identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AF182F1B21051323A9FD00FF2A29C0.text	03AF182F1B21051323A9FD00FF2A29C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Charletonia cameroonensis Haitlinger & Kekeunou	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Charletonia cameroonensis Haitlinger &amp; Kekeunou sp. nov.</p>
            <p>(Figs. 1–11)</p>
            <p>Diagnosis. Four setae between coxae II and III, fD 90–92, fV 30–34, NDV 120–126, Ta I 226–237, Ti I 335–363, Ti III 412–444, fn Bf 4-4-3.</p>
            <p>Description. Larva (Holotype). Dorsal idiosoma with 92 (90–92) weakly barbed setae. One pair of eyes, cornea 23 x 21 (Fig. 1). Dorsal scutum punctate, a little wider than long (in two paratypes distinctly wider than long, in one paratype longer than wide), anterior border concave, posterior border slightly concave. Three pairs of scutalae (AL&gt;ML&gt;PL), all slightly barbed. Two pairs of sensillae with fine setules in distal half only (Fig 3).</p>
            <p>Venter of idiosoma bears a pair of setae between coxae I, a pair of setae between coxae II, four setae between coxae II and III and 34 (30–32) setae behind coxae III; all setae setulose. Coxalae setulose, coxalae 1b longer than other coxalae (Fig. 2).</p>
            <p>Gnathosoma. Galeala (cs) spiniform, nude, anterior hypostomala (as) nude, over twice shorter than posterior hypostomala (bs), bs with relatively long setules. Palpfemur and palpgenu each with one barbed seta, palptibia with three barbed setae. Odontus divergent (Fig. 4). Palptarsus bears five normal setae, two covered with setules, three smooth (two are long 55 and 58), and one proximal solenidion and distal eupathidium (Fig. 5). Supracoxal peg-like seta present (eP).</p>
            <p>Leg setal formula: Leg I: Ta 1ω, 1ε, 2ζ, 29 (25–29); Ti 2φ, 1κ, 18, Ge 1σ, 1κ, 12, Tf 5, Bf 4, Tr 1, Cx 1 (Figs. 6, 7). Leg II: Ta 1ω, 2ζ, 23 (26–27); Ti 2φ, 18; Ge 1κ, 12; Tf 5; Bf 4; Tr 1; Cx 2 (Figs. 8, 9). Leg III: Ta 1ζ, 26 (24– 26); Ti 1φ, 18 (18–19); Ge 12; Tf 5; Bf 3; Tr 1; Cx 2 (Figs. 10, 11).</p>
            <p>Leg lengths. Leg I 1339 holotype, 1278, 1236, 1333 paratypes, leg II 1206, 1129, 1137, 1184, leg III 1507, 1413, 1436, 1506. IP = 4062 holotype, 3820, 3809, 4023 paratypes.</p>
            <p>Measurements are given in Table 1.</p>
            <p>Etymology. Named after its occurrence in Cameroon.</p>
            <p> Type material. All larvae were collected by S. Kekeunou in 7 October 2012 from  Z. variegatus . Holotype and 3 paratypes were collected in Zamakoe (3º34’44” N, 11º39’33” E). The holotype is deposited in the Museum of Natural History, Wrocław University and three paratypes are deposited in the Museum of Natural History, Wrocław University of Environmental and Life Sciences, Poland. </p>
            <p> Remarks.  Charletonia cameroonensis Haitlinger &amp; Kekeunou sp. nov. belongs to the species group with four setae between coxae II and III, Ti III&gt; 320 Μm and Ta I&gt; 190 Μm. This group includes  C. dorotae Haitlinger, 1987 ,  C. taiwanensis Tsai &amp; Chow, 1988 , C.  ishii Southcott, 1991 ,  C. domawiti Haitlinger, 2004 ,  C. ramoni Haitlinger, 2007 ,  C. rajmundi Haitlinger, 2007 and  C. mehranensis Haitlinger &amp; Saboori, 2007 (Haitlinger 1987, 2004, 2007a, c; Tsai &amp; Chow 1988; Southcott 1991; Haitlinger &amp; Saboori 2007). Moreover, to this group  C. brunni (Oudemans, 1910) and  C. volzi (Oudemans, 1910) can be included—length of tarsus I and tibia III partly lies in the settled limit for this group. It differs from  C. dorotae in fD (90–92 vs. 104), fV (30–34 vs. 46–54), the longer GL (180–185 vs. </p>
