taxonID	type	description	language	source
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	description	(Figs 15 – 26)	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	materials_examined	Type locality. Yemen, Socotra Island, wadi Dineghen, 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m a. s. l. Type material. HOLOTYPE: J (NMPC), ‘ YEMEN, SOCOTRA island / wadi DENEGEN, 6 km SE / Hadibo; 140 m a. s. l. / 12 ° 36 ʹ 42 ʺN; 54 ° 03 ʹ 41 ʺE / 4. V. 2004 lgt. A. REITER [printed] ’. PARATYPES: 1 J 5 ♀♀, same label data as holotype (NMPC); 3 ♀♀, ‘ Yemen, Soqotra Is., 1. - 2. xii. 2003 / Dixam plateau, SIRHIN area / N 12 ° 31 ʹ 08 ″ E 53 ° 59 ʹ 09 ″ [GPS] / Jan Farkač lgt., 812 m // YEMEN – SOCOTRA 2003 / Expedition; Jan Farkač / Petr Kabátek & David Král [printed] ’ (NMPC); 1 ♀, ‘ YEMEN, SOCOTRA island / wadi FAR / 45 m a. s. l. / 12 ° 25 ʹ 59 ″ N; 54 ° 11 ʹ 42 ″ E / 14. V. 2004 lgt. A. REITER [printed] ’ (NMPC); 1 ♀, ‘ YEMEN, Socotra Island / wadi Ayhaft, 200 m / 12 º 36.5 ʹN, 53 º 58.9 ʹE / Jiří Hájek leg. 7 - 8. xi. 2010 // used for DNA extraction / voucher number IBE-RA 625 [printed] ’ (IBEB); 3 JJ 2 ♀♀, ‘ YEMEN, Socotra Island / wadi Madar, 1180 – 1230 m / 12 ° 33.2 ʹN, 54 ° 00.4 ʹE, / Jiří Hájek leg. 12 – 14. xi. 2010 ’ (IBEB, NMPC) [one male with additional label: ‘ used for DNA extraction / voucher number IBE-RA 612 [printed] ’ (IBEB)].	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	description	Description of male holotype. Habitus. Body shape globose, rounded. Pronotum trapezoidal, broadest basally; sides of pronotum regularly rounded. Maximum width of pronotum distinctly smaller than maximum width of body, angle between pronotum and elytra fairly distinct. Body surface shiny (Fig. 15). Colouration. Body colouring testaceous to ochraceous with dark (brownish-black) colour pattern. Head somewhat darkened posterior to eyes; pronotum largely darkened with broad longitudinal testaceous band along sides except posterior angles; elytra with separate rounded dark lateral subhumeral spot, and irregular longitudinal markings on disc (cf. Figs 15 – 16); legs testaceous with pro- and mesotarsomeres III, and metatarsomeres III – V darkened; ventral surface darkened except head and prosternum. Surface structure and sculpture. Head with indistinct microreticulation, chiefly anterior to eyes. Punctation fine, rather sparse, smaller on vertex, lacking posteriorly to eyes; punctures irregularly distributed, distance between punctures usually bigger than their diameter; punctures in frontolateral depressions somewhat coarser, and sometimes confluent. Clypeus broadly bordered, its anterior margin rounded (Fig. 17); frontolateral depressions shallow. Pronotum smooth, with shallow medial longitudinal scratch, and fine punctation; punctures sparser on disc and denser along lateral margins, punctation double, but differences in size of punctures rather small. Distance between punctures on disc much larger than their diameter; along margins distance between punctures approximately equal to diameter of coarse punctures. Sides of pronotum rounded, bordered except for anterior angles. Elytra smooth, without reticulation; punctation double: size differences small, coarser punctures much sparser than finer punctures, distance between punctures larger than their diameter. Discal puncture line well marked in basal half, other puncture lines almost imperceptible. Epipleura finely punctate. Ventral surface with reticulation only present laterally on head posterior to eyes; pro- and mesocoxae with areolate reticulation. Prosternal apophysis lanceolate, with longitudinal line of long golden setae medially. Metanepisternum (= metasternal wing) narrow, laciniate. Punctation of metaventrite consisting of fine punctures; punctation of metacoxal plates coarser and denser. Abdominal ventrites uniformly finely punctate; medially and along posterior margins with long golden setae. Apical ventrite with subapical carina raised medially to form distinct transverse swelling (Fig. 19); posterior margin with line of setigerous punctures, each with golden seta. Pro- and mesotarsi slightly broadened, with adhesive setae. Protrochanters incised. Median lobe of aedeagus in ventral view subparallel, only slightly narrowing to apex; apex deeply incised, with several short setae (Figs 21 – 22). Parameres (lateral lobes) triangularly shaped with apical tuft of setae (Fig. 23). Female. Similar to males (Fig. 16); pro- and mesotarsi not broadened; protrochanters not incised. Microreticulation of head more distinct and punctation of elytra finer than in male; disc of elytra very finely microreticulate, thus surface rather matt. Tubercle on apical ventrite absent. Female genitalia as in Figs 24 – 26. Variability. The specimens of the type series vary in the shape and extent of dark body colouration. Measurements: TL 3.9 – 4.6 mm (holotype 4.3 mm), TL-h 3.4 – 4.1 mm (holotype 3.7 mm); TW 2.5 – 3.0 mm (holotype 2.7 mm).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	diagnosis	Differential diagnosis. Based on the combination of the following characters, Hyphydrus dioscoridis sp. nov. belongs undoubtedly to the Hyphydrus signatus species group sensu BIS- TRÖM (1982): 1) longer spine of metatibia not serrate; 2) border of head foremargin without a furrow; 3) elytral punctation double; 4) metatibia externally with a distinct row of punctures; 5) median lobe symmetric in ventral view. The new species is similar to Hyphydrus pictus Klug, 1834 from the Arabian Peninsula and north-eastern Africa, but differs from this species in the significantly coarser, deeper and denser body punctation (distance between punctures equal to diameter of punctures or even less); clypeus more broadly bordered (cf. Figs 17 – 18); head between punctures microreticulated; tubercle on apex of apical ventrite without notch (with shallow notch in H. pictus (Fig. 20 )), and elytral disc of female microreticulate. Molecular data. The two barcode sequences of the specimens from wadi Madar and wadi Ayhaft were identical, but with a 5 % difference from the specimen from continental Yemen (33 out of 568 positions). This genetic difference is larger than that of many closely related species pairs of Dytiscidae (e. g. RIBERA & VOGLER 2004), and at a standard evolutionary rate of Adephagan Coleoptera would correspond to a separation of more than 1 million years (e. g. ANDÚJAR et al. 2012).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	etymology	Etymology. The new species is named after ‘ Dioscoridus’, an ancient Latin name of Socotra Island. The specific epithet is a noun in the genitive case, standing in apposition. Collection circumstances. The new species was collected in residual pools of drying-up streams and springs in wadis (e. g. Fig. 13).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF94FFB1DFF9F4CBCB1DFE00.taxon	distribution	Distribution. So far known only from several localities in Socotra Island, Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF91FFB1DFC6F448CC7DFCDA.taxon	distribution	Distribution. An eastern African species, spreading from Zambezi river in the south to Eritrea and mountains of south-west Arabia in the north (cf. BRINCK 1955).