taxonID	type	description	language	source
03AA81215339FFDA51366E8EB99BF8B5.taxon	materials_examined	Material examined. PHILIPPINES: L ඎඓඈඇ: 1 ♀, Banaue, Ifugao, vii. 2018, local coll. (CNCW). M ං ඇൽൺඇൺඈ: 1 J, Dominorog, Bukidnon, i. 2019, local coll. (LMCN); 1 ♀, Davao City, Talomo Mts., Mt. Talomo, 7 ° 04 ′ 36.9 ″ N 125 ° 20 ′ 13.2 ″ E, 1100 m., 25. – 30. iii. 2019, Medina MND. leg. (UMCRC); 1 ♀, Lake Holon, vi. 2019 (DUBC); 1 ♀, South Cotabato, T’Boli, Lake Holon Mts., Lake Holon, 1466 m., 6 ° 05 ′ 48.1 ″ N 124 ° 52 ′ 56.7 ″ E, 12. vi. 2019, Torrejos C. leg. (UMCRC).	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFDA51366E8EB99BF8B5.taxon	diagnosis	Diagnosis. Large species: body length 6.0 – 7.4 mm (Figs 1 a – c). Head depicted in Fig. 7; epipharynx as in Fig. 13. Pronotum with anterior lobes slightly larger than intermediate; median intercosta with dense punctation, second intercosta without punctures, third intercosta with moderately dense punctation; paramedian costae of pronotum can be basally slightly inwardly bent. All intercostae of elytra with two rows of medium-sized punctures. Internal and median protrusion of caudal bulbs connected, with very weak sinuation between them, external protrusion rather distinctly developed (Fig. 19). Surface between external protrusion of caudal bulb of elytra and lateral margin of elytra distinctly sinuate – external protrusion more or less developed; however, median and internal protrusions always with same proportions. Aedeagus as in Figs 25 a – b.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFDA51366E8EB99BF8B5.taxon	distribution	Distribution. Species recorded from the Philippines for the first time. Previously was known from Taiwan (Lanyu Island).	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	description	(Figs 2 a – c, 8, 14, 20, 26 a – b)	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	materials_examined	Type material. H ඈඅඈඍඒඉൾ: J, labelled: ʻMindanao, Lake Holon, vi. 2019 ʼ (DUBC).	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	description	Description. Large species, body length 7.0 mm. Body surface matt, only pygidium with slightly shiny elevated area; black, except for brown or reddish-black preapical glandular area, more densely covered with yellow setae dorsally compared to other species (Figs 2 a – c). Head as observed dorsally and frontally (Figs 2 c and 8, respectively): Clypeus anteriorly not sinuate, with indistinct teeth on sides, laterally distinctly emarginated and next with widely rounded angle, separated by rather small emargination from distinctly protruding, rounded genae; clypeal disc distinctly convex, ringed by deep groove, with pair of distinct, rounded tubercles. Frons with four short, blunt, posteriorly vanishing longitudinal costae, median pair shorter than lateral pair and separated one from another by narrow, deep furrow, distance between neighboring median and lateral costae twice as broad as that between two median costae. Head covered with irregularly spaced fine punctures bearing small setae. Epipharynx as in Fig. 14. Pronotum subquadrate, 1.13 times as wide as long; with six distinctly (but not sharply) convex costae and seven longitudinal intercostae, surface densely covered with yellow setae. Lateral margin with moderately protruding two lobes, anterior lobe triangular, median lobe obtusely angled, slightly wider than anterior lobe. Costae on anterior part present anteriorly as rounded swellings with dense concentration of setae; costae 1 (paramedian pair), 3 and 4 (lateral pairs) – continuous, costae 2 widely interrupted by transverse depression with deep caviform fossae, and another fossa in middle of third furrow at base of costae. First (median) inter-costa covered with distinct medium-sized punctures, very densely concentrated in middle to form elongated depression; lateral intercostae 1 – 3 abundantly punctured only in anterior part, intercostae 2 and 3 smooth in basal two thirds. Elytra (Figs 2 a – c, 20), elongate, 1.73 times as long as wide, each with five costae of similar height and width from elytral base to apex, and one very long, well defined accessory costa present between costae 3 and 4, terminating posteriorly before preapical depressed area; costae slightly shiny on top, covered with yellow setae: costae 1 (sutural) with one row of setae in middle, costae 