identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4A564D22BA9C23D270FFA927595B782A.text	4A564D22BA9C23D270FFA927595B782A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipodarctus sp.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p>Taxon classification Animalia Arthrotardigrada Halechiniscidae</p>
            <p> Dipodarctus sp. Fig. 1 </p>
            <p>Material examined.</p>
            <p>Two four-clawed juveniles found in sediment sample 4 (Table 1).</p>
            <p>Remarks.</p>
            <p> The species resembles  Dipodarctus borrori Pollock, 1995 and  Dipodarctus susannae Jørgensen , Boesgaard,  Møbjerg &amp; Kristensen, 2014 by having digits of unequal length on legs  I–III and the lack of lateral processes between legs III and IV. It is distinguished from the two species by the lateral cirrus lack of scapus, which is present in the both  Dipodarctus borrori and  Dipodarctus susannae . It is also distinguished from  Dipodarctus borrori by the shorter digit 1 of legs  I–III and from  Dipodarctus susannae by its shorter papillate leg IV sense organ. These observations are based on comparing juveniles with descriptions of adults so while this species is probably an undescribed species observation of an adult specimen is required for confirmation. </p>
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	https://treatment.plazi.org/id/4A564D22BA9C23D270FFA927595B782A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Fujimoto, Shinta	Fujimoto, Shinta (2015): Halechiniscidae (Heterotardigrada, Arthrotardigrada) of Oura Bay, Okinawajima, Ryukyu Islands, with descriptions of three new species. ZooKeys 483: 149-166, DOI: http://dx.doi.org/10.3897/zookeys.483.8936, URL: http://dx.doi.org/10.3897/zookeys.483.8936
C79ABF49FC12CCEED17B6DD8B7AFD59D.text	C79ABF49FC12CCEED17B6DD8B7AFD59D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Florarctus wunai	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p>Taxon classification Animalia Arthrotardigrada Halechiniscidae</p>
            <p> Florarctus wunai sp. n. Figs 2, 3, Table 2 </p>
            <p>Diagnosis.</p>
            <p> Florarctus with smooth dorsal cuticle; six alae with continuous caestus; frontal ala with no caestus; antero-lateral alae caestus with small projection at levels of leg I and II and developed projection at posterior end; postero-lateral alae caestus with parallelogram-shaped projection at posterior end; caudal ala divided into four lobes; caudal alae caestus with pair of small projections; longitudinally elongate secondary clava with internally directed weak swelling. </p>
            <p>Material examined.</p>
            <p>Holotype: KUZ Z705: adult female found in sediment sample 3 (Table 1).</p>
            <p>Paratypes: KUZ Z706: adult female found in sediment sample 2 (Table 1); KUZ Z707: adult male found with holotype; KUZ Z708: four-clawed specimen of undetermined status found from sediment sample 5 (Table 1); KUZ Z709: four clawed juvenile found from sediment sample 2 (Table 1).</p>
            <p>Type locality.</p>
            <p>Water depth of 6 m, off Camp Schwab, Oura Bay, Okinawajima, one of the Ryukyu Islands, Japan (26°31'51.78"N, 128°3'10.74"E). Collected by the author on 25th January 2014.</p>
            <p>Type depository.</p>
            <p>The type series is deposited in the Zoological Collection of Kyoto University (KUZ).</p>
            <p>Description of holotype.</p>
            <p> Adult female, body length: 257  μm , excluding alae (Fig. 2A, 3A). Cephalic region clearly separated from trunk. Dorsal surface smooth with folds. Ventral surface smooth. Lateral margin of body surrounded by aliform expansions with continuous caestus, which consists of frontal ala, pair of antero-lateral alae, pair of postero-lateral alae and caudal ala. Frontal ala spreads across entire anterior margin of cephalic region. Scapi of internal cirri continuous with ala. Base of lateral cirri and primary clavae enveloped together in ala. Antero-lateral ala spreads from approximately level with median cirrus to level of leg III with four slight indentations. Antero-lateral alae caestus with small projections