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF91FFB0DFC8F2C5CB3EFE39.taxon	materials_examined	Material examined. OMAN: DHOFAR: 2 ♀♀, wadi Heisher, 1. x. 2002, L. Kolářová leg. (JSCL, NMPC); 1 J, Taqah env., 270 – 350 m, 18. – 21. ix. 2003, R. Červenka leg. (JSCL). YEMEN: SOCOTRA ISLAND: Yemen, Socotra Island: 2 JJ 1 ♀, 6 km NE Hadibo, wadi Denegen, 12 ° 37 ʹ 01 ʺN, 54 ° 03 ʹ 50 ʺE, 10 – 100 m, 19. – 20. ii. 2000, V. Bejček & K. Šťastný leg. (CULS, NMPC); 1 J, same data, but 12 ° 36 ʹ 55 ʺN, 54 ° 03 ʹ 49 ʺE, 85 m, 27. xi. 2003, J. Farkač; 1 J 1 ♀, same data, but 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m, 4. v. 2004, A. Reiter leg.; 1 J, wadi Ayhaft, 15. iii. 2000, V. Bejček & K. Šťastný leg.; 2 JJ 5 ♀♀, same data, but 12 ° 36 ʹ 38 ʺN, 53 ° 58 ʹ 49 ʺE, 190 m, 24. – 26. xi. 2003, P. Kabátek leg.; 2 JJ, same data, but 12 ° 36.5 ʹN, 53 ° 58.9 ʹE, pool, 200 m, 8. xi. 2010, J. Hájek leg.; 1 J, wadi Faar, 12 ° 25 ʹ 59 ʺN, 54 ° 11 ʹ 42 ʺE, 69 m, 1. iv. 2001, V. Bejček & K. Šťastný leg.; 1 J, Hadibo env., 12 ° 36 ʹ 02 ʺN, 54 ° 02 ʹ 04 ʺE, 10 – 100 m, 21. xi. – 12. xii. 2003, J. Farkač leg.; 5 JJ 2 ♀♀, Homhil protected area, 12 ° 34 ʹ 27 ʺN, 54 ° 18 ʹ 32 ʺE, 364 m, 28. – 29. xi. 2003, D. Král leg.; 1 ♀, Qalansiyah env., N slopes of Khayrha Mts., 12 ° 38 ʹ 50 ʺN, 53 ° 27 ʹ 45 ʺE, 85 – 592 m, 9. – 10. xii. 2003, J. Farkač leg.; 1 J, Dixam plateau, wadi Esgego, 12 ° 28 ʹ 09 ʺN, 54 ° 00 ʹ 36 ʺE, 300 m, 2. – 3. xii. 2003, P. Kabátek leg.; 2 JJ, same data, but 13. v. 2004, A. Reiter leg.; 1 J 1 ♀, Dixam plateau, wadi Zeriq, 16. vi. 2009, V. Hula leg.; 2 ♀♀, Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Collected in pools of drying up streams in wadis.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF91FFB0DFC8F2C5CB3EFE39.taxon	distribution	Distribution. Widespread species, occurring almost in entire Africa south of Sahara, southern Algeria and Libya, and Egyptian oases; widely distributed also in Arabia (BRINCK 1955).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF90FFB0DF71F422CB5FFC51.taxon	description	(Figs 27 – 28) Published records. GAHAN (1985: 286): Yemen (as? Dineutus aereus); RÉGIMBART (1907: 142): Yemen (original description); data repeated by BALFOUR- BROWNE (1951: 195).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF90FFB0DF71F422CB5FFC51.taxon	materials_examined	Type material. SYNTYPES: 1 J, ‘ Type [round label with red margin, printed] // Hadramaut. [yellow underlined] / Arabia. / 94 - 235 [printed] // Dineutus / arabicus Rég. / n. sp. [Régimbart’s handwriting] ’ (BMNH); 1 ♀, ‘ ♀ [printed] // Hadramaut. [yellow underlined] / Arabia. / 94 - 235 [printed] // No [printed] 461 [handwritten] / examined by / Prof. Thaxter for / Laboulbeniaceae. [printed] // Dineutus / arabicus / co-type Rég. [handwriten] ’ (BMNH). Notes. A species known only from the type series from Hadramaut. With apical margin of elytra not serrate, it is more similar to Oriental D. indicus Aubé, 1838 rather than D. aereus. However D. arabicus differs from both mentioned species in absence of postero-lateral elytral sinuation (Fig. 27), deeply impressed elytral reticulation and different shape of male aedeagus (Fig. 28).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF96F7B3CD2DFD7A.taxon	description	(Fig. 29)	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF96F7B3CD2DFD7A.taxon	materials_examined	Material examined. YEMEN: AL HUDAYDAH GOV.: 1 ♀, Jabal Bura valley forest NP, 14 ° 52 ʹN, 43 ° 24 ʹE, 261 – 600 m, 9. – 11. iv. 2007, S. Kadlec leg. (NMPC). Notes. A single female (body length 15 mm) cannot be identified within any of the known Dineutus species from the region. It is characterised with relatively elongate habitus with sides of elytra only weakly rounded and feeble subapical elytral sinuation; greenish lustre of dorsal surface and well visible elytral striae. Finding more specimens, especially males, is necessary for solving its classification.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF35F6EBCFC3FEBF.taxon	distribution	Distribution. A species distributed in north-east Africa and the Arabian Peninsula (cf. BRINCK 1955).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF1EF4E1CC11FB94.taxon	materials_examined	Material examined. OMAN: DHOFAR: 1 J 1 ♀, Hasik, wadi 4 km to S, 17 ° 25 ʹN, 55 ° 17 ʹE, 46 m, 1. iv. 2012, A. Reiter leg. YEMEN: SOCOTRA ISLAND: 1 ♀, Dixam plateau, Dixam lake, 12 ° 31 ʹ 23 ʺN 53 ° 57 ʹ 12 ʺE, 1000 m, 12. v. 2004, A. Reiter leg.; 1 J 1 ♀, wadi Ireh, pool, 12 ° 23.4 ʹN, 53 ° 59.8 ʹE, 65 m, 13. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg.; 3 ♀♀, Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Similarly to D. aereus, collected in pools of drying up streams in wadis.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF1EF4E1CC11FB94.taxon	distribution	Distribution. Widespread African species, occurring through the continent except in the extreme south and north; abundant in Madagascar (BRINCK 1955). It is recorded also from Oman (BRANCUCCI 1985). Other records from the Arabian Peninsula and the Near East need to be verified. First record from Yemen (Socotra Island).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDFC9F25FCD76FAD3.taxon	distribution	Distribution. A species endemic to the Arabian Peninsula, so far recorded only from Iraq, Saudi Arabia and Yemen (cf. ROCCHI 1985).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF1CF367CCB5F98B.taxon	materials_examined	Material examined. YEMEN: AL MAHWIT GOV.: 1 spec., Al Ahjur, Badia vill. env., Al Khaltabi waterfall, 15 ° 28.4 ʹN, 43 ° 51.5 ʹE, 2500 m, 2. xi. 2010, J. Hájek leg. SANA’ A GOV.: 5 spec., 5 km SW Matnah vill., 15 ° 13 ʹ 41 ʺN, 43 ° 59 ʹ 40 ʺE, 2750 m, stream, 1. v. 2004, A. Reiter leg. (all NMPC). Habitat. Collected in high altitude streams (e. g. Fig. 4).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8FFFAFDF1CF367CCB5F98B.taxon	distribution	Distribution. Widely distributed West Palaearctic species, known from the Canary Islands, north Africa, most of Europe to the Near and Middle East (VONDEL 2005).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDF1FF53BCDBAFBE5.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 1 J 2 ♀♀, 5 km SW Matnah vill., 15 ° 13.7 ʹN, 43 ° 59.7 ʹE, 2750 m, stream, 19. xi. 2010, J. Hájek leg. (IBEB, NMPC). Habitat. Collected under stones in a mountainous stream densely covered with filamentous algae (Fig. 4).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDF1FF53BCDBAFBE5.taxon	distribution	Distribution. Widely distributed and common species in the whole Western Palaearctic, from the Canary Islands to north-west China (cf. NILSSON & HÁJEK 2013). First record from Yemen. Colymbetinae	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDFD8F733CCCDFD47.taxon	materials_examined	Material examined. OMAN: DHOFAR: 26 spec., 30 km E Salalah, 6 km E Taqah, wadi Darbat, pools near sea, 26. ii. + 1. iii. 1998, G. Wewalka leg. et det. (GWCW, NMPC, ONMM); 1 spec., 40 km E km Salalah, Wadi Darbat, upper valley, rivulet, 26. ii. 1998, G. Wewalka leg. et det. (CWCW); 3 spec., Jufa, haunt, 17 ° 08 ʹ 15 ʺN, 55 ° 01 ʹ 41 ʺE, 175 m, 29. x. 2009, A. Reiter leg.; 2 spec., wadi Darbat, N part near Shihayt, 17 ° 09 ʹN, 54 ° 28 ʹE, 325 m, 28. iii. 2012, A. Reiter leg.; 7 spec., 13 km E Mirbat, wadi Ain, 17 ° 01 ʹN, 54 ° 47 ʹE, 59 m, 29. iii. 2012, A. Reiter leg. YEMEN: ABIJAN GOV.: 7 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. (all NMPC). Habitat. Collected in relatively large and permanent pools on the wadi bottom with dense littoral vegetation and partially running water (e. g. Fig. 2), or in a shallow vegetated puddle under a livestock watering place.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDFD8F733CCCDFD47.taxon	distribution	Distribution. Afrotropical species occurring in whole Sub-Saharan Africa, north-easterly reaching Yemen and southern Oman in the Arabian Peninsula (cf. NILSSON 2005).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDFFAF27DCFE0FB0D.taxon	distribution	Distribution. A species occurring in Egypt, the Arabian Peninsula and Iran (NILSSON & HÁJEK 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFAEDFC5F336CE7DFA44.taxon	distribution	Distribution. A species described from Saudi Arabia and subsequently recorded also from Yemen (ROCCHI 1985).