taxonID	type	description	language	source
03AB87B8FF8DFFD9FF66FC7BFA3ADD5A.taxon	description	The main morphological characters of the tribe Piesocorynini are: rostrum wider than long, underside of rostrum without deep transverse sulcus, antennae inserted laterally, antennal scrobes foveiform, not visible from above, antennal club three segmented, antennomere II as long as or longer than antennomere I, eyes entire or only weakly truncate anteriad, eyes coarsely faceted, maximum 26 rows of ommatidia perpendicular to long axis, pronotum with dorsal transverse carina (completely basal, sub-basal or antebasal) or this carina absent (in the case of some undescribed species of Phaenotheriopsis, and both species of Afrophaenotherion) (see also Valentine 1960, 1991, 1999; Legalov 2018; Trýzna et al. 2022; keys to the genera: Jordan 1906; Orellana & Barrios 2021).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF8DFFD9FF66FC7BFA3ADD5A.taxon	discussion	Note on Phaenotheriopsis: in Trýzna et al. 2022: 249, figs 16 – 20, a male of Phaenotheriopsis vauriei Valentine, 1955 (B. D. Valentine det. 2008, MTDC) is depicted as an example of a species with sub-basal dorsal transverse carina of pronotum (sub-basal sensu Holloway 1982: 187, fig. 13; in Valentine 1991: 332 it is mentioned as antebasal). However, as Valentine (1991: 332) mentions, undescribed species still exist in this genus in which the transverse carina of the pronotum is completely absent. In this work, we therefore show an example of such a species with a completely absent carina (Figs 1 – 5). This species comes from “ Insel Guadeloupe ”, and although the specimen is determined as P. bierigi Wolfrum, 1931 (R. Frieser det. 1956, MTDC), it belongs to an undescribed species (moreover, very small eyes are also noticeable in this species). Phaenotheriopsis bierigi is illustrated in Rheinheimer (2004: 226, fig. 156).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF8DFFD9FF66FC7BFA3ADD5A.taxon	discussion	Note on Piesocorynus: in Trýzna et al. 2022: 251, figs 26 – 31, a male of Piesocorynus dispar (Gyllenhal, 1833) is depicted as a representative of the type species of the genus. As stated in the cited work, there are two types of position of the dorsal transverse carina of the pronotum in this genus: 1) either it is completely basal (e. g. P. dispar), or 2) it is distinctly antebasal, but in this case approaching the base of the pronotum at the lateral sides of the pronotum, so it is distinctly curved anteriorly (e. g. P. aspis (Erichson, 1847) (= P. gracilicornis (Jekel, 1855), synonymy according to Queiroz et al. 2017). For completeness, an example of such a species with antebasal dorsal transverse carina is illustrated as P. aspis (female, Figs 12 – 16). Species with a distinct antebasal carina that is parallel to the posterior margin of the pronotum do not occur in this genus.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FEA6FF40DF9F.taxon	description	(Figs 17 – 21)	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FEA6FF40DF9F.taxon	type_taxon	Type species: Afrophaenotherion aereum Frieser, 1984 (by original designation).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FEA6FF40DF9F.taxon	description	Frieser (1984: 176) described the genus Afrophaenotherion based on one species, A. aereum Frieser, 1984 (by monotypy and original designation). Frieser pointed out the similarity of the newly described genus to Phaenotheriolum, from which it mainly differs by: 1) absence of dorsal transverse carina of pronotum; 2) elytra without striae, with punctures irregularly distributed over the entire surface of elytra. Other typical morphological characters are the shiny metallic surface of the body, and pronotum and elytra with setae only very sparse or absent.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FBE3FE1BDB28.taxon	description	(Figs 43 – 86)	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FBE3FE1BDB28.taxon	type_taxon	Type species: Piezonemus durus Jordan, 1904 (by original designation).