taxonID	type	description	language	source
03AC87B2FF86FFADE12FFC6BFC6080C1.taxon	description	Typespecies: Systenostomatamlod PanhaetBurch 1999, byoriginaldes- ignation. Content: concava (ThompsonetUpatham, 1997), dominikae Páll- Gergely etHunyadi, 2015, elevata (ThompsonetUpatham, 1997), huoyani Jochum, SlapniketPáll- Gergely, 2014, fabella Páll- GergelyetHunyadi, 2015, singuladentis InkhavilayetPanha, 2016, subelevata Páll- GergelyetHunyadi, 2015, stochi Páll- GergelyetJochum, sp. n., szekeresi Páll- GergelyetHunyadi, 2015, tamlod (PanhaetBurch, 1999). Distribution: ThisgenusisdistributedinnorthernThailand, southern ChinaandnorthernVietnam (Fig. 7). Angustopilastochi Páll- GergelyetJochumsp. n. (Figs 1 C, 3 A – H) Typematerial: C 03 - N 39 (localitycode), NorthernVietnam, HaGiangProvince, Quan BaDistrict, TamSon, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, atthe entrance, goursofdrippingwater, leg. G. TOMASIN, 28.01.2010., HolotypeMZUFGC / 49405; C 04 - N 74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, internalcanyon, goursof drippingwater, leg. G. TOMASIN, 29.01.2010., MZUFGC / 49412 (figuredparatype, juvenile). (umbilicalside), G = sculptureofthebodywhorl. Diagnosis: atiny, convexlyconicspecieswithanovate- subquadrateap- erture, atuberculatedparietalandalowpalataldenticle. (umbilicalside), H = sculptureofthebodywhorl Description: Shellminute, greyishwhite, veryfragile, spireconvexlyconicwithob- tuseapex; whorlsshouldered; spiretiltedslightlyleft; fineequidistantrowsofstriaeradiate fromthenuclearwhorl, startingatazoneofgranulosewrinklesandcontinuingontothe teleoconch; teleoconchsculpturefinelyretiformwithirregularly- spacedradialgrowthlines crossedbyfinerowsofspiralthreads; the 4.75 rounded, evenlyincreasingwhorlsaresepa- ratedbyadeepsuture; apertureovate- subquadrate; parietaldenticledeeply- set, consisting ofthreetuberclesrunninginside, thefirstconspicuouslylowerthanthesubsequenttwo; oppositeoftheparietaldenticleisasmall, deeply- set, lowpalatalplica; distalportionofbody whorlformsashortdetachedtuba; peristomethinatuppermostrightregion, basallyslightlyreflected; umbilicalzonehighlyreticulate; umbilicusnarrow; penultimatewhorlbulges decidedlybeyondtheplaneoftheaperturebyca, 1 / 6 themax. breadthofshell (lateralview). Measurements (in mm): SH = 1.26; SW = 1.06; AH = 0.43; AW = 0.42 (holotype). Differentialdiagnosis: Angustopilastochi sp. n. differsfromthesympa- tric A. subelevata by the larger shell and the presence of two denticles in the aperture. Three Angustopila speciespossesstwoaperturalteethsimilarto A. stochi sp. n., namely, A. huoyani, A. tamlod and A. dominikae. These species are considerably smaller than A. stochi. Moreover, A. huoyani, A. tamlod and A. dominikae bearamorepronounced, tuberculatedparietaldenticlecausingthe aperturetoappearheart-shaped. A. dominikae hasamoreaccentuatedpalatal denticleandasubglobose- conicalshell. Etymology: ThisnewspeciesisdedicatedtoDr. FabioStoch, whosentthesamplesto Dr. MarcoBodonwhokindlyprovidedusthematerial. Distribution: Angustopilastochi sp. n. isknownfromsedimentsamples fromtheCongTroiCaveinNorthernVietnam. Remarks: Thelastquarterwhorloftheholotype (theonlyknownadult shell) got damaged during the SEM imaging. Hence, these two shell frag- mentsaredepositedintheMZUF. Thisbreakageisprobablyduetoshellde- cayonaccountofitspreviouspreservationinformaldehyde (MarcoBodon, pers. comm.). Angustopilasubelevata Páll- GergelyetHunyadi, 2015 (Figs 1 B, 2 A – G, 7 E – F.) Angustopila subelevata Páll- Gergely et Hunyadi, In: PÁLL-GERGELY et al. 2015, ZooKeys 523: 39 – 42, Fig. 4. Materialexamined: C 05 - N 5 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, attheentrance, lakewithdrippingwater, leg. G. TOMASIN, 29.01.2010., 2 stronglycorroded shells, MZUFGC / 49398; C 03 - N 39 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, tureheightandmultipliedby 100. at the entrance, gours of dripping water, leg. G. TOMASIN, 28.01.2010., MZUF GC / 49404 (1 shell); C 04 - N 74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, internalcanyon, goursof