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFADDFD2F38FCDA0FE6C.taxon	materials_examined	Material examined. YEMEN: AL MAHWIT GOV.: 5 JJ 15 ♀♀, 10 km S Shibam, Aruz, 15 ° 27 ʹ 43 ʺN, 43 ° 55 ʹ 03 ʺE, 2560 m, 1. – 2. xi. 2007, A. Reiter leg.; 1 J 2 ♀♀, Al Ahjur, Badia vill. env., Al Khaltabi waterfall, 15 ° 28.4 ʹN, 43 ° 51.5 ʹE, 2500 m, 2. xi. 2010, J. Hájek leg. SANA’ A GOV.: 1 ♀, 5 km SW Matnah vill., 15 ° 13 ʹ 41 ʺN, 43 ° 59 ʹ 40 ʺE, 2750 m, stream, 1. v. 2004, A. Reiter leg.; 1 ♀, Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Collected in various stagnant (pools, tank) and running (stream) water bodies, both natural and manmade, but always at high altitude (2400 – 2750 m a. s. l.) (e. g. Figs 4 – 5).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8EFFADDFD2F38FCDA0FE6C.taxon	distribution	Distribution. A species occurring in north-east Africa and the Arabian Peninsula (BALKE 1992). Copelatinae	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8DFFADDFD6F7E6CE88FDB4.taxon	distribution	Distribution. A species occurring in eastern Africa north to Ethiopia, and in south-west Arabia (GUIGNOT 1961).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8DFFADDF86F4BFCD55FBB3.taxon	materials_examined	Material examined. OMAN: DHOFAR: 1 J, Mudhai oasis, spring, concrete pool, 17 ° 28 ʹ 38 ʺN, 53 ° 21 ʹ 09 ʺE, 548 m, 24. x. 2009, A. Reiter leg.; 4 spec., Sarfayt env., 16 ° 41 ʹ 23 ʺN, 53 ° 08 ʹ 02 ʺE, 580 m, 4. x. 2013, P. Kučera leg. (PKCL); 2 spec., 12 km NE of Sarfayt, 16 ° 44 ʹ 13 ʺN, 53 ° 13 ʹ 30 ʺE, 600 m, 4. x. 2013, P. Kučera leg. (PKCL). YEMEN: AL HUDAYDAH GOV.: 1 J, Jabal Bura valley forest NP, 14 ° 52 ʹN, 43 ° 24 ʹE, 225 – 600 m, 30. x. – 1. xi. 2005, P. Kabátek leg. HADRAMAUT GOV.: 1 ♀, W of Saywun, Shibam, 15 ° 54 ʹN, 48 ° 39 ʹE, 681 m, 4. iv. 2007, S. Kadlec leg.; 1 ♀, Al Khurabayah env., wadi Dawan, 15 ° 08.8 ʹN, 48 ° 25.6 ʹE, 1005 m, 19. – 20. x. 2005, D. Král leg. SANA’ A GOV.: 2 ♀♀, wadi Moor, 15 ° 08.0 ʹN, 43 ° 36.4 ʹE, 744 m, stream, 5. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Always associated with running water, usually in side pools of springs and streams.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8DFFADDF86F4BFCD55FBB3.taxon	distribution	Distribution. A species described from Ethiopia and subsequently recorded from Egypt (including Sinai Peninsula), Israel, Saudi Arabia and southern Oman (WEWALKA 1974, NILSSON & HÁJEK 2013). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8DFFACDE9FF2B8CD8CFE40.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 1 J 2 ♀♀, Dixam plateau, Sinhin area, 12 ° 31 ʹ 08 ʺN, 53 ° 59 ʹ 09 ʺE, 812 m, 1. – 2. xii. 2003, J. Farkač leg.; 5 JJ 5 ♀♀, Dixam plateau, Dixam lake, 12 ° 31 ʹ 23 ʺN 53 ° 57 ʹ 12 ʺE, 1000 m, 12. v. 2004, A. Reiter leg.; 4 JJ 6 ♀♀, Al Haghier Mts., wadi Madar, 12 ° 33.2 ʹN, 54 ° 00.4 ʹE, 1180 – 1230 m, springs, 12. – 14. xi. 2010, J. Hájek leg. (all NMPC). Notes. Matt females with striolate whole surface of elytra prevail in Socotra (Fig. 30) – the form was described by SHARP (1882) from Mesopotamia as a separate species under the name C. strigulosus, but later synonymised with C. pul- Fig. 30. Copelatus pulchellus (Klug, 1834), female chellus by BALFOUR- BROWNE (1939). habitus. Habitat. Collected in small pools of temporary stream, and in the littoral zone of larger permanent pool.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8DFFACDE9FF2B8CD8CFE40.taxon	distribution	Distribution. A species occurring most probably in entire Sub-Saharan Africa, the Arabian Peninsula and Iraq (GUIGNOT 1961). First record from Yemen (Socotra Island). Dytiscinae	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8CFFACDF27F79ACCB5FDA3.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 1 ♀, 60 km SW Sana’a, wadi Anis, 15 ° 00 ʹN, 44 ° 09 ʹE, 1522 m, 7. x. 2005, D. Král leg. (NMPC).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8CFFACDF27F79ACCB5FDA3.taxon	distribution	Distribution. A species known from Ethiopia and Sudan in eastern Africa, and Saudi Arabia and Yemen in the Arabian Peninsula (GUIGNOT 1961, NILSSON & HÁJEK 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8CFFACDE9CF4A8CC7EFA77.taxon	materials_examined	Material examined. OMAN: DHOFAR: 2 JJ, Al Mughsayl, 16 ° 53 ʹ 46 ʺN, 53 ° 46 ʹ 24 ʺE, 45 m, wadi, pools 2 km from seashore, 26. x. 2009, A. Reiter leg.; 1 J, Jufa, haunt, 17 ° 08 ʹ 15 ʺN, 55 ° 01 ʹ 41 ʺE, 175 m, 29. x. 2009, A. Reiter leg. (all NMPC); 1 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 380 m, 29. ix. 2011, P. Kučera leg. (PKCL); 7 spec., same data, but 24. viii. 2012 (PKCL); 1 spec., same data, but 13. iv. 2013, R. Fouqué leg. (JSCL); 2 spec., same data, but 10. x. 2013, P. Kučera leg. (PKCL); 2 JJ, 13 km E Mirbat, wadi Ain, 17 ° 01 ʹN, 54 ° 47 ʹE, 59 m, 29. iii. 2012, A. Reiter leg. (NMPC); 1 J, wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg. (NMPC); 1 spec., Ajdarawt, wadi Ra’s Sajir, 30 m, 22. viii. 2012, P. Kučera leg. (PKCL); 3 spec., wadi Muqshin, 19 ° 35 ʹ 06 ʺN, 54 ° 53 ʹ 04 ʺE, 9. iv. 2013, P. Kučera leg. (PKCL). YEMEN: ABIJAN GOV.: 1 ♀, Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. HADRAMAUT GOV.: 5 JJ 1 ♀, N of Al Mukalla, Nueima, 14 ° 36.4 ʹN, 49 ° 05.7 ʹE, 279 m, 20. – 21. x. 2005, D. Král leg. SOCOTRA ISLAND: 1 ♀, wadi Ayhaft, 12 ° 36 ʹ 38 ʺN, 53 ° 58 ʹ 49 ʺE, 190 m, 24. – 26. xi. 2003, P. Kabátek leg.; 1 J, same data, but 12 ° 36.5 ʹN, 53 ° 58.9 ʹE, pool, 200 m, 7. – 8. xi. 2010, J. Hájek leg.; 1 ♀, Homhil protected area, 12 ° 34 ʹ 27 ʺN, 54 ° 18 ʹ 32 ʺE, 364 m, 28. – 29. xi. 2003, P. Kabátek leg.; 2 JJ, 10 km E of Hadibo, Kam vill., 12 ° 33 ʹ 42 ʺN, 54 ° 07 ʹ 05 ʺE, 60 m, 5. v. 2004, A. Reiter leg.; 1 J, Dixam plateau, Dixam lake, 12 ° 31 ʹ 23 ʺN 53 ° 57 ʹ 12 ʺE, 1000 m, 12. v. 2004, A. Reiter leg.; 1 J 2 ♀♀, Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Collected in various aquatic habitats, mostly larger pools both natural and manmade (e. g. Figs 2, 14); frequently attracted also to light traps. In the Arabian Peninsula so far not documented from higher altitudes.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8CFFACDE9CF4A8CC7EFA77.taxon	distribution	Distribution. Cybister tripuncatus occurs in subtropical and tropical areas of the whole Old World. The subspecies C. t. africanus occurs in entire continental Africa, reaching south Europe in the north and Arabian Peninsula in the east (GUIGNOT 1961).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8CFFABDF2AF3FCCB4AFDCD.taxon	materials_examined	Published records. GAHAN (1985: 286): Yemen; BALFOUR- BROWNE (1951: 195): Yemen; ROCCHI (1985: 449): Yemen. Material examined. OMAN: DHOFAR: 1 J, Rakhyut, 0 – 50 m, 13. – 14. i. 2007, S. Jákl leg.; 1 J, Ain Tabruq, spring, 17 ° 06 ʹ 02 ʺN, 54 ° 19 ʹ 36 ʺE, 115 m, 28. x. 2009, A. Reiter leg.; 2 JJ, Hagarir, pasture, haunt, 16 ° 41 ʹ 34 ʺN, 53 ° 09 ʹ 21 ʺE, 658 m, 25. x. 2009, A. Reiter leg.; 2 JJ, 13 km E Mirbat, wadi Ain, 17 ° 01 ʹN, 54 ° 47 ʹE, 59 m, 29. iii. 2012, A. Reiter leg.; 1 ♀, wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg. (all NMPC); 3 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 380 m, 27. ix. 2011, P. Kučera leg. (PKCL); 1 spec., same data, but 13. iv. 2013, R. Fouqué leg. (JSCL). YEMEN: HADRAMAUT GOV.: 4 JJ 3 ♀♀, N of Al Mukalla, Nueima, 14 ° 36.4 ʹN, 49 ° 05.7 ʹE, 279 m, 20. – 21. x. 2005, D. Král leg.; 1 ♀, NE Ghayl Ba Wazir, 14 ° 47 ʹ 33 ″ N, 49 ° 22 ʹ 46 ″ E, 118 m, bottom of karstic abyss, 6. – 7. xi. 2007, A. Reiter leg. (all NMPC). Habitat. Collected in similar habitats as previous species with which it may occur syntopically. Distribution. Widely distributed Afrotropical species. Cybister vulneratus is known from entire Africa with the exception of Cape provinces, southern Spain and Sicily (Italy) in southern Europe, the Arabian Peninsula and Iran (GUIGNOT 1961, NILSSON & HÁJEK 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFABDF87F477CB78FCFF.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 1 ♀, 5 km N of Beni Al Salal, 15 ° 10.9 ʹN, 43 ° 23.4 ʹE, 350 m, artificial lights in petrol station, 4. – 5. xi. 2010, J. Hájek leg. (NMPC). Habitat. A single Yemeni specimen was collected at light.