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF88FFD9FF66FBE3FE1BDB28.taxon	description	Redescription. Rostrum very short, distinctly wider than long (ratio of rostrum length to maximum width 0.3 – 0.4), flat, not carinate, indistinctly emarginate at apex, with only shallow depression before eyes. Apical half of rostrum oblique (converged) laterally. Mandibles more or less bent along the entire length, or along a substantial part, obtuse at apex. Antennal scrobes lateral, located underneath posterior part of eyes (not in front of them), upper edges cariniform. Eyes dorso-lateral, very large, occupying a considerable part of head, elongate, oblique, not emarginate or truncate in anterior part, coarsely granulate, distance between eyes as long as half of length of eye. Frons trapeziform, more or less flattened. Pronotum always wider than long, conical, from dorsal transverse carina narrowed anteriorly, disc of pronotum moderately convex. Dorsal transverse carina sub-basal to basal, always complete. Lateral carinae of pronotum present, extending at most to middle of pronotum, or shorter. Basal longitudinal carinulae (sensu Jordan 1928: 151, Trýzna & Baňař 2021: 558) present. Pygidium of male distinctly longer than wide, convex, rounded at apex. Pygidium of female differently shaped: in P. durus with distinct promontory before apical edge, this projection divided basally; in P. lateralis this promontory is not so high and not so deeply divided as in P. durus; in P. espoch sp. nov. without any promontory, rather flat. Mesotibia of male with small, short spine at apex in all species. Tarsomere I of mesotarsus widely dilated at apex on inner side in P. durus and P. lateralis; tarsomere I of protarsus and mesotarsus with distinct spur at apex in male of P. espoch sp. nov.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF82FFD3FF66FF66FC59DB9D.taxon	description	(Figs 43 – 61, 70 – 72)	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF82FFD3FF66FF66FC59DB9D.taxon	materials_examined	Type locality. Brazil: “ Jatahy, Prov. Goyaz ” Type material. Lectotype, male (designated here, Figs 43 – 48, 102): ‘ Type [p, typical round label with red margin used in BMNH] ’ // ‘ Jatahy. / Prov. Goyas. [p] / Piezonemus / durus. / Type. 1904 / N. Z. p. 276 [= Novitates Zoologicae, page 276] [h] ’ // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ // ‘ LECTOTYPE / Piezonemus durus / Jordan, 1904 / M. Trýzna design., 2023 ’ [red, p] (BMNH). Female: ‘ Jatahy’ / (GOYAZ) [p] // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH); ‘ PARALECTOTYPE / Piezonemus durus / Jordan, 1904 / M. Trýzna design., 2023 ’ [red, p] (BMNH). Other paralectotypes not examined directly by the first author: Unsexed, 1 ex.: ‘ Jatahy’ / (GOYAZ) [p] // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH). Unsexed, 3 exx.: ‘ Jatahy. / Prov. Goyas. [p] // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH). Unsexed, 2 exx.: ‘ Jatahy. / Prov. Goyas. Brésil [p, green paper] // 1 with ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ and the other with ‘ Pres. E. Gowing-Scopes. / Brit. Mus. 1968 - 131 // Piezonemus / durus / 1904 Jord. [h] / Karl Jordan det. [p] ’ (BMNH). Note. Jordan (1904: 276) stated only “ a series ”, without the exact number of specimens of the type series. Lectotype and 7 Paralectotypes are listed above. Other material: 1 male: ‘ Sta. Cruz / Bolivia [p] ’ // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH). 1 male (Figs 49 – 55): Ecuador, Sucumbios prov., Shushufindi, S 0 ° 42´, W 77 ° 19´, 200 – 400 m, 4. – 18. xii. 2004, Zd. Mráček leg. (MTDC). 