2 and 3 bordered by row of setae on each side. Intercostae 1 and 2 flat with 3 rows of punctures, with punctures in lateral rows larger than punctures in median row, punctures of intercostae 3 a, 3 b and 4 arranged in two rows, distances between neighboring punctures not uniform, but larger than or approximately equal to puncture diameter. Apical bulbous area of second costa densely covered with yellow macrosetae. Caudal bulbs weakly developed. Internal and median protrusion of caudal bulbs connected; external protrusion moderately developed. Pygidium with elevated area at base, extended as costa to pygidial apex, areas on each side of costa being strongly depressed. All legs sturdy, matt, covered with small punctures and triangular short setae (except for simple and smooth area along outer margin), greatest quantity of setae on mesotibia. Profemora strongly expanded, widest near basal third, with deep groove anteriorly; surface with medium-sized punctures throughout. Protibiae slenderer than mesotibiae, only slightly dilated apically and noticeably incurved apically; apex of protibiae with three small teeth. Mesofemora slightly narrower than profemur, posterior margin bilobed, surface punctate throughout, punctures smaller compared to those on profemora. Mesotibiae narrow at base, gradually expanded from base to about one third of length, and then narrowed to apex; apex of mesotibiae with large projection and tooth (larger than protibiae apical teeth). Metafemora long, slightly narrower than mesofemora; narrow at base, gradually expanded, apical part narrowed; punctures as on mesofemora. Metatibiae larger than mesotibiae, but similar in shape, but not narrowed preapically, only slightly widened to apex. Venter matt, black. Prosternum with long anterior process with rounded apex and hastate posterior process. Mesoventrite with shallow setigerous punctures and small setae. Meso-metaventral plate with distinct, shallow medi- an impression; surface otherwise with irregular medium- sized setigerous punctures. Abdomen with six abdominal ventrites, all separated by impressed sutures; surface punctate-setose. Ventrites 3 – 5 triangularly grooved on each lateral broadened area, groove noticeably large on ventrite 5; ventrites 5 – 6 fused, medially with costae. Aedeagus as in Figs 26 a – b.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	diagnosis	Differential diagnosis. The new species can be distinguished from all Rhyparus species worldwide based on the combination of very long accessory costa, dense setation of dorsal surface, and number of rows of punctures on intercostae.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	etymology	Etymology. The specific epithet is composed of the Latin adjectives flavus, - a, - um and hirtus, - a, - um, and alludes to the dense covering of yellow setae on the dorsum of body.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215339FFD853BE6A15BEB2F8A7.taxon	distribution	Distribution. Species known only from type locality in Mindanao Island, the Philippines.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533BFFDF53B66A76BAF7F7A7.taxon	description	(Figs 3 a – c, 9, 15, 21, 27 a – b)	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533BFFDF53B66A76BAF7F7A7.taxon	materials_examined	Material examined. PHILIPPINES: M ං ඇൽൺඇൺඈ: 1 J 3 ♀♀, Kalatungan, Bukidnon, ix. 2016, local coll. (LMCN); 4 ♀♀, Bukidnon, Kitanglad Mts., Mt. Kitanglad, 8 ° 03 ′ 24.8 ″ N 125 ° 00 ′ 22.9 ″ E, 1200 m, 23. – 24. x. 2018, Medina MND. leg. (UMCRC); 1 J, Davao City, Talomo Mts., Mt. Talomo, 7 ° 04 ′ 36.9 ″ N 125 ° 20 ′ 13.2 ″ E, 1100 m, 25. – 30. iii. 2019, Medina MND. leg. (UMCRC); 1 J, Araibo, Pantukan, Compostela Valley, Candalaga Mts., 7 ° 16 ′ 35.3 ″ N 126 ° 10 ′ 12.8 ″ E, 900 m, 15. – 20. x. 2019, Anichtchenko A. leg. (DUBC); 1 J, New Bataan, Cagan, 7 ° 16 ′ 25 ″ N 126 ° 13 ′ 23.7 ″ E, 18. – 21. x. 2019, Anichtchenko A. leg. (DUBC)	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533BFFDF53B66A76BAF7F7A7.taxon	diagnosis	Diagnosis. Medium-sized to large species: body length 5.2 – 7.0 mm (Figs 3 a – c). Head depicted in Fig. 9; epipharynx as in Fig. 15. Intermediate lobes of pronotum as large as anterior ones; median intercosta with dense punctation; second and third intercostae of pronotum with dense punctation basally. First, second and fourth intercostae of elytra with two rows of medium-sized punctures, third intercosta with three rows of medium-sized punctures basally – medium row presenting to about midlength of elytra. All protrusions of caudal bulb connected, weakly developed (Figs 3 a – c, 21) (see also S ඍൾൻඇ ං ർΚൺ 1998). Aedeagus as in Figs 27 a – b.