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFABDF87F477CB78FCFF.taxon	distribution	Distribution. A species recently revalidated and distinguished from E. sticticus. It occurs in most subtropical and tropical areas of the Old World, except Australia (cf. MILLER 2002).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFABDF83F565CE27FA83.taxon	materials_examined	Material examined. OMAN: DHOFAR: 2 ♀♀, Qitbit, spring, 19 ° 09 ʹ 20 ″ N, 54 ° 30 ʹ 31 ″ E, 164 m, 23. x. 2009, A. Reiter leg. (NMPC); 1 J 1 ♀, Difa, wadi, 16 ° 45 ʹN, 53 ° 14 ʹE, 435 m, 23. iii. 2012, A. Reiter leg. (NMPC); 1 ♀, Muqshin, oasis, at light, 19 ° 35 ʹN, 54 ° 53 ʹE, 155 m, 29. iii. 2011, A. Reiter leg. (NMPC); 1 spec., same data, but 9. iv. 2013, P. Kučera leg. (PKCL); 2 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 380 m, 27. ix. 2011, P. Kučera leg. (PKCL); 3 JJ, wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg. (NMPC); 1 J 1 ♀, Hasik, wadi 4 km to S, 17 ° 25 ʹN, 55 ° 17 ʹE, 46 m, 1. iv. 2012, A. Reiter leg. (NMPC); 1 spec., W of Al Mughsayl, wadi Raʹs Sajir, 16.84497 ° N, 53.68615 ° E, 20. – 31. viii. 2013, P. Kučera leg. (PKCL). YEMEN: AL MAHWIT GOV.: 1 J, 10 km S Shibam, Aruz, 15 ° 27 ʹ 43 ″ N, 43 ° 55 ʹ 03 ″ E, 2560 m, 1. – 2. xi. 2007, A. Reiter leg. (NMPC). SOCOTRA ISLAND: 1 ♀, Dixam plateau, Dixam lake, 12 ° 31 ʹ 23 ″ N 53 ° 57 ʹ 12 ″ E, 1000 m, 12. v. 2004, A. Reiter leg. (NMPC). Habitat. Collected predominantly in deeper pools in wadis, mostly with muddy bottom, and without vegetation; frequently attracted also to light trap.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFABDF83F565CE27FA83.taxon	distribution	Distribution. A cosmopolitan species occurring in the Old World in Africa including the Cape Verde and Canary Islands, southern Europe, and the Near and Middle East in Asia (MILLER 2002).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFAADF3AF348CC7EFEA0.taxon	materials_examined	Material examined. OMAN: DHOFAR: 2 JJ, 12 km NW Mirbat, wadi Hannah, 17 ° 03 ʹN, 54 ° 37 ʹE, 310 m, 30. iii. 2012, A. Reiter leg. (NMPC); 25 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 24. viii. 2012, P. Kučera leg. (JSCL, PKCL); 13 spec., same data, but 13. iv. 2013, R. Fouqué leg. (JSCL); 5 spec., same data, but 10. x. 2013, P. Kučera leg. (PKCL); 1 spec., wadi Al Mughsayl, 16 ° 52 ʹN, 53 ° 43 ʹE, 120 m, 10. iv. 2013, P. Kučera leg. (PKCL); 3 spec., Sarfayt env., 16 ° 41 ʹ 23 ʺN, 53 ° 08 ʹ 02 ʺE, 580 m, 4. x. 2013, P. Kučera leg. (PKCL). YEMEN: 1 J, Hadramaut (BMNH). Habitat. Found in a small concrete dam on a stream on the bottom of a forested wadi (Fig. 8), and in larger pools in vegetated wadis.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8BFFAADF3AF348CC7EFEA0.taxon	distribution	Distribution. A species described from Somalia and subsequently recorded also from Saudi Arabia (BRANCUCCI 1985). The single record of similar species H. pictus from Yemen (GAHAN 1895) belongs in fact also to H. africanus. First records from Oman.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8AFFA9DF1DF7ABCCFBFEE0.taxon	description	(Figs 31 – 34) Published records. GAHAN (1895: 286): Yemen (as Hydaticus histrio Clark, 1864); data repeated by BALFOUR- BROWNE (1951: 192) (as H. histrio); GUIGNOT (1951: 21) (original description of H. arabicus); and BRANCUCCI (1981: 229). Type material. HOLOTYPE: J, ‘ Hadramaut / Arabia / 94 - 235. [printed] // arabicus Rég. / n. sp. (Brit. Mus.) [handwritten] // TYPE [printed, red label] // HOLOTYPE [printed, red label] // Hydaticus / arabicus / Guign. Type [handwritten] ’ (MNHN). PARATYPE: J, ‘ Hadramaut / Arabia / 94 - 235. [printed] // J [printed] // Paratype [printed, red label with black frame] ’ (MNHN).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8AFFA9DF1DF7ABCCFBFEE0.taxon	materials_examined	Additional material examined. 1 J 1 ♀, with the same locality data (BMNH). Notes. A species known only from the type series from Hadramaut, Yemen, originally from BMNH. It was first studied by M. Régimbart, who kept two specimens for his collection, but did not manage to describe it. Later, F. Guignot discovered two males in Régimbart’s collection (MNHN) with manuscript name “ Hydaticus arabicus ” and described it (GUIGNOT 1951) without knowledge of two additional specimens still in BMNH. Here we present those additional specimens, one of which is the only known female. Whereas all males have reduced longitudinal yellow stripes on disc of elytra (Fig. 31), female has stripes complete (Fig. 33), much resembling the widely distributed Oriental Hydaticus histrio. However, both species differ in shape of median lobe of aedeagus, whose apex is distinctly curved in H. arabicus (Fig. 32; see also BRANCUCCI 1981: fig. 8).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF8AFFA9DF1DF7ABCCFBFEE0.taxon	distribution	Distribution. Hadramaut region, Yemen. The occurrence of the species in Oman (cf. NILSSON & HÁJEK 2013) was based on an unpublished record by G. Wewalka, which was however misidentified H. satoi dhofarensis Pederzani, 2003 (G. Wewalka, pers. comm.).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DFD8F752CE44FD9D.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 1 J, wadi Dahr, 2199 m, at light, 5. x. 2005, P. Kabátek leg. (NMPC). Habitat. A single recorded specimen was collected at light near adrying up stream in a wadi (P. Kabátek, pers. comm.).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DFD8F752CE44FD9D.taxon	distribution	Distribution. A species occurring in north-east Africa (Egypt, Sudan) and western Arabia (cf. ROCCHI 1985).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DFC4F44DCC3CFCAD.taxon	materials_examined	Material examined. YEMEN: AL MAHWIT GOV.: 1 ♀, 10 km S Shibam, Aruz, 15 ° 27 ʹ 43 ʺN, 43 ° 55 ʹ 03 ʺE, 2560 m, 1. – 2. xi. 2007, A. Reiter leg. SANA’ A GOV.: 2 JJ, Beni Mansour vill. env., 15 ° 06.1 – 4 ʹN, 43 ° 52.8 – 53.2 ʹE, 1520 – 1550 m, stream valley, 3. + 5. + 19. xi. 2010, J. Hájek leg.; 1 J, Bab Bahel, 15 ° 07.0 ʹN, 43 ° 40.9 ʹE, 1195 m, river valley and pool, 4. xi. 2010, J. Hájek leg. (all NMPC). Habitat. All recorded specimens were collected from pools at altitude higher than 1000 m a. s. l. Distribution. Widely distributed species, occurring from Madagascar through whole tropical Africa to Arabia (GUIGNOT 1961). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DFD1F585CC10FB05.taxon	materials_examined	Material examined. YEMEN: AL HUDAYDAH GOV.: 4 JJ 3 ♀♀, Jabal Bura valley forest NP, 14 ° 53 ʹN, 43 ° 26 ʹE, 557 m, 19. – 21. iii. 2007, P. Kabátek leg. (NMPC). Habitat. The specimens were collected at light near a drying up stream in a wadi (P. Kabátek, pers. comm.).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DFD1F585CC10FB05.taxon	distribution	Distribution. Widely distributed Afrotropical species. Recorded from Arabia under the name H. jucundus Reiche, 1849 (BRANCUCCI 1980, ROCCHI 1985).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DF12F33DCC83FA2A.taxon	materials_examined	Material examined. YEMEN: AL HUDAYDAH GOV.: 1 J, Jabal Bura valley forest NP, 14 ° 53 ʹN, 43 ° 26 ʹE, 557 m, 19. – 21. iii. 2007, P. Kabátek leg. (NMPC) Habitat. A single recorded specimen was collected at light near a drying up stream in a wadi (P. Kabátek, pers. comm.).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF89FFA9DF12F33DCC83FA2A.taxon	distribution	Distribution. Widely distributed Afrotropical species, occurring in whole sub-Saharan Africa and reaching Arabia in Asia (GUIGNOT 1961). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA8DF52F6EBCCC3FD52.taxon	materials_examined	Material examined. OMAN: DHOFAR: 1 ♀, 30 km E Salalah, 6 km E Taqah, Wadi Darbat, pool near sea, 26. ii. 1998, G. Wewalka leg. et det. (GWCW); 2 spec., wadi Darbat, 17 ° 05 ʹN, 54 ° 26 ʹE, 160 m, 9. x. 2013, P. Kučera leg. (PKCL); 1 ♀, Rakhyut, 0 – 50 m, 13. – 14. i. 2007, S. Jákl leg. (NMPC); 1 ♀, Difa, wadi, 16 ° 45 ʹN, 53 ° 14 ʹE, 435 m, 23. iii. 2012, A. Reiter leg. (NMPC); 9 spec., Ajdarawt, wadi Ra’s Sajir, 30 m, 22. viii. 2012, P. Kučera leg. (JSCL, PKCL); 2 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 380 m, 24. viii. 2012, P. Kučera leg. (PKCL); 4 spec., Sarfayt env., 16 ° 41 ʹ 23 ʺN, 53 ° 08 ʹ 02 ʺE, 580 m, 4. x. 2013, P. Kučera leg. (PKCL); 6 spec., 12 km NE of Sarfayt, 16 ° 44 ʹ 13 ʺN, 53 ° 13 ʹ 30 ʺE, 600 m, 4. x. 2013, P. Kučera leg. (PKCL). Habitat. Drying up pools in vegetated wadis on the SSE oriented slope of Jabal Al Qamar (Fig. 9).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA8DF52F6EBCCC3FD52.taxon	distribution	Distribution. Hydaticus satoi Wewalka, 1975 is a widely distributed Oriental species. Hydaticus s. dhofarensis is a subspecies endemic to southern Oman (PEDERZANI 2003).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA8DF1CF499CDBDFBF3.taxon	materials_examined	Material examined. OMAN: DHOFAR: 21 spec., 30 km E Salalah, 6 km E Taqah, Wadi Darbat, pools near sea, 24. ii. + 1. iii. 1998, G. Wewalka leg. et det. (GWCW, ONMM). YEMEN: SOCOTRA ISLAND: 1 J, wadi Ayhaft, 12 ° 36.5 ʹN, 53 ° 58.9 ʹE, pool, 200 m, 7. – 8. xi. 2010, J. Hájek leg. (NMPC). Habitat. Collected from larger residual pools of drying up streams in wadis.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA8DF1CF499CDBDFBF3.taxon	distribution	Distribution. This species is widely distributed in the whole Afrotropical Region, mainly in central Africa and Madagascar (GUIGNOT 1961). In the Arabian Peninsula known from the United Arab Emirates, Oman and Yemen (NILSSON & HÁJEK 2013). Hydroporinae	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA7DFC0F207CC20FEC0.taxon	materials_examined	Material examined. OMAN: DHOFAR: 8 spec., 30 km W Salalah, Wadis near Al Mughsayl, pools and lagoons, 27. ii. 1998, G. Wewalka leg. et det. (GWCW, ONMM); 2 spec., Qitbit, spring, 19 ° 09 ʹ 20 ″ N, 54 ° 30 ʹ 31 ″ E, 164 m, 23. x. 2009, A. Reiter leg. (NMPC). YEMEN: ABIJAN GOV.: 2 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. AL HUDAYDAH GOV.: 2 spec., ca. 12 km SE Al Mawkir vill., wadi Zabid, 14 ° 09 ʹ 35 ʺN, 43 ° 29 ʹ 33 ʺE, 264 m, 30. – 31. x. 2007, A. Reiter leg. LAHJ GOV.: 13 spec., W Lahj Al Hutah by road, wadi Am Rija, 13 ° 01 ʹ 57 ʺN, 44 ° 33 ʹ 30 ʺE, 297 m, 25. – 26. x. 2007, A. Reiter leg. SANA’ A GOV.: 2 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. TA’ IZZ GOV.: 2 spec., Najd an Nashamah env., Mashgab vill., 13 ° 21 ʹ 29 ″ N, 43 ° 57 ʹ 26 ″ E, 1182 m, 26. – 27. x. 2007, A. Reiter leg. SOCOTRA ISLAND: 1 spec., Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Collected predominantly from muddy littoral of larger pools, both natural and manmade (e. g. Figs 2, 5 – 7, 14).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF88FFA7DFC0F207CC20FEC0.taxon	distribution	Distribution. This species is widely distributed in the entire northern half of Africa (BISTRÖM 1986). In Asia it is known from Turkey, Iraq, the Arabian Peninsula, Iran and Pakistan (NILS- SON & HÁJEK 2013). First precise data from Oman – the occurrence of this species in Oman in Palaearctic catalogue (NILSSON & HÁJEK 2013) was based on unpublished records by G. Wewalka mentioned above. First record from Socotra Island.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFFBF70BCE36FCF4.taxon	materials_examined	Material examined. OMAN: DHOFAR: 7 spec., 30 km E Salalah, 6 km E Taqah, Wadi Darbat, pools near sea, 26. ii. 1998, G. Wewalka leg. et det. (GWCW, ONMM); 1 spec., 30 km W Salalah, Wadis near Al Mughsayl, pools and lagoons, 27. ii. 1998, G. Wewalka leg. et det. (GWCW). YEMEN: ABIJAN GOV.: 2 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. HAJJAH GOV.: 1 spec., NE Hajjah by road, Halhal vill. env., 15 ° 43 ʹ 42 ʺN, 43 ° 37 ʹ 25 ʺE, 998 m, 2. – 3. xi. 2007, A. Reiter leg. LAHJ GOV.: 16 spec., W Lahj Al Hutah by road, wadi Am Rija, 13 ° 01 ʹ 57 ʺN, 44 ° 33 ʹ 30 ʺE, 297 m, 25. – 26. x. 2007, A. Reiter leg. TA’ IZZ GOV.: 6 spec., Najd an Nashamah env., Mashgab vill., 13 ° 21 ʹ 29 ″ N, 43 ° 57 ʹ 26 ″ E, 1182 m, 26. – 27. x. 2007, A. Reiter leg. (all NMPC). Habitat. Habitat requirements identical with those of H. angularis; both species were often collected together.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFFBF70BCE36FCF4.taxon	distribution	Distribution. North African species, occurring predominantly in areas surrounding Sahara desert (cf. BISTRÖM 1986); known also from the Near and Middle East (cf. NILSSON & HÁJEK 2013). First precise data from Oman – the occurrence of this species in Oman in Palaearctic catalogue (NILSSON & HÁJEK 2013) was based on unpublished records of G. Wewalka mentioned above.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFD2F57FCE55FB16.taxon	materials_examined	Material examined. YEMEN: AL MAHWIT GOV.: 1 spec., Al Ahjur, Badia vill. env., Al Khaltabi waterfall, 15 ° 28.4 ʹN, 43 ° 51.5 ʹE, 2500 m, 2. xi. 2010, J. Hájek leg. SANA’ A GOV.: 12 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg.; 7 spec., 5 km SW Matnah vill., 15 ° 13.7 ʹN, 43 ° 59.7 ʹE, 2750 m, stream, 19. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Collected both in running (stream) and stagnant (tank) water bodies. All Yemeni records came from high altitude above 2400 m a. s. l. (Figs 4 – 5).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFD2F57FCE55FB16.taxon	distribution	Distribution. Widely distributed African species occurring in most areas of southern and eastern Africa and reaching the Arabian Peninsula in the north (BISTRÖM 1986, HÁJEK & WEWALKA 2009).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFF4F2ECCB79F9E4.taxon	materials_examined	Published records. BALFOUR- BROWNE (1951: 185): Yemen (as Guignotus major); ROCCHI (1985: 446): Yemen (as G. major); PEDERZANI (2003: 24): Dhofar; WEWALKA (2004: 465): Socotra (Samha); HÁJEK & WEWALKA (2009: 99): Dhofar. Material examined. OMAN: DHOFAR: 5 spec., Difa, wadi, 16 ° 45 ʹN, 53 ° 14 ʹE, 435 m, 23. iii. 2012, A. Reiter leg.; 6 spec., Hasik, wadi 4 km to S, 17 ° 25 ʹN, 55 ° 17 ʹE, 46 m, 1. iv. 2012, A. Reiter leg. YEMEN: SANA’ A GOV.: 5 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Yemeni specimens were collected in muddy littoral of a tank (Fig. 5). Omani specimens came from residual pools on the bottom of wadis (Fig. 9).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF87FFA7DFF4F2ECCB79F9E4.taxon	distribution	Distribution. The species occurs in the Saharan Africa from Niger, Algeria and Chad to the countries along the Red Sea (Egypt, Sudan, Eritrea, Djibouti, Somalia, Saudi Arabia and Yemen), reaching Israel in the north and southern Oman in the east (HÁJEK & WEWALKA 2009).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF86FFA6DFCAF6EBCC9FFCE2.taxon	materials_examined	Material examined. OMAN: DHOFAR: 10 spec., Muntasar, spring, 19 ° 27 ʹ 14 ʺN, 54 ° 37 ʹ 14 ʺE, 135 m, 23. x. 2009, A. Reiter leg.; 1 spec., Qitbit, spring, 19 ° 09 ʹ 20 ʺN, 54 ° 30 ʹ 31 ʺE, 164 m, 23. x. 2009, A. Reiter leg.; 2 spec., Hagarir, pasture, haunt, 16 ° 41 ʹ 34 ʺN, 53 ° 09 ʹ 21 ʺE, 658 m, 25. x. 2009, A. Reiter leg.; 10 spec., Muqshin, oasis, salty pools, 19 ° 35 ʹN, 54 ° 53 ʹE, 155 m, 29. iii. 2011, A. Reiter leg.; 3 spec., wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg. YEMEN: ABIJAN GOV.: 6 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. SOCOTRA ISLAND: 2 spec., 6 km SE of Hadibo, wadi Denegen, 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m, 4. v. 2004, A. Reiter leg.; 12 spec., 10 km E of Hadibo, Kam vill., 12 ° 33 ʹ 42 ʺN, 54 ° 07 ʹ 05 ʺE, 60 m, 5. v. 2004, A. Reiter leg.; 18 spec., Zemhon area, 12 ° 30 ʹ 58 ʺN, 54 ° 06 ʹ 39 ʺE, 270 – 350 m, at light, 3. – 4. ii. 2010, L. Purchart & J. Vybíral leg.; 3 spec., Noged plain (sand dunes), Sharet Halma vill. env., 12 ° 21.9 ʹN, 54 ° 05.3 ʹE, 20 m, at light, 10. – 11. xi. 2010, J. Hájek leg.; 3 spec., Qadub, coastal salt marsh, 12 ° 38.3 ʹN, 53 ° 57.3 ʹE, 8 m, 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Collected predominantly from puddles and small pools, often salty – both in desert and close to the sea shore (e. g. Figs 10 – 11); frequently attracted also to light trap.