at level of leg I and leg II and developed projection at posterior end. Slightly overlapping antero-lateral ala, postero-lateral ala spreads from level of leg III to level of cirrus E with two indentations: anterior slight indentation and posterior relatively strong indentation. Postero-lateral alae caestus with developed projection parallelogram-shaped at posterior end. Caudal ala spread between pair of cirri E with pair of lateral indentations (26  μm deep) and medial indentation (40  μm deep). Caudal caestus with pair of small projections. Unpaired median cirrus (36  μm ) with scapus (10  μm ), tubular portion (22  μm ) and flagellum (4  μm ) inserted dorsally 27  μm from frontal margin. Pair of internal cirri (46  μm ) each with scapus (13  μm ) tubular portion (30  μm ) and flagellum (3  μm ) inserted at anterior margin. Internal structure of internal cirrus arise  25 μm from frontal margin. Pair of external cirrus (44  μm ) with scapus (18  μm ), tubular portion (20  μm ) and flagellum (6  μm ) inserted ventrally 30  μm from frontal margin. Internal structure of external cirrus arise 39  μm from frontal margin. Lateral cirrus (43  μm ) with scapus (15  μm ), tubular portion and flagellum and primary clava arise from same cirrophore. Boundary between tubular portion and flagellum of lateral cirrus indistinct in holotype. Lateral cirrus inserted dorso-posteriorly to primary clava. Primary clava (101  μm ) thicker near base with basal van der  Land’s body. Secondary clavae in shape of longitudinally elongated, flat sac with internally directed weak swelling on each side of ventrally protruded mouth cone (Figs 2B, 3B). Bucco-pharyngeal apparatus not visible except for pharyngeal bulb (32  μm × 24  μm ). No bacterial vesicles visible. Leg I sense organ (29  μm ) consists of tapering spine and distal flagellum. Leg II and leg III sense organ (28  μm and 24  μm respectively) each consists of unsegmented tapering spine. Leg IV sense organ (29  μm ) consists of tapering spine with basal van der  Land’s body, distal constricted portion and distal  pore . Pair of cirri E (46  μm ) each with proximal portion and flagellum arise from between postero-lateral and caudal alarum caesti. Rosette-like female gonopore opens 27  μm anterior to anus. Pair of seminal receptacles sited laterally at a level between gonopore and anus. Seminal receptacle consists of sinuous duct, which opens 29  μm laterally from gonopore and terminates in spherical vesicle 9  μm in diameter. Each  leg terminates in four digits with proximal wrinkles and distal claws. External digits with hook-shaped peduncle. Internal digits longer than external digits. Internal claws with dorsal spur. External claws with calcar and avicularia. Internal claws smaller than external claws. </p>
            <p>Etymology.</p>
            <p> The specific epithet, wunai, is a Ryukyuan word for  “sister” (Nakamoto 1981) referring to the new species as a sister of  Florarctus antillensis Van der Land, 1968, a species with similar morphology. </p>
            <p>Remarks on paratypes.</p>
            <p> The adult male, KUZ Z707, was smaller than adult females, KUZ Z705 and Z706, but had longer primary clavae relative to its body length (Table 2). The male gonopore of KUZ Z707 opens 10  μm anterior to the anus. The precise shape of male gonopore was not visible but spermatozoa were present inside the body. Excluding the lack of the genital structure, the paratypic four-clawed juvenile, KUZ Z709, was identical to the adults. A pair of bacterial vesicles is present in the paratypic four-clawed juvenile (Fig. 3C). For antero-lateral and postero-lateral alae, the number of slight indentations varied among specimens. There are two projections at the level of leg II in the paratypic specimen, KUZ Z708. The pair of small projections of the caudal alae caestus was better observed in the paratypes (Fig. 3D). </p>
            <p>Differential diagnosis.</p>