            <p> 167), 1a (63–67 vs. 48–60), Ta I (226–237 vs. 194–200), Ti I (335–363 vs.270–294), Ta II (212–226 vs. 182–184), Ti II (288–308 vs. 228–254), Ta III (229–245 vs. 192–202), Ti III (412–444 vs. 330–360), IP (3809–4052 vs. 3058–3256) and fnBf (4-4-3 vs. 4-4-2); from  C. taiwanensis in fD (90–92 vs. 70), the longer L (122–135 vs. 94), W (125–135 vs. 100–118), AP (61–67 vs. 44–47), AL (86–98 vs. 63–75), ML (71–83 vs. 44–47), Ta II (226–237 vs. 196–197), Ti III (412–444 vs. 331–338), IP (3809–4052 vs. 3163–3197) and fnBf (4-4-3 vs. 4-4-2); from  C. ishii in fD (90–92 vs. 71), fV (30–34 vs. 24), the longer L (122–135 vs. 109–118), W (125–135 vs. 118–121), AP (61–67 vs. 51), cs (51–56 vs. 36), bs (52–69 vs. 36), Ta I (226–237 vs. 198–218), Ti I (335–363 vs. 294–322), Ti III (412– 444 vs. 355–387) and fnBf 4-4-3 vs. 4-4-2; from  C. domawiti [based on specimens from Brazil and Costa Rica (Mayoral &amp; Barranco, 2011)] in fD (90–92 vs. 44), fV (30–34 vs. 18), the longer Ta I (226–237 vs. 202–207), Ti I (335–363 vs. 288–315), Ta III (229–245 vs. 203–213), Ti III (412–444 vs. 365–405), IP (3809–4062 vs. 3349– 3604), 1b (127–130 vs. 100–117), 2b’ (97–102 vs. 84–94) and 2b’’ (69–74 vs. 54–63); from  C. ramoni in fD (90– 92 vs. 68), fV (30–34 vs. 24), the longer L (122–135 vs. 104–110), W (125–135 vs. 112–120), AP (51–67 vs. 40– 46), ML (71–83 vs. 42–50), 1a (63–67 vs. 44–58), 1b (127–130 vs. 96–114), Ta I (226–237 vs. 196–206) and Ti III (412–444 vs. 336–366); from  C. rajmundi in fD (90–92 vs. 148–152), fV (30–34 vs. 53), the shorter AW (82–89 vs. 98–104), longer AP (61–67 vs. 52–54), 1b (127–130 vs. 98–120), Ti I (335–363 vs. 284–304), Ti III (412–444 vs. 346–364) and IP (3809–4052 vs. 3158–3444); from  C. mehranensis in fD (90–92 vs. 102), the longer AP (61– 67 vs. 50–56), AL (86–98 vs. 72–84), 1a (63–67 vs. 52–60), 2a (82–93 vs. 72–80), 1b (127–130 vs. 108–114), 2b’ (97–102 vs. 84–94), Ti I (335–363 vs. 296–310), Ti III (412–444 vs. 374–386), IP (3809–4062 vs. 3510 –3616) and fnBf (4-4-3 vs. 4-4-2); from  C. volzi in the longer AP (61–67 vs. 35–48), L (122–135 vs. 100–116), W (125–134 vs. 108–124), GL (180–185 vs. 164–172), 1a (63–67 vs. 40–56), 1b (127–130 vs. 92–108), 2b’ (97–102 vs. 80– 92), PsFd (101–112 vs. 78–92), Ta I (226–237 vs. 178–196), Ti I (335–363 vs. 248–294) and Ti III (412–444 vs. 302–360) and from  C. brunni in the longer 1a (63–67 vs. 42–50), 2a (82–93 vs. 52–64), 1b (127–130 vs. 92–110), Ta I (226–237 vs.184–208), Ti I (335–363 vs. 244–268), Ti III (412–444 vs. 310–332) and shorter AW (82–89 vs. 92–102). </p>
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	https://treatment.plazi.org/id/03AF182F1B21051323A9FD00FF2A29C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Haitlinger, Ryszard;Kekeunou, Sévilor;Łupicki, Dariusz	Haitlinger, Ryszard, Kekeunou, Sévilor, Łupicki, Dariusz (2014): Charletonia cameroonensis Haitlinger & Kekeunou sp. nov. and the first record of C. justynae Haitlinger, 1987 (Acari: Erythraeidae) from Cameroon with redescription of the species. Zootaxa 3760 (1): 39-53, DOI: 10.11646/zootaxa.3760.1.2
03AF182F1B24051723A9FA23FAF72F5B.text	03AF182F1B24051723A9FA23FAF72F5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Charletonia justynae Haitlinger 1987	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Charletonia justynae Haitlinger, 1987</p>
            <p>(Figs. 12–20)</p>
            <p>Diagnosis. Larva. Four setae between coxae II and III, fnBf 4-4-2, fD 65–94, Ta I 169–194, Ti III 245–296, σI placed at κ, both distally located on genu I.</p>