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF86FFA6DFCAF6EBCC9FFCE2.taxon	distribution	Distribution. Widely distributed African and Palaearctic species. In Sub-Saharan Africa, it is recorded from Senegal, Gambia, Mali, Sudan, Ethiopia, Djibouti, Somalia and Kenya (BISTRÖM 1986). In the Palaearctic Region, it occurs from North Africa and southern Europe throughout the Near East to Central Asia; in the east it reaches Tajikistan, Afghanistan and Pakistan (cf. NILSSON & HÁJEK 2013). First record from Socotra Island.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF86FFA6DF22F569CE33FA87.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 2 spec., Dixam plateau, Sinhin area, 12 ° 31 ʹ 08 ʺN, 53 ° 59 ʹ 09 ″ E, 812 m, 1. – 2. xii. 2003, J. Farkač leg.; 2 spec., Dixam plateau, wadi Esgego, 12 ° 28 ʹ 09 ʺN, 54 ° 00 ʹ 36 ʺE, 300 m, 2. – 3. xii. 2003, P. Kabátek leg.; 6 spec., same data, but 13. v. 2004, A. Reiter leg.; 3 spec., Dixam plateau, wadi Zerriq, 12 ° 31 ʹ 08 ʺN, 53 ° 59 ʹ 09 ʺE, 750 m, 3. xii. 2003, P. Kabátek leg.; 4 spec., 6 km SE of Hadibo, wadi Denegen, 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m, 4. v. 2004, A. Reiter leg.; 2 spec., Dixam plateau, Dixam lake, 12 ° 31 ʹ 23 ʺN, 53 ° 57 ʹ 12 ʺE, 1000 m, 12. v. 2004, A. Reiter leg.; 5 spec., wadi Ayhaft, 12 ° 36 ʹ 26 ʺN, 53 ° 59 ʹ 22 ʺE, 220 m, 16. v. 2004, A. Reiter leg.; 18 spec., same data, but pool, 7. – 8. xi. 2010, J. Hájek leg.; 3 spec., Aloove area, Hassan vill. env., 12 ° 31.2 ʹN, 54 ° 07.4 ʹE, 221 m, at light, 9. – 10. xi. 2010, J. Bezděk leg.; 2 spec., Dixam plateau, Firmihin (Dracaena forest), 12 ° 28.6 ʹN, 54 ° 01.1 ʹE, 490 m, at light, J. Bezděk leg.; 2 spec., Qadub, coastal salt marsh, 12 ° 38.3 ʹN, 53 ° 57.3 ʹE, 8 m, 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (all NMPC). Habitat. Collected frequently in puddles and both natural and manmade pools, from sea level to Dixam plateau (1000 m a. s. l.); found also in salty coastal marsh, and frequently attracted also to light trap.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF86FFA6DF22F569CE33FA87.taxon	distribution	Distribution. So far known only from the type series (WEWALKA 2004); a species endemic to Socotra Island.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF86FFA5DFC1F34DCB79FEA0.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 2 JJ, 1 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. (NMPC). Habitat. It was collected in muddy littoral of a tank (Fig. 5). Distribution. Species widely distributed in eastern and south-eastern Africa (cf. BISTRÖM 1988). Another species – Uvarus occultus (Sharp, 1882) – was described from Saudi Arabia, and may represent its junior synonym. However the single known specimen of U. occultus differs from U. peringueyi in slightly bigger and broader body and in almost uniformly ferruginous colouration of the dorsal surface (HÁJEK & BISTRÖM 2009). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF85FFA5DFA9F7ABCB44FD44.taxon	materials_examined	Material examined. YEMEN: ABIJAN GOV.: 8 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. LAHJ GOV.: 2 spec., Kadamat Al Abdali vill. env., wadi Tuban, 13 ° 07 ʹ 49 ʺN, 44 ° 51 ʹ 02 ʺE, 200 m, 24. – 25. x. 2007, A. Reiter leg. (all NMPC). Habitat. Collected at the edge of wadi pools with partially running water (e. g. Fig. 2). Distribution. Arabian species known only from Saudi Arabia and Yemen (BISTRÖM 1983).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF85FFA5DEBAF48FCDB4FB4C.taxon	materials_examined	Material examined. YEMEN: AL HUDAYDAH GOV.: 6 spec., Jabal Bura valley forest NP, 14 ° 52.4 – 5 ʹN, 43 ° 24. – 25.2 ʹE, 240 – 350 m, stream valley, 4. xi. 2010, J. Hájek leg. (NMPC). Notes. The specimens from Yemen have slightly more extensive yellow colour on pronotum: anterior half of pronotum yellow, except for thin blackish stripe along the anterior margin (Fig. 32); otherwi- se, they fit perfectly to the original description (BRANCUCCI 1985). Habitat. Collected in small gravelly residual pools of a temporary stream in a wadi (Fig. 3).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF85FFA5DEBAF48FCDB4FB4C.taxon	distribution	Distribution. A species described and so far known only from Saudi Arabia. First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF85FFA5DEF0F2F7CE5CFA40.taxon	distribution	Distribution. A species described from Sudan and subsequently recorded also from Yemen (BALFOUR- Fig. 35. Yola buettikeri Brancucci, 1985, habitus. - BROWNE 1951).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF85FFA4DFFDF38BCE4BFE0A.taxon	materials_examined	Material examined. YEMEN: AL HUDAYDAH GOV.: 4 spec., ca. 12 km SE Al Mawkir vill., wadi Zabid, 14 ° 09 ʹ 35 ʺN, 43 ° 29 ʹ 33 ʺE, 264 m, 30. – 31. x. 2007, A. Reiter leg. LAHJ GOV.: 13 spec., W Lahj Al Hutah by road, wadi Am Rija, 13 ° 01 ʹ 57 ʺN, 44 ° 33 ʹ 30 ʺE, 297 m, 25. – 26. x. 2007, A. Reiter leg. TA’ IZZ GOV.: 8 spec., Najd an Nashamah env., Mashgab vill., 13 ° 21 ʹ 29 ″ N, 43 ° 57 ʹ 26 ″ E, 1182 m, 26. – 27. x. 2007, A. Reiter leg. (all NMPC). Habitat. Collected in residual pools in wadis (e. g. Fig. 7) as well as manmade reservoirs out of the valley (Fig. 6); it seems to prefer habitats with muddy substrate, and without vegetation. Distribution. A species occurring in Africa in (semi) arid belt between Sahara desert and tropical rain forests, from Mauritania and Senegal to Sudan, Ethiopia and Djibouti (cf. BI- STRÖM 1983); in the Arabian Peninsula recorded from Saudi Arabia (BRANCUCCI 1985). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF84FFA4DFF2F431CE30FCE4.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 1 spec., Dixam plateau, wadi Esgego, 12 ° 28 ʹ 09 ʺN, 54 ° 00 ʹ 36 ʺE, 300 m, 2. – 3. xii. 2003, P. Kabátek leg.; 2 spec., same data, but D. Král leg.; 7 spec., same data, but 13. v. 2004, A. Reiter leg.; 4 spec., 6 km SE of Hadibo, wadi Denegen, 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m, 4. v. 2004, A. Reiter leg.; 10 spec., wadi Ayhaft, 12 ° 36.5 ʹN, 53 ° 58.9 ʹE, 200 m, pool, 7. – 8. xi. 2010, J. Hájek leg.; 4 spec., Al Haghier Mts., wadi Madar, 12 ° 33.2 ʹN, 54 ° 00.4 ʹE, 1180 – 1230 m, 12. – 14. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Collected in spring and stream pools in wadis originating from the Hagher Mountains. Distribution. A species endemic to Socotra Island, so far known only from the type series (WEWALKA 2004).