            <p> The presence of the continuous caestus and the absence of dorsal mammilla-like ornamentation, are shared by  Florarctus antillensis ,  Florarctus glareolus Noda, 1987,  Florarctus pulcher De Zio Grimaldi, Lamarca,  D’addabbo Gallo &amp; Pietanza, 1999 and  Florarctus wunai sp. n. The new species is distinguished from these three species by the two small projections of the caudal alae caestus, which are long projections in  Florarctus glareolus , long projections with swollen tips in  Florarctus pulcher and absent in  Florarctus antillensis (using Renaud-Mornant [1970] for information on the caestus morphology of  Florarctus antillensis ). </p>
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	https://treatment.plazi.org/id/C79ABF49FC12CCEED17B6DD8B7AFD59D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Fujimoto, Shinta	Fujimoto, Shinta (2015): Halechiniscidae (Heterotardigrada, Arthrotardigrada) of Oura Bay, Okinawajima, Ryukyu Islands, with descriptions of three new species. ZooKeys 483: 149-166, DOI: http://dx.doi.org/10.3897/zookeys.483.8936, URL: http://dx.doi.org/10.3897/zookeys.483.8936
60B2A3F543E3206113BD56759A2489AA.text	60B2A3F543E3206113BD56759A2489AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halechiniscus churakaagii	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p>Taxon classification Animalia Arthrotardigrada Halechiniscidae</p>
            <p> Halechiniscus churakaagii sp. n. Figs 4, 5, Table 2 </p>
            <p> Diagnosis . </p>
            <p> Halechiniscus with cephalic region consisting of antero-medial lobe and dorsal lobe; median cirrus inserted on long robust cirrophore; dorsal internal cirrus inserted on robust cirrophore; ventral external cirrus inserted on short cirrophore; lateral cirrus and primary clava inserted on large lateral cirrophore; large scapular process with flat oval tip; cirrus E with proximal portion with distal dark portion and distal flagellum; bipartite leg I sense organ; large, unsegmented legs II and III sense organs; papillate leg IV sense organ; all claws with calcar. </p>
            <p>Material examined.</p>
            <p>Holotype: KUZ Z710: adult female found in sediment sample 1 (Table 1).</p>
            <p>Paratype: KUZ Z711: adult female found in sediment sample 4 (Table 1).</p>
            <p>Type locality.</p>
            <p>Water depth of 6 m, off Sedake, Oura Bay, Okinawajima, one of the Ryukyu Islands, Japan (26°32'39.06"N, 128°2'52.8"E). Collected by the author on 24th January 2014.</p>
            <p>Type depository.</p>
            <p>The type series is deposited in the Zoological Collection of Kyoto University (KUZ).</p>
            <p>Description of holotype.</p>
            <p> Adult female, body length: 170  μm (Figs 4, 5A, B). Dorsal and ventral surface smooth. Cephalic region divided into two lobes: antero-ventrally protruded round medial lobe and dorsal lobe. Unpaired median cirrus with scapus (42  μm ) constricted at distal end, tubular portion (20  μm ) and flagellum (4  μm ) inserted on robust cirrophore (28  μm ); positioned dorsally 28  μm from frontal margin on dorsal lobe. Pair of internal cirri each with scapus (16  μm ), tubular portion (16  μm ) and flagellum (5  μm ) inserted on cirrophore; positioned dorsally on basal margin of medial lobe. Base of internal cirri arise postero-internal to base of scapi beneath cuticle. Pair of external cirri each with scapus (9  μm ), tubular portion (9  μm ) and flagellum (4  μm ) inserted on cirrophore; positioned ventrally on medial lobe. Base of external cirri arise posterior to scapi beneath cuticle. Lateral cirrus with scapus (35  μm ), tubular portion (23  μm ) and flagellum (10  μm ) and elongate primary clava (34  μm ) inserted on each large, lateral cirrophore; positioned slightly anterior to level of median cirrus on dorsal lobe. Primary clava with basal van der  Land’s body inserted antero-ventrally to lateral cirrus. Secondary clava absent. Mouth cone protruded antero-ventrally. Bucco-pharyngeal apparatus not visible except for pharyngeal bulb (17  μm × 19  μm ). Laterally protruded scapular process (42  μm )  with flat oval tip (Figs 4, 5C). No other process present. Cirrus E (86  μm ) segmented into proximal portion and distal flagellum. Under phase contrast microscopy distal end of proximal portion appears dark (Fig. 5D). This could be a distinct