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533BFFDF53B66A76BAF7F7A7.taxon	distribution	Distribution. Species known from Papua New Guinea: Milne Bay, Central Province, Eastern Highlands, Gulf, Madang, Morobe, Oro, Western Highlands, Western Province, Sepik; and Indonesia: West Papua (M ൾඇർඅ et al. 2013). First record from the Philippines: Mindanao Island.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533CFFD253F36843B826F950.taxon	description	(Figs 4 a – c, 10, 16, 22, 28 a – b)	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533CFFD253F36843B826F950.taxon	materials_examined	Material examined. PHILIPPINES: L ඎඓඈඇ: 1 J, Camirines Sur, Lagonoy, ii. 2019, local coll. (CNCW). M ං ඇൽൺඇൺඈ: 3 JJ 1 ♀, Araibo, Pantukan, Compostela Valley, Candalaga Mts., 7 ° 16 ′ 35.3 ″ N 126 ° 10 ′ 12.8 ″ E, 900 m, 15. – 20. x. 2019, Anichtchenko A. leg. (DUBC); 4 JJ 4 ♀♀, New Bataan, Cagan, 7 ° 16 ′ 25 ″ N 126 ° 13 ′ 23.7 ″ E, 18. – 21. x. 2019, Anichtchenko A. leg. ʼ (DUBC). 1 ♀, Maragusan, Marangig Falls, Candalaga Mts., 7 ° 20 ′ 28.4 ″ N 126 ° 10 ′ 19.8 ″ E, 700 m, 3. – 4. v. 2019, A. Anichtchenko leg. (DUBC). 1 ♀, S Cotobato, Lake Holon, 5. – 10. x. 2019, Anichtchenko A. leg. (DUBC). 1 ♀, Bukidnon, Kitanglad Mt., 8 ° 03 ′ 24.8 ″ N 125 ° 00 ′ 22.9 ″ E, 1200 m, 23. – 24. x. 2018, Medina MND. leg. (UMCRC); 1 ♀, Davao City, Marilog Mts., Marilog, 7 ° 27 ′ 16.2 ″ N 125 ° 14 ′ 52.7 ″ E, 1357 m, 15. – 17. vii. 2018, Medina MND. leg. (UMCRC); 1 J, Dominorog, Bukidnon, iv. 2019, local coll. ” (LMCN).	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533CFFD253F36843B826F950.taxon	diagnosis	Diagnosis. Small species: body length 3.4 – 4.5 mm (Figs 4 a – c). Head as depicted in Fig. 10; epipharynx as in Fig. 16. Intermediate lobes of pronotum slightly larger than anterior one; median intercosta with dense punctation medially; second and third intercostae without or only with simple punctation basally. All intercostae of elytra with two rows of large punctures. Internal and median protrusion of caudal bulbs connected (Fig. 22); external protrusion weakly developed (see also TIJ ඣ * © ං & B ඈ * © ൽൺඍ intervals 1 – 4; anterior part of pronotal costae with minute 2012). Aedeagus as in Figs 28 a – b. appressed microsetae as described below, but otherwise Distribution. Australia (introduced): New South Wales, glabrous; black, except for dark brown antennae and mouth Queensland; Indonesia: Central Java, West Kalimantan, parts (Figs 5 a – c). West Papua; Japan: Honshu, Kyushu, Shikoku, Ryukyus; Head, as observed dorsally and frontally (Figs 5 c and 11, Malaysia: Sabah; New Caledonia; Papua New Guinea: respectively), with anterior clypeal emargination with weak, Central Province, Eastern Highlands, Morobe: the Philippi- very obtuse tooth on each side, distinct lateral emargination nes: Luzon, Negros; Vanuatu; Taiwan; ʻBornéoʼ (M ൾඇർඅ sinuate before widely rounded lateral angle; clypeal margin et al. 2013). First record from Mindanao Island. with upper and lower edges: upper edge concave, sharp and distinct, lower edge obsolete and not observable from above; Rhyparus mindanaoensis sp. nov. clypeal disc very convex, ringed by deep groove, with pair (Figs 5 a – c, 11, 17, 23, 29 a – b) of small rounded tubercles. Frons with four relatively long, Type material. H ඈඅඈඍඒඉൾ: J, labelled: ʻPhilippines, Mindanao, Maragu- sharp, longitudinal ridges of equal length; lateral ridges san, 700 m, Marangig Falls, Candalaga Mts., 7 ° 20 ′ 28.4 ″ N 126 ° 10 ′ 19.8 ″ E, slightly less elevated; distance between median and lateral 3 – 4. v. 2019, A. Anichtchenko leg. ʼ (DUBC). P ൺ * © ൺඍඒඉൾඌ: 1 J, labelled: ridges slightly larger than that between two median ones. ʻPhilippines, Mindanao, Bukidnon, Kalatungan, ix. 2016, local coll. ʼ Head between genae almost as wide as pronotum; genae (LMCN). 