taxonID	type	description	language	source
03AD87BB1534F70037FEFC68FE3FFE93.taxon	description	Characters. The first antennal flagellomere bears dorsally a few socketed setae, apart from the non-socketed setae that are somewhat shorter and cover the node entirely. The maxillary palpus is five-segmented, with the third segment elongate, a little longer and thicker than the second segment and bearing a very few short hyaline sensilla on its mesial surface. The aedeagus consists of a rod-like sclerotized apodeme that is widened and bearing numerous long trichia apically; this spatulate widening extends dorsally into a voluminous membranous portion that merges almost imperceptibly with the hypoproct (Fig. 1 C). The hypoproct, discernible by the two setae on either side, is apically two-pointed and laterally strong, thereby giving the entire structure the clear contours that otherwise might be mistaken for the lateral margins of the tegmen (Figs 1 A and C). As suggested by both Laštovka and Matile (1972) and Papavero (1977), nigripalpis should be classified with Adidocidia, which is based on the following characters: the anepisternum bears five to six setae on its dorso-anterior margin; the first antennal flagellomere is four or more times longer than wide; the apical margin of ninth tergite does not bear spine-like setae; the gonostylus has a broad and deeply bifurcate apex (Fig. 1 B); the anterior tarsus of female is not enlarged; and the female cercus is somewhat widened subapically (Fig. 1 D). The gonostylus in nigripalpis is diagnostic in bearing subapically two large macrosetae (Fig. 1 B). Compared with the other species classified with Adidocidia, nigripalpis comes closest to the Nearctic D. borealis Coquillett (with one short macroseta subapically on the gonostylus), and D. valida Mik (with one long macroseta in some distance from the apex of gonostylus) and D. trispinosa Polevoi (with three moderately long and thin macrosetae on the mesial surface of gonostylus), both Palearctic. As regards the female terminalia, nigripalpis and borealis are similar in bearing five or so spines dorso-subapically on the cercus, which are absent in valida (the female of trispinosa is not known). These spines are larger in nigripalpis than in borealis (see Laštovka & Matile 1972: fig. 23). Altogether, nigripalpis and borealis appear to be sister species.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1534F70037FEFC68FE3FFE93.taxon	discussion	Discussion. Specimens from Costa Rica are unquestionably conspecific with the type specimens of nigripalpis from Brazil. In two male specimens from Costa Rica with complete antennae, the first flagellomeres are four times as long as wide and thus slightly shorter than in the male syntype specimen. The balsammounted terminalia of the latter are very pale, which makes perception of structural details of the aedeagustegmen complex difficult.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1534F70037FEFC68FE3FFE93.taxon	materials_examined	Material studied. 1 male (on slide), Costa Rica, Cartago province, Finca los Lagos, Madreselva, alt. 2600 m, December 1993, by Malaise trap, M. M. Chavarria (INBio sample # 2602); 1 male, 1 female (both on slides), Cartago province, Parque Nacional Tapantí-Macizo de la Muerte, Estación La Esperanza, alt. 2200 - 2600 m, 3 – 22 November 2003, by Malaise trap, M. & C. Jaschhof; 1 male (pinned, with terminalia mounted on a plastic sheet in Canada balsam, labelled “ Type ”), “ Brasilien, Nova Teutonia, 27 ° 11 ’ B / 52 ° 23 ’ L, 10.7.1938, Fritz Plaumann ” (Brit. Mus. 1938 - 682, BMNH (E) # 253409); 1 male (pinned, with terminalia in glycerol, labelled “ Syntype ”), same data as previous specimen but “ 9.7.1939 ” (Brit. Mus. 1939 - 628, BMNH (E) # 253410); 1 female, abdomen missing (pinned, labelled “ Syntype ”), same data but “ 9.6.1939 ” (BMNH (E) # 253411).	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1536F70037FEFE3AFBD2F81D.taxon	description	Description. Male. Body length 2.8 – 3.1 mm. Head. Antennal flagellum bearing short non-socketed setae intermingled with a few sensory spines; longest setae as long as half flagellomere width. First flagellomere 2.3 to 2.7 times as long as wide; bearing dorsally 1 – 3 short socketed setae (Fig. 2 A). Fourth flagellomere 1.5 times as long as wide (Fig. 2 A). Maxillary palpus 5 - segmented; third segment 1.8 times as long as second segment, bearing mesially a few short hyaline sensilla apart from ordinary setae elsewhere; fourth and fifth segments elongate cylindrical, fifth segment 1.6 times as long as fourth segment. Thorax. Anepisternum with 2 – 3 setae in upper portion, other pleura non-setose. Legs. Antero-apical depression of fore tibia with distinct semicircular rim, bearing 20 – 25 non-socketed, stiff setae; apical setae arranged in line. Wing. Sc entering C well before level of ta. Sc-r very weak. R 1 entering C well beyond level of M-furcation. A 1 reaching wing margin. A 2 absent. Terminalia. Ninth tergite with broadly rounded apical margin, bearing setae of various sizes. Gonocoxites ventrobasally interlinked through membranous, non-setose portion, otherwise bearing setae of various sizes. Gonostylus rather