38 – 4.46, 1.2 – 1.57, 2.8 – 3.7, 3.57 – 4.5, 1.7 – 2.2; leg III: 3.2 – 4.3, 1.08 – 1.5, 2.27 – 2.9, 3.5 – 4.5, 1.78 – 2.25; leg IV: 3.8 – 5.26, 1.3 – 1.68, 3.9 – 5.1, 4.6 – 6.0, 2.14 – 2.58. COLOR. Carapace yellow to orange and brown, lighter around fovea and in patches behind PME; lateral bands brown to grey, radiated, fused posteriorly; ocular area black; cephalic and thoracic regions separated by sharp, black V-shaped line; perimeter with spots of white setae. Clypeus black near AME, light laterally, with white setae on margin. Chelicerae proximally yellow to orange with grey or brown reticulation connecting to oblique brown band in middle, retrolateral margin white. Legs yellow, proximal part of femur and coxa usually grey. Palps yellow to brownish (femur and tarsus darker). Sternum grey to brown, margin black, sometimes with light radiation. Abdomen dorsum yellow to whitish, sometimes with white spots. Cardiac mark yellow to brown, outlined with black spots, surrounded by posteriorly extending broken light band, outlined with dark spots, not reaching spinnerets. Abdominal venter margins yellow to whitish, center irregularly bordered, grey to black with yellow spots. Book lungs white. Spinnerets yellow (Fig. 4 E). CARAPACE. Ocular area elevated. Fovea area slightly elevated. CHELICERAL TEETH. 3 promarginal, 2 retromarginal. GENITALIA. Palpal organs weakly sclerotized. Tegulum keel-shaped. Part of sperm duct visible through tegulum bent at 90 °. Tegular apophysis small, translucent, oriented distad, with subapical process oriented ventrad; slightly elevated over tegulum (Figs 6 E, 7 E, 8 E). LEGS. Tarsus without pseudoarticulation, but in some specimens tarsi bent. Metatarsus I ventral spination: 2 pairs of long spines (pair I not reaching pair II) + short apical triplet; tibia I ventral spination: 5 pairs (all long except 5 th pair) + short apical pair. Distance between pairs 2 – 3 greatest. Female MEASUREMENTS (n = 6). AME diameter: 0.1 – 0.19; PME diameter: 0.33 – 0.42; carapace length: 3.28 – 4; carapace width: 2.3 – 3.07; abdomen length: 3.57 – 5.1; leg I (Fe, Pa, Ti, Mt, Tr): 2.95 – 3.6, 1.24 – 1.56, 2.5 – 3.09, 1.84 – 2.31, 1.22 – 1.5; leg II: 2.64 – 3.18, 1.22 – 1.3, 2.15 – 2.47, 1.69 – 2.2, 1.1 – 1.3; leg III: 2.46 – 3.14, 1.1 – 1.26, 1.5 – 2.9, 1.9 – 3.1, 1.37 – 1.6; leg IV: 2.99 – 3.86, 1.28 – 1.58, 2.79 – 3.8, 2.69 – 3.25, 1.48 – 1.78. COLOR. Carapace yellow to brownish-orange, covered with white setae, lighter around fovea and in patches behind PME; lateral bands grey, radiated, fused posteriorly; cephalic and thoracic regions separated by sharp, black V-shaped line. Clypeus yellow to brown, covered with leaf-like setae, with dark lateral bar. Chelicerae orange, with leaf-like setae on proximal half, sometimes with grey reticulation and oblique grey band. Legs yellow, usually, greyish on proximal side of segment, sometimes with grey line on dorsal side. Palps yellow, with some grey on femur and tarsus. Sternum brown. Abdomen dorsum yellow to whitish. Cardiac mark grey, usually with two lines of gray spots, connecting above spinnerets. Abdomen venter yellow, grey or white, darker on center. Book lungs white. Spinnerets yellow (Fig. 5 E). CARAPACE. Ocular area elevated. Due to the females’ strongly raised cephalic region and short legs they resemble a juvenile Lycosa rather than an Evippa. CHELICERAL TEETH. 3 promarginal, 2 retromarginal. GENITALIA. Epigyne wider than long, septum wide, atria distinct, shallow, narrow, widening proximally. Spermathecae longer than atria, bent retrolaterally, shaped like a crookneck squash (Figs 10 E, 11 E, 13 B, D). LEGS. No pseudoarticulation of the tarsus. Metatarsus I ventral spination: 2 long pairs (pair I reaching base of pair II); tibia I ventral spination: 5 pairs (long, except 5 th) + short apical pair.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	biology_ecology	Natural history Adult Evippomma simoni were collected April through June, August and September (Table 3) and may have a yearly cycle similar to what was suggested for E. arenaria, with a period of lowered activity in the colder months of the year (see Discussion). A female with eggs (HUJ INV-Ar 20422) was collected in June. The species mostly inhabits sandy desert habitats (Fig. 15 B). An anecdotal report raises the possibility that the species also exists along the coastal dune strip, similar to Evippa arenaria (Y. Salaviz, pers. com.). Specimen HUJ INV-Ar 20421 was found by S. Aharon in a silk-lined burrow (Fig. 2), similar to what has been reported for E. rechenbergi (Bayer et al. 2017).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	distribution	Distribution Sudan (Alderweireldt 1992), Israel. Israel: Negev (Be’er Mash’abbim, Holot Agur, Nahal Sekher, Mamshit), Dead Sea area (Near Sedom) (Fig. 16).	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
03B787E5BA48C124FDD11C76763203AB.taxon	discussion	Remarks This is the first record of Evippomma in Israel, and may also be the first record outside of Africa (depending on the dubious report of the species from India (Roewer 1955 )). If there is a continuous distribution with the type locality at the northern Sudanese border, this species is to be present in the sandy desert of northern Sinai and in eastern Egypt. It may be distributed in other hot desert environments in the region, such as the sandy deserts of Jordan. Most of the specimens were collected in dune areas, but a specimen from the Sedom area, with a clayrich substrate, suggests that this species is not an obligate psammophile. Phylogenetic relationships of Evippinae based on COI and NADH The ML tree topology recovered the genus Evippa as monophyletic (bootstrap support [BS] = 89 %). Evippinae was only moderately supported in our tree (Fig. 17; BS = 63 %), yet the topology in our tree accords with the tree topology for Lycosidae in Piacentini & Ramírez 2019 and with the composition of Evippinae sensu Alderweireldt 1991.	en	Steinpress, Igor Armiach, Alderweireldt, Mark, Cohen, Mira, Chipman, Ariel, Gavish-Regev, Efrat (2021): Synopsis of the Evippinae (Araneae, Lycosidae) of Israel, with description of a new species. European Journal of Taxonomy 733: 87-124, DOI: 10.5852/ejt.2021.733.1225