2 males: Ecuador, Morona-Santiago [province], 30 km NE of Macas, 21. ii. 2022, Cascada de Musap, 1085 m, S 02 ° 07´45 ´´, W 77 ° 55´58 ´´, M. Trýzna & J. C. Carrasco leg. (1 male with dissected terminalia (Figs 56 – 61) in MTDC, 1 male in MERC). Identification. Coloration of the cuticle dark, black to dark brown, basic setae of the upper part of the body dark, with sparse tomentum of whitish grey setae, denser on the head and scutellum. Lower part of body with dense greyish setae. Small irregular clusters of greyish setae on even-numbered elytral intervals, rest of intervals dark, most noticeable on elytral intervals II, IV and VI. Antennomeres rufous brown, only antennomeres IX and X dark brown to black. Legs dark brown, femora, tibiae covered with grey setae. Colour pattern as in Figs 43 – 55, 72. Male tarsomere I of mesotarsus shorter than others together (Figs 49, 53) and dilated at apex on inner side. Pygidium of male elongate, ratio of the maximum length to maximum width 1.24 (Figs 46, 54), pygidium of female with distinct promontory before apical edge, this projection high and deeply divided basally (Fig. 70). Eyes shorter (cf. Figs 44, 50 — P. durus; and 63 — P. lateralis), dorsal ocular index 0.91 in male. Body size ca. 5.3 – 5.5 mm. Lectotype, male (Figs 43 – 48). Measurements (in mm): Total body length — 5.36. Head: total length — 0.86, length of rostrum — 0.19, maximum width of rostrum — 1.07, length of eye — 0.71, maximum width across eyes — 1.31, minimum distance between eyes — 0.41. Antenna: length of segments: II — 0.21, III — 0.34, IV — 0.28, V — 0.26, VI — 0.21, VII — 0.18, VIII — 0.21, IX — 0.21, X — 0.11, XI — 0.20, width of segment IX — 0.14. Pronotum: maximum length — 1.38, maximum width — 1.95, minimum width — 1.25. Elytra: maximum length — 3.20, maximum width — 1.90. Pygidium: maximum length — 0.46, maximum width — 0.37. Male genitalia and associated structures (Figs 57 – 61). Aedeagus (Fig. 59) long, narrow, widest in apical part, this part rectangular. Pedon narrow, its base 2 x wider than the rest of pedon, this apical part almost the same width along its entire length. Tectum broadly triangular, form equilateral triangle. Apodemes of aedeagus long, slightly longer than the part from bridge to apex of aedeagus, almost parallel. Bridge of aedeagus inconspicuously developed. Tegmen (Fig. 61) robust, moderately wide, with strongly sclerotized basal piece, apodeme of tegmen wide, straight, shorter than body of tegmen. Apex of tegmen moderately broad, almost bare. Structure of internal sac as in Fig. 60. Segment VIII (Fig. 57) moderately sclerotized, longer than wide, trapezoidal, anterior part wider than basal part, anterior margin of tergite VIII slightly sinusoidal outside. Sternal lobes of sternite VIII very small, irregularly triangular. Apodeme of sternite VIII present, but indistinct. Sternite IX (= spiculum gastrale) (Fig. 58) with symmetrical divergent arms, apodeme straight. Length of apodeme ca. 2.75 x as long as arms.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF82FFD3FF66FF66FC59DB9D.taxon	distribution	Distribution. Brazil (Jordan 1904, Rheinheimer 2004), Ecuador, Bolivia.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFCFFF66FF66FC0DD8AC.taxon	description	(Figs 62 – 69)	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFCFFF66FF66FC0DD8AC.taxon	materials_examined	Type locality. Brazil: “ Jatahy, Prov. Goyaz ” Type material. Lectotype, male (designated here, Figs 62 – 67, 103): ‘ Type [p, typical round label with red margin used in BMNH] ’ // ‘ Jatahy. / Prov. Goyas. [p] / Piezonemus / lateralis / Type. 1904 / N. Z. p. 276 [= Novitates Zoologicae, page 276] [h] ’ // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ // ‘ LECTOTYPE / Piezonemus lateralis / Jordan, 1904 / M. Trýzna design., 2023 ’ [red, p] (BMNH). Paralectotypes not examined directly by the first author: Unsexed, 5 exx.: ‘ Jatahy. / Prov. Goyas. [p] // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH). Unsexed, 2 exx.: ‘ Jatahy’ / GOYAZ [p] // 1 with ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ and the other with ‘ Pres. E. Gowing-Scopes. / Brit. Mus. 1968 - 131 // Piezonemus / lateralis / 1904 Jord. [h] / Karl Jordan det. [p] ’ (BMNH). Note. Jordan (1904: 276) stated only “ a series ”, without the exact number of specimens of the type series. Lectotype and 7 Paralectotypes are listed above. Other material. Male (Figs 68 – 69): ‘ Hansa Humboldt / Sta. Catherina / Brasilien Reitter [p] ’ // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ // ‘ 3. Piezonemus / lateralis Jord. [h] ’ (BMNH). Male: ‘ Salobro prov de Bahia / Bresil / E. Gounelle, 6.7.1885 [p] ’ // ‘ Karl Jordan Coll. / B. M. 1940 - 109. [p] ’ (BMNH). Identification. The coloration of the body is very similar to almost identical to that in P. durus. Colour pattern as in Figs 62 – 68. Male tarsomere I of mesotarsus distinctly longer than others together (Fig. 69). Pygidium of male elongate, ratio of the maximum length to maximum width 1.36 (Fig. 66), pygidium of female also with distinct promontory before apical edge, but this projection is not so high and not deeply divided basally as in P. durus. Eyes longer (cf. Figs 44, 50 — P. durus; and 63 — P. lateralis), dorsal ocular index 0.66 in male. Body size 4.5 – 4.9 mm. Lectotype, male (Figs 62 – 67). Measurements (in mm): Total body length — 4.86. Head: total length — 0.86, length of rostrum — 0.23, maximum width of rostrum — 0.93, length of eye — 0.66, maximum width across eyes — 1.21, minimum distance between eyes — 0.30. Antenna: length of segments: II — 0.21, III — 0.36, IV — 0.36, V — 0.26, VI — 0.24, VII — 0.20, VIII — 0.21, IX — 0.19, X — 0.13, XI — 0.19, width of segment IX — 0.14. Pronotum: maximum length — 1.13, maximum width — 1.63, minimum width — 1.13. Elytra: maximum length — 2.90, maximum width — 1.60. Pygidium: maximum length — 0.64, maximum width — 0.47.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFCFFF66FF66FC0DD8AC.taxon	distribution	Distribution. Brazil, Paraguay (Jordan 1904, Rheinheimer 2004).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	description	(Figs 73 – 89)	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	materials_examined	Type locality. Ecuador, Orellana province, Loreto env. near El Coca [= Puerto Francisco de Orellana], remnants of natural forest [ca. S 0 ° 42´, W 77 ° 19´] Type material. Holotype, male (Figs 73 – 80): ‘ Ecuador, Napo distr. / Loreto [now in Orellana province] / 17.2. - 14.3.1996 / J. Strnad leg. [p] ’ // ‘ Piezonemus / lateralis / Jord. [h] / det. R. Frieser 199 [p] 6 [h] ’ (NMPC). Allotype, female: (Figs 81 – 86): ‘ Ecuador, 4. - 18. xii. 2004 / prov. Sucumbios, Shushufindi / (S 0 ° 11´, W 76 ° 38´) / 200 - 400 m, Zd. Mráček leg. ’ // Piesocorynus / sp. [h] / det: B. D. Valentine [p]´10 [h] [= 2010] (MTDC). Red label [p] HOLOTYPE (or ALLOTYPE) / Piezonemus espoch sp. nov. / M. Trýzna, V. Caballero-Serrano / & J. C. Carrasco det., 2024. Identification. The coloration of the body is similar to that in both previous species, but in general, coloration of head, pronotum, elytra and legs is more yellowish (rather more greyish in the previous two species). Elytra yellowish in apical and lateral parts, while disc of elytra black. Male tarsomere I of protarsus with spine on lower part (Fig. 78), tarsomere I of mesotibia continuously flowing to spur with a wide base (Figs 73, 79) (dilated at apex on inner side in both previous species). Pygidium of female without any promontory.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	description	Description. Holotype, male (allotype, female) (Figs 73 – 86). Measurements (in mm): Total body length — 4.41 (4.41). Head: total length — 0.86 (0.86), length of rostrum — 0.30 (0.33), maximum width of rostrum — 0.79 (0.77), length of eye — 0.60 (0.57), maximum width across eyes — 1.04 (1.09), minimum distance between eyes — 0.27 (0.27). Antenna: length of segments: II — 0.20 (0.17), III — 0.27 (0.26), IV — 0.21 (0.21), V — 0.21 (0.20), VI — 0.16 (0.14), VII — 0.17 (0.14), VIII — 0.13 (0.10), IX — 0.34 (0.33), X — 0.19 (0.17), XI — 0.21 (0.20), width of segment IX — 0.20 (0.20). Pronotum: maximum length — 0.98 (1.00), maximum width — 1.50 (1.55), minimum width — 0.88 (0.90). Elytra: maximum length — 2.55 (2.63), width in humeral part — 1.50 (1.55). Pygidium: maximum length — 0.50 (0.50), maximum width — 0.44 (0.53). Coloration of the cuticle of entire body generally dark brown to black. Disc of elytra black, margins and apical part brown, abdominal ventrites and pygidium dark brown. Antennae light brown, antennomeres IX – XI black, only apical half of XI dark brown. Labrum distinctly yellow, mandibles light brown with black