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF84FFA4DFFEF57ECD84F9FC.taxon	materials_examined	Published records. BALFOUR- BROWNE (1951: 186): Yemen; ROCCHI (1985: 446): Yemen; BISTRÖM (1983: 60): Yemen. Material examined. OMAN: DHOFAR: 5 spec., Difa, wadi, 16 ° 45 ʹN, 53 ° 14 ʹE, 435 m, 23. iii. 2012, A. Reiter leg.; 1 spec., wadi Darbat, N part near Shihayt, 17 ° 09 ʹN, 54 ° 28 ʹE, 325 m, 28. iii. 2012, A. Reiter leg.; 8 spec., 13 km E Mirbat, wadi Ain, 17 ° 01 ʹN, 54 ° 47 ʹE, 59 m, 29. iii. 2012, A. Reiter leg.; 7 spec., wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg.; 18 spec., Hasik, wadi 4 km to S, 17 ° 25 ʹN, 55 ° 17 ʹE, 46 m, 1. iv. 2012, A. Reiter leg. YEMEN: ABIJAN GOV.: 1 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. AL HUDAYDAH GOV.: 3 spec., ca. 12 km SE Al Mawkir vill., wadi Zabid, 14 ° 09 ʹ 35 ʺN, 43 ° 29 ʹ 33 ʺE, 264 m, 30. – 31. x. 2007, A. Reiter leg.; 12 spec., Jabal Bura valley forest NP, 14 ° 52.4 – 5 ʹN, 43 ° 24. – 25.2 ʹE, 240 – 350 m, stream valley, 4. xi. 2010, J. Hájek leg. HAJJAH GOV.: 3 spec., NE Hajjah by road, Halhal vill. env., 15 ° 43 ʹ 42 ʺN, 43 ° 37 ʹ 25 ʺE, 998 m, 2. – 3. xi. 2007, A. Reiter leg. LAHJ GOV.: 10 spec., W Lahj Al Hutah by road, wadi Am Rija, 13 ° 01 ʹ 57 ʺN, 44 ° 33 ʹ 30 ʺE, 297 m, 25. – 26. x. 2007, A. Reiter leg. SANA’ A GOV.: 2 spec., Beni Mansour vill. env., 15 ° 06.1 – 4 ʹN, 43 ° 52.8 – 53.2 ʹE, 1520 – 1550 m, stream valley, 3. + 5. + 19. xi. 2010, J. Hájek leg.; 1 spec., wadi Moor, 15 ° 08.0 ʹN, 43 ° 36.4 ʹE, 744 m, stream, 5. xi. 2010, J. Hájek leg.; 1 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. TA’ IZZ GOV.: 2 spec., Najd an Nashamah env., Mashgab vill., 13 ° 21 ʹ 29 ʺN, 43 ° 57 ʹ 26 ʺE, 1182 m, 26. – 27. x. 2007, A. Reiter leg.; NNW Ash Shuqayrah by road, wadi Bani Khawlan, 13 ° 19 ʹ 57 ʺN, 43 ° 43 ʹ 19 ʺE, 460 m, 27. – 28. x. 2007, A. Reiter leg. (all NMPC). Habitat. Associated mostly with running water, especially with side pools of streams and rivers in wadis (e. g. Figs 2 – 3, 7); rarely also in pools (tank), especially at higher altitude (Figs 5 – 6). Yemeni localities came from lower parts of wadis up to 2400 m a. s. l. in the mountains; in Oman, Yolina insignis was found in pools in middle and lower parts of wadis on slopes of Jabal Al Qamar in western part of Dhofar to Jabal Jinawt on eastern coast of the province (e. g. Fig. 9).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF84FFA4DFFEF57ECD84F9FC.taxon	distribution	Distribution. A species occurring in eastern Africa and the Arabian Peninsula (Saudi Arabia, Yemen) (BISTRÖM 1983). First record from Oman.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DF8EF6EBCD76FE15.taxon	description	Notes. The species was first published by BALFOUR- BROWNE (1951) under the name Hydroporus inscitus Sharp, 1882; however, WEWALKA (1992) recognised the series from Yemen distinct from ‘ true inscitus ’ from Iraq and described it as a separate species.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DF8EF6EBCD76FE15.taxon	distribution	Distribution. A species endemic to Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DF8BF7C6CE22FD72.taxon	distribution	Distribution. A species with confirmed occurrence in the Arabian Peninsula and Egypt (TOLEDO 2009).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DFC3F4F9CCDAFC46.taxon	materials_examined	Material examined. OMAN: DHOFAR: 1 spec., Hagarir, pasture, haunt, 16 ° 41 ʹ 34 ʺN, 53 ° 09 ʹ 21 ʺE, 658 m, 25. x. 2009, A. Reiter leg. (NMPC). Habitat. Collected in a concrete tank at a livestock watering place (Fig. 10).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DFC3F4F9CCDAFC46.taxon	distribution	Distribution. A species described from northern Oman and subsequently recorded also from the United Arab Emirates, Iran and Afghanistan (HÁJEK & BRANCUCCI 2011).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DF21F5FCCB5DFB34.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 4 JJ 2 ♀♀, 5 km SW Matnah vill., 15 ° 13.7 ʹN, 43 ° 59.7 ʹE, 2750 m, stream, 1. v. 2004, A. Reiter leg., H. Fery det. (HFCB, NMPC); 2 J J 2 ♀♀, same data, but 19. xi. 2010, J. Hájek leg. (IBEB, NMPC). Habitat. Collected in a mountainous stream with gravelly bottom, densely covered with filamentous algae (Fig. 4), together with the following species.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DF21F5FCCB5DFB34.taxon	distribution	Distribution. A species known so far only from two localities in Yemen (TOLEDO 2009).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DFF2F32FCB78F9CC.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 2 JJ 3 ♀♀, 5 km SW Matnah vill., 15 ° 13.7 ʹN, 43 ° 59.7 ʹE, 2750 m, stream, 19. xi. 2010, J. Hájek leg. (IBEB, NMPC). Habitat. Collected in a mountainous stream with gravelly bottom, densely covered with filamentous algae (Fig. 4), together with the previous species.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF83FFA3DFF2F32FCB78F9CC.taxon	distribution	Distribution. Arabian species recorded only from Saudi Arabia and Yemen (TOLEDO 2009).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DFCEF4FECFA9FBF9.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 1 spec., Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (NMPC). Habitat. The single specimen recently collected in Socotra came from a large pool surrounded with Juncus socotranus marsh in a wadi (Fig. 14).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DFCEF4FECFA9FBF9.taxon	distribution	Distribution. Widely distributed species known from most of African territory except for Sahara, reaching also Mediterranean area and the Arabian Peninsula (cf. BISTRÖM & NILSSON 2002).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DFDFF262CB5FFB30.taxon	distribution	Distribution. Widely distributed Palaearctic species occurring in the whole Mediterranean area and subtropical parts of Asia eastward to western China and India (NILSSON & HÁJEK 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DF20F6EBCC3BFD75.taxon	materials_examined	Material examined. OMAN: DHOFAR: 32 spec., 30 km E Salalah, 6 km E Taqah, Wadi Darbat, pools near sea, 26. ii. 1998, G. Wewalka leg. et det.; 8 spec., 40 km E Salalah, Wadi Darbat, upper valley, rivulet, 26. ii. 1998, G. Wewalka leg. et det.; 8 spec., 30 km W Salalah, Wadis near Al Mughsayl, pools and lagoons, 27. ii. 1998, G. Wewalka leg. et det. (all GWCW); 1 J, Ain Tabruq, spring, 17 ° 06 ʹ 02 ″ N, 54 ° 19 ʹ 36 ″ E, 115 m, 28. x. 2009, A. Reiter leg.; 1 spec., Jufa, haunt, 17 ° 08 ʹ 15 ″ N, 55 ° 01 ʹ 41 ″ E, 175 m, 29. x. 2009, A. Reiter leg.; 4 spec., wadi Darbat, N part near Shihayt, 17 ° 09 ʹN, 54 ° 28 ʹE, 325 m, 28. iii. 2012, A. Reiter leg. (all NMPC). YEMEN: SOCOTRA ISLAND: 1 ♀, Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (NMPC). Habitat. In Oman collected in shallow vegetated puddles on the wadi bottom, below a karstic spring or below a livestock watering place, with partially running water; a single specimen from Socotra was found in a large pool surrounded with Juncus socotranus (Buchenau) Snogerup marsh in a wadi (Fig. 14).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DF20F6EBCC3BFD75.taxon	distribution	Distribution. Widely distributed species occurring in most areas of tropical Africa and Asia (cf. BISTRÖM 1997). First record from Yemen (Socotra Island).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DF09F33BCFE0F998.taxon	materials_examined	Material examined. YEMEN: HAJJAH GOV.: 1 spec., Thula castle, 15 ° 34 ʹ 32 ʺN, 43 ° 53 ʹ 54 ʺE, 2967 m, 2. xi. 2007, A. Reiter leg. SANA’ A GOV.: 2 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg.; 1 spec., 5 km SW Matnah vill., 15 ° 13.7 ʹN, 43 ° 59.7 ʹE, 2750 m, stream, 19. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Collected both in a mountainous stream (Fig. 4) and a manmade tank (Fig. 5), but always at a very high altitude exceeding 2400 m a. s. l.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF82FFA2DF09F33BCFE0F998.taxon	distribution	Distribution. A Palaearctic species distributed in the whole of Europe (except the most northern parts), northern Africa, and Asia eastward to Afghanistan and Kashmir (NILSSON & HÁJEK 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DFC8F6EBCFFBFDD5.taxon	materials_examined	Material examined. OMAN: DHOFAR: 3 spec., Muqshin, oasis, salty pools, 19 ° 35 ʹN, 54 ° 53 ʹE, 155 m, 29. iii. 2011, A. Reiter leg. (NMPC). Habitat. Recorded from a very isolated oasis in the desert separating two fertile parts of Oman. Pools without vegetation were situated in a depression with artesian springs; high salinity of water is proven by efflorescence of salts on the banks (Fig. 11).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DFC8F6EBCFFBFDD5.taxon	distribution	Distribution. A species occurring in arid areas from Egypt and the Arabian Peninsula, through the Near and Middle East to Central Asia (cf. NILSSON & HÁJEK 2013). First record from Oman.