dripping water, leg. G. TOMASIN, 29.01.2010., Specimen 1 (figured shell), Specimen 2 (figured shell), additional 26 shells, MZUF GC / 49411; Same data, HNHM 99733 / 1; Same data, PGB / 2. Measurements (in mm): SH = 0.82 – 0.94; SW = 0.78 – 0.86; AH = 0.32 – 0.36; AW = 0.33 – 0.35 (n = 12) (Table 1). Distribution: Angustopilasubelevata wasdescribedfromGuangxi, south- ernChina. Thelocalityreportedinthispaperrepresentsthefirstlocalityfrom Vietnam. Remarks: TheVietnamesespecimensareslightlylargerthanthetype materialfromGuangxi, China (Min. SH: 0.83, maxSH: 0.91, mean: 0.87: Páll- Gergely etal. (2015), butnootherconchologicaldifferenceswerefound. The parietalcallusofoneoftheshellsformsatooth- likeprojection (Fig. 7 F). Thematerialexaminedherehasspiralthreadsontheumbilicusandat the base of the shell in sync with PÁLL-GERGELY et al. (2015). PANHA and BURCH (2005) didnotobservethischaracteristicintheirinvestigationsofthemost similarThaispecies Angustopilaelevata (ThompsonetUpatham, 1997).	en	Páll-Gergely, Barna, Jochum, Adrienne, Asami, Takahiro (2017): Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 63 (3): 327-341, DOI: 10.17109/AZH.63.3.327.2017, URL: http://dx.doi.org/10.17109/azh.63.3.327.2017
03AC87B2FF82FFA2E169FAF9FBE281DB.taxon	description	Differentialdiagnosis: Dentisphaera differsfrom Angustopila bythepres- enceofsixteethintheaperture, andtheelongationofthetuba (mostcon- spicuouslyvisiblefromdorsalview). Etymology: Thename Dentisphaera referstothepresenceofmultipleteethinthe peristomeandtheovoid, almostsphericalshapeoftheshell. Dentisphaeramaxema Páll- GergelyetJochumsp. n. (Figs 1 D, 4, 7 C – D) Typematerial: C 04 - N 74 (localitycode), NorthernVietnam, HaGiangProvince, Quan BaDistrict, TamSon, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, internal canyon, goursofdrippingwater, leg. G. TOMASIN, 29.01.2010., HolotypeMZUFGC / 49408; Samedata, MZUFGC / 49409 / 12 paratypes; Samedata, HNHM 99732 / 1 paratype; Same data, PGB / 2 paratypes; Samedata, MZUFGC / 49410 / 1 juvenileshell (notparatype); C 05 - N 5 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, Cong TroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, attheentrance, lakewithdrippingwater, leg. G. TOMASIN, 29.01.2010., 2 corrodedparatypes, MZUFGC / 49397; C 03 - N 39 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, TamSon, Cong TroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, attheentrance, goursofdripping water, leg. G. TOMASIN, 28.01.2010., Specimen 1 (figuredparatype), additional 8 paratypes, MZUFGC / 49402; Samedata, 4 shellfragments (2 apical, 2 lowerparts; notparatypes), MZUF GC / 49403. Diagnosis: Shelltiny, ovate- conoidwithshoulderedwhorls, umbilicus narrow, tubaprotruding, sixrobustteeth (1 parietal, 1 angular, 2 palatal, 2 columellar). Description: Shellminute, ovate, veryfragile, lightgreywithspireconvexlyconicwith obtuseapex; protoconchrecessedtotherightatantepenultimatewhorl; nuclearwhorlbe- ginswithapatternofgranulosewrinkles, mergingintoapatternofdense, shallowpits; the 4.5 convex, unevenlyincreasingwhorlsareseparatedbyadeepsuture; teleoconchsculpture finelyretiformwithraisedequidistantly- spacedspiralthreads; smallaperturepiriform; six prominent, deeplysetdenticlesformrobustbarriers; parietaldenticleslightlycontortedand aligneddirectlyoppositethedeeper- set, palataldenticle; angulardenticleslantstotheright; bodywhorlthick; apertureobliquetoshellaxis; tubaendofbodywhorldetachedandpro- jectingslightlydownwardpastpenultimatewhorl (lateralview); tubadeflectstotherightof theumbilicus; umbilicalzonefaintlyreticulate; umbilicusnarrowanddeep. Measurements (in mm): SH = 0.87 – 1.06; SW = 0.85 – 0.96; AH = 0.38 – 0.45; AW = 0.38 – 0.44 (n = 18) (Table 2). Etymology: ThespecificepithetderivesfromtheHungarianword “ mákszem ” (poppy seed), referring to the tiny shell. This name is to be used as a noun in apposition. tionsseeTable 1. Distribution: Dentisphaeramaxema sp. n. isknownfromsedimentsam- plesfromtheCongTroiCaveinNorthernVietnam. Fig. 5. Holotype of Tonkinospira tomasini sp. n. A – D = different views of the shell, E = proto- conch, F = sculpture of the base of the shell (umbilical side), G = sculpture of the body whorl	en	Páll-Gergely, Barna, Jochum, Adrienne, Asami, Takahiro (2017): Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 63 (3): 327-341, DOI: 10.17109/AZH.63.3.327.2017, URL: http://dx.doi.org/10.17109/azh.63.3.327.2017