portion with accordion-like folds but scanning electron microscopy is required for confirmation. Rosette-like female gonopore opens ventrally 20  μm anterior to anus. Seminal receptacle ducts open postero-lateral to gonopore. Vesicles of seminal receptacles not visible. Leg I sense organ (11  μm ) consists of scapus and flagellum. Leg II and III sense organs (36  μm , 32  μm ) each consists of unsegmented, large spine. Papillate leg IV sense organ (16  μm ) with basal van der  Land’s body and terminal constriction inserted on slender cirrophore. Each leg terminates in digits with wrinkles and distal claws. All claws with small calcar. </p>
            <p>Etymology.</p>
            <p>The specific epithet, churakaagii, is a Ryukyuan word for "beautiful woman" (Tojo 1930) referring to the well-defined cephalic morphology of the new species.</p>
            <p>Differential diagnosis.</p>
            <p> The robust cirrophores of the median and lateral cirri are present in  Halechiniscus chafarinensis De Zio Grimaldi &amp; Villora Moreno, 1995,  Halechiniscus macrocephalus Grimaldi de Zio,  D’Addabbo Gallo &amp; Morone De Lucia, 1988,  Halechiniscus paratuleari Grimaldi de Zio,  D’Addabbo Gallo &amp; Morone De Lucia, 1988,  Halechiniscus tuleari Renaud-Mornant, 1979 and  Halechiniscus churakaagii sp. n. Among these species,  Halechiniscus paratuleari and the new species are the only species with large scapular processes. The new species is distinguished from  Halechiniscus paratuleari by the flat oval tip of the scapular process, which is acute in the latter and by the dark portion on cirrus E, which is absent in the latter. </p>
            <p> I have interpreted the dorsally positioned cirrus as internal cirrus and the ventrally positioned cirrus as external cirrus in the new species, which is opposite to the interpretation of these features in the original descriptions of  Halechiniscus tuleari and  Halechiniscus paratuleari . </p>
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	https://treatment.plazi.org/id/60B2A3F543E3206113BD56759A2489AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Fujimoto, Shinta	Fujimoto, Shinta (2015): Halechiniscidae (Heterotardigrada, Arthrotardigrada) of Oura Bay, Okinawajima, Ryukyu Islands, with descriptions of three new species. ZooKeys 483: 149-166, DOI: http://dx.doi.org/10.3897/zookeys.483.8936, URL: http://dx.doi.org/10.3897/zookeys.483.8936
92B4858E6BEF1971AED8D9E9179A61F6.text	92B4858E6BEF1971AED8D9E9179A61F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halechiniscus yanakaagii	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p>Taxon classification Animalia Arthrotardigrada Halechiniscidae</p>
            <p> Halechiniscus yanakaagii sp. n. Figs 6, 7, Table 2 </p>
            <p>Diagnosis.</p>
            <p> Halechiniscus with no distinct cephalic lobes; only lateral cirrus and primary clava inserted on cirrophore; laterally protruded arched, double processes with acute tips at level of leg I; unsegmented cirrus E; bipartite leg I sense organ; unsegmented legs II and III sense organs; papillate leg IV sense organ; claws of internal digits with dorsal spur. </p>
            <p>Material examined.</p>
            <p>Holotype: KUZ Z712: adult female found in sediment sample 5 (Table 1).</p>
            <p>Type locality.</p>
            <p>Water depth of 6 m, Off Thima, Oura Bay, Okinawajima, one of the Ryukyu Islands, Japan (26°32'0.81"N, 128°3'49.61"E). Collected by the author on 28th January 2014.</p>
            <p>Type depository.</p>
            <p>The holotype is deposited in the Zoological Collection of Kyoto University (KUZ).</p>
            <p> Description of holotype. </p>