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DF94F41ECDD4FBB9.taxon	materials_examined	Published records. BALFOUR- BROWNE (1951: 183): Yemen; ROCCHI (1985: 445): Yemen; PEDERZANI (2003: 24): Dhofar. Material examined. OMAN: DHOFAR: 9 spec., 30 km E Salalah, 6 km E Taqah, Wadi Darbat, pools near sea, 26. ii. 1998, G. Wewalka leg. et det. (GWCW, ONMM); 1 J, Taqah env., 270 – 350 m, 18. – 21. ix. 2003, R. Červenka leg.; 8 spec., Mudhai oasis, spring, concrete pool, 17 ° 28 ʹ 38 ʺN, 53 ° 21 ʹ 09 ʺE, 548 m, 24. x. 2009, A. Reiter leg.; 1 ♀, Hagarir, pasture, haunt, 16 ° 41 ʹ 34 ʺN, 53 ° 09 ʹ 21 ʺE, 658 m, 25. x. 2009, A. Reiter leg.; 1 J, wadi Ash Shuwaymiyah, 17 ° 56 ʹN, 55 ° 32 ʹE, 31 m, 1. iv. 2012, A. Reiter leg. (all NMPC). YEMEN: AL BAYDA GOV.: 1 spec., NW Al Bayda by road, At Taghiq vill. env., 14 ° 08 ʹ 26 ʺN, 45 ° 25 ʹ 53 ʺE, 1968 m, 4. – 5. xi. 2007, A. Reiter leg. AL MAHWIT GOV.: 1 spec., 10 km S Shibam, Aruz, 15 ° 27 ʹ 43 ʺN, 43 ° 55 ʹ 03 ʺE, 2560 m, 1. – 2. xi. 2007, A. Reiter leg. SANA’ A GOV.: 12 spec., Beni Mansour vill. env., 15 ° 06.1 – 4 ʹN, 43 ° 52.8 – 53.2 ʹE, 1520 – 1550 m, stream valley, 3. + 5. + 19. xi. 2010, J. Hájek leg. (1 spec. used for DNA extraction, specimen voucher IBE-RA 609). TA’ IZZ GOV.: 10 spec., Najd an Nashamah env., Mashgab vill., 13 ° 21 ʹ 29 ″ N, 43 ° 57 ʹ 26 ″ E, 1182 m, 26. – 27. x. 2007, A. Reiter leg. (all NMPC). Habitat. A species most probably without special habitat requirements; found both in residual pools of drying up streams in wadis, and various natural and manmade stagnant water bodies (e. g. Figs 6, 10).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DF94F41ECDD4FBB9.taxon	distribution	Distribution. A species occurring in north-eastern Africa, the Arabian Peninsula and Iran (BISTRÖM 1982). Laccophilinae	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DF32F2B1CB16F9F4.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 1 J, Dixam plateau, wadi Zerig, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. (NMPC). Notes. Identification of this species has been verified by Olof Biström (Helsinki, Finland). The single Socotran male slightly differs from the African specimens in shape of median lobe (O. Biström, pers. comm. 2014). However, as the variability in median lobe is known also in African population of the species (O. Biström, pers. comm. 2014), and only a single specimen is known from Socotra, we prefer to maintain the species as L. continentalis. Habitat. The single specimen recently collected in Socotra came from a large pool surrounded with Juncus sp. marsh in a wadi (Fig. 14).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF81FFA1DF32F2B1CB16F9F4.taxon	distribution	Distribution. A species known from southern Africa, and eastern Africa north to Somalia (GUIGNOT 1959, OMER- COOPER 1965). First record from Yemen (Socotra Island).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FFA0DF70F6EBCCF0FE32.taxon	materials_examined	Material examined. OMAN: DHOFAR: 4 spec., Hagarir, pasture, haunt, 16 ° 41 ʹ 34 ʺN, 53 ° 09 ʹ 21 ʺE, 658 m, 25. x. 2009, A. Reiter leg. (NMPC); 1 spec., 15 km W of Jufa (Laga Shalia), 17 ° 11 ʹ 05 ʺN, 54 ° 56 ʹ 38 ʺE, 380 m, 24. viii. 2012, P. Kučera leg. (PKCL). Habitat. Collected in a concrete tank at a livestock watering place (Fig. 10), and in small pools in a wadi.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FFA0DF70F6EBCCF0FE32.taxon	distribution	Distribution. The nominotypical subspecies was described from northern Oman and subsequently recorded also from the United Arab Emirates (HÁJEK & BRANCUCCI 2011).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FFA0DF31F439CC60FC87.taxon	materials_examined	Material examined. YEMEN: SOCOTRA ISLAND: 1 ♀, Dixam plateau, Sinhin area, 12 ° 31 ʹ 08 ʺN, 53 ° 59 ʹ 09 ″ E, 812 m, 1. – 2. xii. 2003, J. Farkač leg.; 1 J, Dixam plateau, wadi Zerriq, 12 ° 31 ʹ 08 ʺN, 53 ° 59 ʹ 09 ʺE, 750 m, 3. xii. 2003, P. Kabátek leg.; 4 spec., same locality, pools, Juncus marsh, 12 ° 29.6 ʹN, 53 ° 59.5 ʹE, 655 m, 13. – 14. vi. 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg.; 8 spec., 6 km SE of Hadibo, wadi Denegen, 12 ° 36 ʹ 42 ʺN, 54 ° 03 ʹ 41 ʺE, 140 m, 4. v. 2004, A. Reiter leg.; 3 spec., Dresmoiten vill., wadi Darho, 12 ° 30 ʹ 18 ʺN, 54 ° 02 ʹ 08 ʺE, 365 m, 13. v. 2004, A. Reiter leg.; 5 spec., wadi Ayhaft, 12 ° 36.5 ʹN, 53 ° 58.9 ʹE, 200 m, pool, 7. – 8. xi. 2010, J. Hájek leg. (all NMPC). Habitat. Collected mostly in larger pools of drying up streams in wadis (e. g. Figs 12, 14). Distribution. Widely distributed Afrotropical species (cf. GUIGNOT 1959)	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FFA0DFC6F54CCFE0FAAB.taxon	materials_examined	Material examined. OMAN: DHOFAR: 4 spec., Difa, wadi, 16 ° 45 ʹN, 53 ° 14 ʹE, 435 m, 26. iii. 2012, A. Reiter leg. YEMEN: ABIJAN GOV.: 16 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. AL BAYDA GOV.: 1 spec., NW Al Bayda by road, At Taghiq vill. env., 14 ° 08 ʹ 26 ʺN, 45 ° 25 ʹ 53 ʺE, 1968 m, 4. – 5. xi. 2007, A. Reiter leg. AL HUDAYDAH GOV.: 9 spec., Jabal Bura valley forest NP, 14 ° 52.4 – 5 ʹN, 43 ° 24. – 25.2 ʹE, 240 – 350 m, stream valley, 4. xi. 2010, J. Hájek leg. HAJJAH GOV.: 3 spec., NE Hajjah by road, Halhal vill. env., 15 ° 43 ʹ 42 ʺN, 43 ° 37 ʹ 25 ʺE, 998 m, 2. – 3. xi. 2007, A. Reiter leg. LAHJ GOV.: 6 spec., W Lahj Al Hutah by road, wadi Am Rija, 13 ° 01 ʹ 57 ʺN, 44 ° 33 ʹ 30 ʺE, 297 m, 25. – 26. x. 2007, A. Reiter leg. SANA’ A GOV.: 6 spec., 60 km SW Sana’a, wadi Anis, 15 ° 00 ʹN, 44 ° 09 ʹE, 1522 m, 7. x. 2005, D. Král leg.; 2 spec., Beni Mansour vill. env., 15 ° 06.1 – 4 ʹN, 43 ° 52.8 – 53.2 ʹE, 1520 – 1550 m, stream valley, 3. + 5. + 19. xi. 2010, J. Hájek leg. TA’ IZZ GOV.: 2 spec., NNW Ash Shuqayrah by road, wadi Bani Khawlan, 13 ° 19 ʹ 57 ʺN, 43 ° 43 ʹ 19 ʺE, 460 m, 27. – 28. x. 2007, A. Reiter leg. (all NMPC). Habitat. Mostly associated with running water; especially in relatively permanent side pools of streams and river, mostly at lower and middle altitudes (ca. 150 – 1500 m) (e. g. Figs 2 – 3, 7, 9). Distribution. A species known from Ethiopia and Egypt in Africa, and Saudi Arabia and Yemen in the Arabian Peninsula (BRANCUCCI 1983, NILSSON & HÁJEK 2013). First record from Oman.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FF9FDFC4F390CCE9FEC0.taxon	materials_examined	Material examined. YEMEN: ABIJAN GOV.: 59 spec., Ba Tays vill. env., wadi Bana, 13 ° 20 ʹ 58 ʺN, 45 ° 17 ʹ 48 ʺE, 168 m, 8. xi. 2007, A. Reiter leg. (NMPC). Habitat. Collected in larger and relatively stable pools in a wadi, with partially running water and dense littoral vegetation (Fig. 2).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FF80FF9FDFC4F390CCE9FEC0.taxon	distribution	Distribution. Widely distributed species in tropical areas of Asia (from Saudi Arabia to Sulawesi in Indonesia) and Australia (BRANCUCCI 1983). First record from Yemen.	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FFBFFF9FDF8BF70BCCE3FDF2.taxon	materials_examined	Material examined. YEMEN: SANA’ A GOV.: 3 spec., Sana’a city, Bait Baws, 15 ° 16.3 ʹN, 44 ° 12.1 ʹE, 2410 m, tank, 6. xi. 2010, J. Hájek leg. (NMPC). Habitat. The species was collected in muddy littoral of a tank (Fig. 5).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FFBFFF9FDF8BF70BCCE3FDF2.taxon	distribution	Distribution. Predominantly Arabian species described from Saudi Arabia and subsequently recorded from Egypt, Sinai Peninsula, Yemen and Iran (cf. SHAVERDO et al. 2013).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
03A8BF79FFBFFF9FDFFBF479CEB1FCC7.taxon	distribution	Distribution. A species described from Saudi Arabia (Hejaz) and ‘ Mesopotamia’. Subsequently its occurrence in Mesopotamia was doubted, and the species was recorded from Yemen (BRANCUCCI 1983).	en	Hájek, Jiří, Reiter, Antonín (2014): Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island. Acta Entomologica Musei Nationalis Pragae 54: 63-99, DOI: 10.5281/zenodo.5314766