03AC87B2FF8EFFA6E137FF3AFCAB8559.taxon	description	Systenostoma — Bavay & Dautzenberg, 1908, JournaldeConchyliologie, 57: 196. (diagnosis). Tonkinospira Jochum, SlapniketPáll- Gergely, 2014: In: JOCHUMetal., ZooKeys 410: 31. Typespecies: Helix (Systenostoma) pauperrima BavayetDautzenberg, 1908 bysubsequentdesignation (PILSbRY 1917). Content: defixa (BavayetDautzenberg, 1912), depressa (Jaeckel, 1950), pauperrima (BavayetDautzenberg, 1908), pulverea (BavayetDautzenberg, 1908), tomasini Páll- GergelyetJochum, sp. n. Distribution: Tonkinospira speciesareknownfromNorthernVietnamso far (PÁLL-GERGELY et al. 2015 and this study). Tonkinospiratomasini Páll- GergelyetJochumsp. n. (Figs 1 A, 5, 6, 7 A – B) Typematerial: C 03 - N 39 (localitycode), NorthernVietnam, HaGiangProvince, Quan BaDistrict, TamSon, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, atthe entrance, goursofdrippingwater, leg. G. TOMASIN, 28.01.2010., Specimen 1 (= holotype, MZUFGC / 49399); Samedata, Specimen 2 (figuredparatype), Specimen 3 (figuredpara- type), additional 4 paratypes, MZUFGC / 49400; Samedata, HNHM 99731 / 1 paratype; Samedata, PGB / 1 paratype; Samedata, 2 juvenileshells (notparatypes), MZUFGC / 49401; C 04 - N 74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23 ° 02 ’ 44.7 ” N, 104 ° 59 ’ 23.9 ” E, WGS 84, internalcanyon, gours ofdrippingwater, leg. G. TOMASIN, 29.01.2010., MZUFGC / 49407 / 1 paratype. Diagnosis: Smallconicalshellwithconvexwhorls, lessthan 1.95 mm broad; apertureedentate. Description: Shellconvexlyconicwithobtuseapex, greyishwhite; nuclearwhorlbe- ginswithapatternofgranulosewrinkles, mergingintoapatternofdense, shallowpits seeTable 1. Fig. 6. Variation of shell shape in Tonkinospira tomasini sp. n. A = n 39, Specimen 2, B = n 39, Specimen 1 (holotype); C = n 39, Specimen 3 mellar, 5 – 6 = palatal withoutspiralthreads, granulartexture; suturedeep; protoconchconsistsofapproximate- ly 1.5 whorls; teleoconchretiformwithfaintspiralthreads; protoconchandteleoconchcon- sistsof 5 evenlyincreasingconvexwhorls; aperturequadrate- reniform, adnate; leftouter margin thickly angular with decreasing thickness descending towards the basal margin andascendingtowardstheupperrightportionoftheperistome; basal- mostsectionof peristomefoldsbackslightly; distalendofthebodywhorldoesnotproceedpastpenul- timatewhorlbutratheralignsevenwiththeventralconvexityofthepenultimatewhorl inprofileview; distalendofbodywhorldeflectstotherightoftheumbilicus; umbilicus narrow and deep. Measurements (in mm): SH = 2.23 – 2.98; SW = 1.7 – 1.93; AH = 0.79 – 0.91; AW = 0.82 – 0.94 (n = 8) (Table 3). Differentialdiagnosis: Themostsimilar Tonkinospira speciesis T. pauperrima, which has a wider shell, a lower spire and shows a more pronounced spiralstriationthan T. tomasini sp. n. Fig. 8. Distributionof Angustopila species. 1 = typelocalityof A. huoyani, 2 = otherlocality of A. huoyani (see PÁLL-GERGELY et al. 2015), 3 = type locality of A. dominikae, A. subelevata and A. szekeresi, 4 = type locality of A. stochi sp. n. (Cong Troi Cave) and other locality of A. subelevata, 5 = typelocalityof A. fabella, 6 = typelocalityof A. tamlod, 7 = typelocalityof A. elevata, 8 = type locality of A. concava Etymology: ThenewspeciesisnamedafterG. Tomasin, whocollectedthematerial fromCongTroiCave. Distribution: Tonkinospiratomasini sp. n. isknownfromsedimentsam- plesfromtheCongTroiCaveinNorthernVietnam.	en	Páll-Gergely, Barna, Jochum, Adrienne, Asami, Takahiro (2017): Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 63 (3): 327-341, DOI: 10.17109/AZH.63.3.327.2017, URL: http://dx.doi.org/10.17109/azh.63.3.327.2017