1 ♀, labelled: ʻPalawan (Central), Sabang, 50 - 100 m., via very large and strongly protruding. Head irregularly punc- Underground River, I. Lobl, 30. xi. [19] 95 ʼ (ISEA). tate, more sparsely so in frons, punctures large, separated Description. Male holotype: Moderately large species, by about 1.5 – 2.0 times their diameter and bearing minute, body length 5.5 mm; robust, matt, except for shiny tops of pale, relatively thin microsetae. Epipharynx as in Fig. 17. Pronotum transverse, 1.26 times as wide as long; opaque, convex, anterior margin weakly bisinuate; disc with eight costae, six central costae sharp and well delimited. Costae 1 (paramedian pair) continuous, very highly elevated anteriorly and posteriorly, in anterior part forming elevated, elongate tubercles (Fig. 11); costae slightly and evenly convergent in central part. Costae 2 high and well delimited, flattened in transverse depression, anterior part short but very high; fossae of transverse depression small and caviform, with two coracoid processes hanging over fossae. Costae 3 continuous, distinctly elevated, weakly sinuous in anterior third. Costae 4 incomplete, flattened and low but distinct in median part, interrupted in apical third by two deep punctiform fossae, and effaced in basal third; one deep punctiform fossa situated at base of anterolateral pronotal process. Median intercosta densely covered with large punctures, punctures not confluent in median line and sparser in anterior half. Lateral intercostae 1 and 2 each with four large punctures in apical part; intercostae 2 and 3 with only few large punctures. Lateral margin with two lobes, anterior lobe large, relatively narrow and with pointed apex, median lobe subangulate. Elytra (Figs 5 a – c; 17) elongate, 1.65 times as long as wide. Humeral callus with large punctures separated by about one puncture diameter. Each elytron with five strong costae. First costa (sutural) straight, flattened on disc and slightly elevated in apical third, without apical tooth. Second costa sharp, very strongly elevated from base to apical bulbous area, and terminated as strong swelling with dense brown trichome at apex. Third costa slightly incurved, sharp, extending from base to a little before end of costa 2, apex slightly swollen and with brown elongate tubercle. Fourth costa high, of almost same length and width as costa 3, shortly curved at base and slightly sinuous near apex, with relatively long brown tubercle and dense trichome. Fifth costa sinuous, elevated, extending from base to apex, with apical portion strongly swollen and forming bisinuate transverse protrusion with three large tubercles in dorsal view. Elytral intercostae very weakly and transversally corrugated, matt, except for very shiny and impunctate apical parts of intercostae 1 – 4. Intercostae 1 – 4 with two longitudinal rows of large and deep punctures of almost equal size, extending from base to near apex, distances between neighboring punctures relatively uniform, about 1.5 – 2.0 times puncture diameter. Caudal bulbs distinctly developed. Internal and median protrusion of caudal bulbs well developed; external protrusion moderately developed. Pygidium with semicircular elevated area at base connected by narrow ridge with pygidial apex and by deep groove with basal margin, with areas on each side of ridge strongly depressed. Legs. Profemora wide, with large punctures throughout. Protibiae thin at base; with slightly sinuate inner margin up to relatively large and slightly inward bent inner apical tooth, weakly dilated apically, with moderately arcuate and then emarginate inner margin in front of large outer tooth. Mesofemora with 4 – 5 tough, fairly long, sharp, pale microsetae along apical half of anterior edge; posterior margin bilobed, with deep groove, its surface punctate throughout, punctures of half size of those on profemoral surface. Mesotibiae narrow at base, inner margin strongly widened from base to about third of its length and then almost parallel to apex, with short, pale, decumbent setae on inner surface and rows of longer, pale, semierect setae on edges; apex with backward directed inferior spine and inward bent tooth. Metafemora long, narrow, posterior margin slightly bilobed, punctation smaller and sparser than on mesofemora. Metatibiae with outer margin concave, setation less developed than on mesotibiae; inferior apical tooth large and directed inward in male. Venter matt, black. Prosternum with long anterior process with rounded apex and hastate posterior process. Mesoventrite with shallow setigerous