outer edges. Legs brown, tarsomeres IV – II with lighter basal part than apical, III – V rather yellowish. Vestiture (Figs 73 – 80 male, 81 – 86 female). Head (Figs 74, 82) with sparse appressed yellowish setae, setae denser on upper margin of eyes. Antennomeres I – VIII (Figs 77, 85) almost bare, only with sporadic fine setae located in the apical half of the antennomeres, IX – XI with short appressed black dense setae. Disc of pronotum (Figs 76, 84) with appressed yellowish setae forming irregular and interrupted longitudinal stripe in central part and also slightly denser on lateral sides. Lower side of pronotum with denser yellowish to whitish setae. Elytra (Figs 73, 80 – 81, 86) covered with appressed yellowish setae particularly in apical and lateral parts, disc of elytra with black setae (surface is not abraded here as it might seems). Meso-, metasternum and all abdominal ventrites covered with dense yellowish to whitish setae. All femora with sparse appressed yellowish setae, each tibia with subdecumbent setae forming two yellowish rings in post-basal and apical part, part between rings covered with dark brown setae. Basal part of tarsomeres I – II with light-coloured setae only in basal part, rest of these tarsomeres brown, III – V with light-coloured setae. Pygidium covered with longer sparse yellowish setae. Structure. Head (Figs 74, 82) noticeably short, rostrum flat, without any carinae, only with indistinct depression before eyes. Sides of rostrum short, only imperceptibly widening apically. Ratio of rostrum length to maximum width 0.38 in male, 0.43 in female. Dorsal and lateral part of rostrum only with very fine sculpture. Eyes strikingly large, dorsal ocular index 0.70 in male, 0.66 in female. Ratio of maximum width across eyes to maximum width of rostrum 1.32 in male, 1.42 in female. Antennae (Figs 73, 77, 81, 85) reaching posterior margin of pronotum, however, do not reach half of elytra in both sexes. Scape and pedicel distinct, at least 2 times as wide as antennomeres III on apical parts, VIII shorter and nearly the same width as VII, IX – XI not forming compact club, but with only separated antennomeres, IX 1.7 times longer than wider (1.65 times in female), X as wide as long (wider than longer in female), X ovoid. Pronotum (Figs 76, 84) conical, sides straight, only weakly convex in female, pronotum transverse, ratio of its length to maximum width ca. 0.65 in both sexes, widest at dorsal transverse carina, from here strongly narrowed anteriad. Dorsal transverse carina basal, slightly bisinuate, widely and shallow emarginate in middle posteriad, continuous to sides, here forming weakly protruding sharp lateral angle. Lateral carinae of pronotum distinct, protruding, not reaching to half of length of pronotum (Figs 80, 86). Basal longitudinal carinulae of pronotum present, but due to basal dorsal transverse carina, extremely short. Disc of pronotum moderately convex. Elytra (Figs 73, 81) sub-oval, sides parallel, humeri developed, apical part of elytra broadly rounded. Ratio of the maximum length to maximum width of elytra ca. 1.70 in both sexes. Elytra with pair of indistinct bumps in sub-basal part. Elytral striae with distinct punctures, elytral intervals ca. 3 times as wide as foveae, fovea often weakly longitudinal, spaces between foveae shorter than individual fovea. Legs. Male: protibia and mesotibia with small, short spine at apex (see Fig. 78 — spine on apical part of protibia). Tarsomere I of protarsus with delicate but visible spine on lower part (Fig. 78), tarsomere I of mesotarsus continuous to spur with a wide base (Figs 73, 79). Female: protibia, mesotibia and tarsomeres simple, without any spine or spur. Abdomen shorter than broad. Pygidium slightly convex, widely rounded, longer than wider in male, wider than longer in female. Ratio of the maximum length to maximum width 1.14 in male, 0.94 in female. Pygidium of female without any promontory.