            <p> Adult female, body length: 170  μm (Figs 6, 7A). Dorsal and ventral surface smooth. Cephalic region not divided into distinct lobes. Tubular portions of cephalic cirri indistinct from flagellum for median, internal and external  cirri . Unpaired median cirrus with scapus (11  μm ) and flagellum (24  μm ); inserted dorsally 16  μm from frontal margin. Pair of internal cirri each with scapus (6  μm ) and flagellum (20  μm ); inserted dorsally close to frontal margin. Pair of external cirri each with scapus (7  μm ) and flagellum (11  μm ); inserted ventrally close to frontal margin. Lateral cirrus with scapus (11  μm ), tubular portion (22  μm ) and flagellum (8  μm ) and elongate primary clava (21  μm ); inserted on each lateral cirrophore positioned at same level as median cirrus. Primary clava with basal van der  Land’s body inserted antero-ventrally to lateral cirrus. Secondary clava absent. Mouth cone protruded antero-ventrally. Bucco-pharyngeal apparatus not visible except for pharyngeal bulb (14  μm × 20  μm ). Laterally protruding arched, double processes (23  μm , 26  μm ) with acute tip situated dorsally at level of leg I (Figs 6, 7B). Unsegmented cirrus E (41  μm ) inserted on short cirrophore. Rosette-like female gonopore present. Seminal receptacles not visible. Leg I sense organ (14  μm ) consists of tapering spine and distal flagellum. Leg II and III sense organs (both 21  μm ) each consists of unsegmented spine. Papillate Leg IV sense organ (15  μm ) with basal van der  Land’s body inserted on small cirrophore. Each leg terminates in digits with wrinkles and distal claws. Claws of internal digits with dorsal spur. </p>
            <p>Etymology.</p>
            <p>The specific epithet, yanakaagii, is a Ryukyuan word for "ugly woman" (Tojo 1930) referring to dirty appearance of the holotype.</p>
            <p>Differential diagnosis.</p>
            <p> Halechiniscus
yanakaagii
 sp. n. and  Halechiniscus tuleari are the only two  Halechiniscus species with double processes at the level of leg I. The new species is distinguished from  Halechiniscus tuleari by the absence of distinct cephalic lobes and robust cephalic cirrophores (which are present in the latter species), the similar length, arched, double processes (23  μm , 26  μm ) in contrast with a short, straight, anterior process (holotype female: 8  μm ; paratype male: 4  μm ) and a long, straight, posterior process (holotype female: 19  μm ; paratype male: 10  μm ) (see: Renaud-Mornant 1979), and the absence of processes at level of leg II and III, which are present in  Halechiniscus tuleari . </p>
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	https://treatment.plazi.org/id/92B4858E6BEF1971AED8D9E9179A61F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Fujimoto, Shinta	Fujimoto, Shinta (2015): Halechiniscidae (Heterotardigrada, Arthrotardigrada) of Oura Bay, Okinawajima, Ryukyu Islands, with descriptions of three new species. ZooKeys 483: 149-166, DOI: http://dx.doi.org/10.3897/zookeys.483.8936, URL: http://dx.doi.org/10.3897/zookeys.483.8936
95C3D4036B604FC554039871808B022A.text	95C3D4036B604FC554039871808B022A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Styraconyx sp.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p>Taxon classification Animalia Arthrotardigrada Halechiniscidae</p>
            <p> Styraconyx sp. Fig. 8 </p>
            <p> Material examined. </p>
            <p>One female adult and one four-clawed juvenile found in sediment sample 4 (Table 1).</p>
            <p>Remarks.</p>
            <p> The individuals found resemble  Styraconyx
nanoqsunguak
 Kristensen &amp; Higgins, 1984 by the dorsal ridges (Fig. 8A, B). However, these specimens are distinguished by the lateral cirrus with no scapus (which is present  Styraconyx nanoqsunguak ), longer peduncles of the external digits and leg IV sense organs consisting of a spherical papilla and a distal spine (which is an elongate papilla and a shorter distal spine in  Styraconyx nanoqsunguak ). While I believe this is a new undescribed species, lack of visible taxonomic characters has hindered providing a complete species description. </p>
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	https://treatment.plazi.org/id/95C3D4036B604FC554039871808B022A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Fujimoto, Shinta	Fujimoto, Shinta (2015): Halechiniscidae (Heterotardigrada, Arthrotardigrada) of Oura Bay, Okinawajima, Ryukyu Islands, with descriptions of three new species. ZooKeys 483: 149-166, DOI: http://dx.doi.org/10.3897/zookeys.483.8936, URL: http://dx.doi.org/10.3897/zookeys.483.8936