punctures. Meso- - metaventral plate with distinct, moderately deep medi- an impression, surface with small setigerous punctures medially and larger punctures laterally. Abdomen with ventrites 1 – 5 with small, shallow and sparse setiferous punctures, punctures along apical and basal margin crenate; ventrites 3 – 5 triangularly grooved on each lateral broadened area, groove distinctly larger on ventrite 5; ventrites 5 and 6 fused, ventrite 6 with large transverse groove with 16 large punctures. Male genitalia. Aedeagus as in Figs 29 a – b. Differential diagnosis. Based on the moderately elongate habitus with elytra widest in basal third, body length 5.5 – 6.0 mm, anterior lateral lobes of pronotum wider than intermediate ones, very high costae of pronotum and elytra with deep intercostae, presence of two rows of punctures in all intercostae of elytra, distinctly divided median and internal protrusion of caudal bulb of elytra with distinct external protrusion, the new species is very similar to Rhyparus denticollis Fairmaire, 1893. However, R. mindanaoensis sp. nov. can be distinguished from the former species by median intercosta of pronotum somewhat narrower and with denser punctation, second and third intercostae of pronotum with sparse / very sparse, irregularly distributed punctation in basal part (versus moderately dense punctation, rather evenly distributed, with similar density to punctation in median furrow), sinuation between internal and median protrusion of caudal bulb of elytra more distinct and tibiae relatively slender. Etymology. The specific epithet is derived from Min- danao Island, where the holotype of the new species was collected; the name is an adjective.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA8121533CFFD253F36843B826F950.taxon	distribution	Distribution. Species known so far only from the Philippines: Mindanao and Palawan islands.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215331FFD1508C6A30B86DF834.taxon	description	(Figs 6 a – c, 12, 18, 24, 30 a-b)	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215331FFD1508C6A30B86DF834.taxon	materials_examined	Type material. S ඒඇඍඒඉൾඌ: 1 J, labelled: ʻPhil. Isl. ʼ [round label] // ʻSyntype ʼ [round label] // ʻNHMUK 014189207 ʼ (BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻ 45 49 ʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189209 ʼ (BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189208 ʼ (BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 013906231 ʼ (BMNH). Additional material examined. PHILIPPINES: L ඎඓඈඇ: 1 ♀, Lagonoy, Camarines Sur, Bicol, vii. 2018, local coll. (CNCW); 1 ♀, Cagayan, Santa Ana, viii. 2018, local coll. (DUBC); 1 J, Barlig Mt. Province, xii. 2018, local coll. (LMCN). M ං ඇൽൺඇൺඈ: 1 ♀, New Albay, Compostela Valley, vii. 2018, local coll. (CNCW); 1 J, Wao, Lanao del Sur, x. 2018, local coll. (CNCW); 1 ♀, Alamada, Lanao del Sur, x. 2018, local coll. (CNCW).	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215331FFD1508C6A30B86DF834.taxon	diagnosis	Diagnosis. Medium-sized species: body length 5.0 – 6.5 mm (Figs 6 a – c). Head as depicted in Fig. 12; epipharynx as in Fig. 18. Intermediate lobes of pronotum as large as anterior one; median intercosta with dense punctation; second and third intercostae with dense punctation basally. All intercostae of elytra with two rows of medium-sized punctures. Internal and median protrusion of caudal bulbs connected (Fig. 24); surface between external protrusion of caudal bulb of elytra and lateral margin of elytra distinctly sinuate – external protrusion more or less developed; however, median and internal ones always with the same proportions. Aedeagus as in Figs 30 a – b. Variability. Lateral lobes of pronotum slightly variable in size and proportions. Punctures of intercostae of pronotum and elytra can be proportionally variable in size between the specimens. Costae of pronotum and elytra are slightly variable in height and width.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
03AA81215331FFD1508C6A30B86DF834.taxon	distribution	Distribution. Philippines: Luzon, Mindanao.	en	Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija, Medina, Milton Norman D. (2021): A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae 61 (1): 99-111, DOI: 10.37520/aemnp.2021.006, URL: http://dx.doi.org/10.37520/aemnp.2021.006