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	etymology	Etymology. The species name espoch is dedicated in honor of the Escuela Superior Politécnica de Chimborazo (acronym ESPOCH) in recognition of the support given for the development of the research project Entomofauna IDIPI 281 (see Acknowledgements). Collecting circumstances. Both specimens of the type series were collected by beating smaller twigs of fallen branches of unidentified deciduous trees inside natural forests.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	distribution	Distribution. Ecuador, Orellana (holotype) and Sucumbíos (allotype) provinces (Fig. 87).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF9EFFC6FF66FAB5FBBBD99E.taxon	diagnosis	Differential diagnosis. The new species differs from both other species of Piezonemus mainly by the male protibia and mesotibia with a small, short spine at apex, and tarsomere I of the protarsus with a delicate spine on lower part, and tarsomere I of mesotibia with a distinct spur. Males of two other species lack these spines or spur. The female of the new species has a simple, convex pygidium, without any promontory, while both other species have a bifurcated promontory on the apical part. For more details see key below.	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
03AB87B8FF97FFC5FF66F9E2FEFBDBA9.taxon	discussion	Notes on Piezonemus and confusion with some species of the genus Piesocorynus During the study of Neotropical species of the tribe Piesocorynini in world collections, misidentifications between the similar genera Piezonemus and Piesocorynus were found, as well as confusion caused by abbreviated scientific names on determination labels. For specimens labelled as ‘ P. lateralis Jordan’, it is not clear whether Piezonemus lateralis Jordan, 1904 or Piesocorynus lateralis Jordan, 1906 is intended. Moreover, the two species, and the two genera, are somewhat similar and are confused in collections. Furthermore, both specimens of the new species described here, Piezonemus espoch sp. nov., have been misidentified in the past. The holotype (male) was previously determined as Piezonemus lateralis (R. Frieser det. 1996), and the allotype (female) had been determined as Piesocorynus sp. (B. D. Valentine det. 2010). In addition, one specimen determined by R. Frieser as Piezonemus lateralis (R. Frieser det. 1953) was found in his collection (now in ZSMC), which, however, belongs to the genus Piesocorynus, specifically to the species P. obliquus Frieser, 1978, which Robert Frieser described himself, but he did not include this specimen in the type series. It follows from the above, that distinguishing the genera Piezonemus and Piesocorynus can be difficult. We help to resolve these inaccuracies by illustrating the type material of the relevant species. Therefore, we studied the holotype (male from Panama) and paratype (male from Guatemala) (both in BMNH) of Piesocorynus lateralis, and the holotype (male from Brazil, in ZSMC) and two paratypes (male and female from Brazil, MTDC) of Piesocorynus obliquus. The main distinguishing features of both species are: Piesocorynus lateralis (Figs 90 – 95): disc of pronotum convex, simple, without bumps, uniform coloured (Figs 92, 95). Dorsal transverse carina of pronotum straight (Fig. 92). Elytra almost flat, with only faint pair of bumps in sub-basal part (Fig. 95). Size 5.5 – 5.8 mm. Distribution: Central America (Panama, Guatemala, Mexico). Piesocorynus obliquus (Figs 96 – 100): disc of pronotum with one central and two lateral bumps, with narrow longitudinal contrasting yellow strip in central part (Figs 98, 100). Dorsal transverse carina of pronotum bisinuate (Fig. 98). Elytra with pair of distinct bumps in sub-basal part (Fig. 100). Size 2.5 – 4.0 mm. Distribution: South America (Brazil).	en	Trýzna, Miloš, Caballero-Serrano, Verónica, Carrasco, Juan Carlos (2024): Review of the Neotropical Piezonemus Jordan with type designations, description of Piezonemus espoch sp. nov. from Ecuador, and notes on the tribe Piesocorynini Valentine with keys to world genera (Coleoptera: Anthribidae). Zootaxa 5415 (3): 401-435, DOI: 10.11646/zootaxa.5415.3.3, URL: http://dx.doi.org/10.11646/zootaxa.5415.3.3