stout, bearing apically strong double-tooth; tooth half as long as gonostylus body, tooth width at base 0.35 times its length (Fig. 2 C). Aedeagus with rod-like apodeme, the latter apically funnellike and bearing numerous long trichia. Parameres fused to form tegmen; tegmen apically rounded, with weak contours; parameral apodemes strong. Hypoproct large, apically incised, bearing 1 large seta on either side. Cerci as large as hypoproct, bearing apically and dorsally setae of various sizes. Female. Body length 3.0 – 3.1 mm. Head. First flagellomere 2.6 times as long as wide. Fourth flagellomere 1.7 times as long as wide. Thorax. Anepisternum bearing 0 – 3 setae. Legs. Tarsus of fore leg not enlarged; three distal tarsomeres bearing sole of dense, stiff setae. Terminalia. Cercus somewhat widened subapically (Fig. 2 E).	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1536F70037FEFE3AFBD2F81D.taxon	discussion	Discussion. The first antennal flagellomere in hybrida is short, as is typical of the subgenus Diadocidia s. str., while the anepisternum is setose, as is typical of Adidocidia. As regards male terminalia in hybrida, the gonostylus is Diadocidia s. str. - like in having its apex slender and bifurcate for just a short distance, while the ninth tergite is Adidocidia - like in lacking spine-like setae at its apex. Female terminalia are Adidocidia - like in that the cercus is somewhat widened subapically. This adds up to a mixed constellation of features, so that hybrida cannot be classified to either Diadocidia s. str. or Adidocidia. It cannot be assigned to the subgenus Taidocidia either, because in globosa, its only species, the subcosta enters the radius, not the costa, and the gonostylus is peculiar in bearing apically a comb of thick, short macrosetae (Papp & Ševčik 2005). Apparently, the subgeneric classification of Diadocidia needs reassessment, which should be addressed further once the extra-Holarctic fauna of diadocidiids becomes better known.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1536F70037FEFE3AFBD2F81D.taxon	etymology	Etymology. From Latin, hybridus, hybrid, referring to the peculiar mix of characters. Types. Holotype. Male (on slide). Costa Rica, Guanacaste province, Nandayure, Bellavista, Cerro Azul, alt. 1050 m, 1 May – 19 July 2003, by Malaise trap, W. Porras & D. Briceño (INBio sample # 74645). Paratypes. 4 males, 2 females, same data as the holotype.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1536F70037FEFE3AFBD2F81D.taxon	materials_examined	Other material. 51 males, 4 females (all in ethanol), same data as the holotype.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1531F70637FEF965FDCAFC6B.taxon	description	Description. Male. Body length 2.8 – 3.1 mm. Head. Antennal flagellum with short non-socketed setae intermingled with a few sensory spines; longest setae as long as half flagellomere width. First flagellomere twice as long as wide (Fig. 2 B). Fourth flagellomere 1.3 times as long as wide (Fig. 2 B). Maxillary palpus 5 - segmented; third segment thickest, stout, 1.6 times as long as second segment, bearing a very few short hyaline sensilla apart from ordinary setae elsewhere; fourth and fifth segments elongate cylindrical, fifth segment 1.6 times as long as fourth segment. Thorax. Anepisternum with 4 (holotype), 8 or 12 (paratypes) setae in upper portion, other pleura non-setose. Legs. Antero-apical depression of fore tibia with distinct semicircular rim, bearing some 15 non-socketed, stiff setae scattered rather than ordered in lines. Wing. Length 2.7 mm. Venation similar to hybrida, but Sc entering C shortly before level of ta. Terminalia. Fig. 2 D. Width at base of gonostylus tooth 0.4 times the length. Otherwise similar to hybrida. Female. Unknown.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1531F70637FEF965FDCAFC6B.taxon	discussion	Discussion. We do not assign similis to any subgenus for the same reasons as explained under hybrida. These two species are very similar and apparently closely related. Compared with hybrida, the gonostylus body in similis is somewhat broader basally and the apical tooth is thicker. The first antennal flagellomere is shorter in similis than in hybrida. The third palpus segment is thickened in similis, while it is not so in hybrida. Setae of the fore tibial anteroapical depression are scattered in similis, while they are partly arranged in a line in hybrida.	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
03AD87BB1531F70637FEF965FDCAFC6B.taxon	etymology	Etymology. From Latin, similis, similar, referring to the similarity to hybrida. Types. Holotype. Male (on slide). Costa Rica, Cartago province, Parque Nacional Tapantí-Macizo de la Muerte, Estación La Esperanza, alt. 2200 - 2600 m, 3 – 22 Nov. 2003, by Malaise trap, M. & C. Jaschhof. Paratypes. 1 male (in ethanol), same data as the holotype; 1 male (in ethanol), Puntarenas province, Parque Internacional La Amistad, Estación Altamira, Cerro Biolley, alt. 1700 m, 11 June – 11 July 2001, by Malaise trap, D. Rubí (INBio sample # 64127 B).	en	Jaschhof, Mathias, Jaschhof, Catrin (2007): On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa 1586: 33-38, DOI: 10.5281/zenodo.178463