            <p>Description. Larva. Dorsal side of idiosoma with 94 weakly barbed setae in holotype (65–83 in other specimens). One pair of eyes placed at level of posterior margin of scutum. Scutum wider than long; anterior border slightly concave, posterior margin with small concavity. Three pairs of scutalae with short setules, AL longest, ML&gt; PL. Anterior sensillae (ASE) ciliated on their distal ¼ length, placed between bases of AL and ML. Posterior sensillae (PSE) ciliated on their distal ¼ length, longer than ASE (Fig. 12).</p>
            <p>Ventral side of idiosoma with a pair of setae 1a placed between coxae I, setae 2a placed between coxae II and four setae placed between coxae III; all these setae with setules. Beyond coxae III 38 barbed setae in holotype (25– 50 in other specimens). NDV = 132 holotype, 93–133 in other specimens. Setae 1b, 2b’, 2b’’, 3b’, 3b’’ located on coxae I, II, III, respectively; all barbed.</p>
            <p>Gnathosoma ventrally with ciliated hypostomal setae bs and shorter than other nude hypostomal setae as (Fig. 13). Dorsally a pair of nude adoral setae cs. Pedipalp formula: O-B-B-BBB-BBBNNωζ. Palpfemur and palpgenu with one setulose seta. Palptibia with three setulose setae (Fig. 13). Palptarsus with five normal setae: two long setae and three short setae (one of them distinctly longer than remaining ones), one solenidion placed in proximal part of segment and one eupathidium placed in distal part of palptarsus (Fig. 14).</p>
            <p>Leg setal formula: Leg I: Ta 1ω, 2ζ, 26 (22–30); Ti 2φ, 1κ, 18; Ge 1σ, 1κ, 12; Tf 5, Bf 4, Tr 1, Cx 1 (Fig. 15– 16). Leg II: Ta 1ω, 2ζ, 23 (24–30); Ti 2φ, 18 (17–19); Ge 1κ, 12; Tf 5; Bf 4: Tr 1; Cx 2 (Fig. 17–18). Leg III: Ta 1ζ, 25 (25–30); Ti 1φ, 18 (17–19); Ge 12; Tf 5; Bf 2; Tr 1; Cx 1 (Fig. 19–20). σI placed at κ, both located distally on genu I.</p>
            <p>Leg lengths: Leg I 874 holotype, 819–927 other specimens; leg II 791, 764–890; leg III 955, 921–1084. IP = 2620 holotype, 2513–2901 other specimens.</p>
            <p>Measurements are given in Table 2.</p>
            <p> Material. 38 larvae from  Z. variegatus , 6 October 2012, Zamakoe (3º34’44’’ N, 11º39’33’’ E) and Mbankomo (3º52’13’’ N, 11º29’42’’ E), Cameroon; collector S. Kekeunou; one larva from  Eupezus rufipes, Bagomojo , Cameroon, date and collector unknown. </p>
            <p> Remarks. To date this species was known only from Madagascar and was obtained from undetermined  Orthoptera (Haitlinger 2007a) . All specimens of  C. justynae from Cameroon were obtained from  Z. variegatus , excluding one specimen obtained from  Eupezus rufipes (  Coleoptera :  Tenebrionidae ). It is the second record of  Charletonia from  Tenebrionidae . Previously  C. aerolata was found on undetermined  Tenebrionidae (Beron 2008) . Specimens of  C. justynae from Cameroon differ from specimens of  C. justynae from Madagascar in some characters, especially in 2b’, fD and fV (Table 2). Tarsi I–III indicate high variability in the number of normal setae (Ta I 22–30, Ta II 24–30, Ta III 24–30); also tibiae II and III have a variable number of setae. These facts indicate that both characters, especially number of normal setae on all tarsi, have unimportant taxonomic value and should not be used in keys as for example by Khanjani et al. (2012). </p>
            <p> Charletonia justynae belongs to the species group with four setae between coxae II–III, Ta I 160–200 Μm, and Ti III 240–310 Μm. This group includes:  C. adellae Haitlinger, 2007 ,  C. arlettae Haitlinger, 1987 ,  C. falcata Southcott, 1966 ,  C. oudemansi Southcott, 1966 and  C. barbata Southcott, 1966 (Southcott 1966; Haitlinger 1987, 2007a). Moreover, partly these dimensions are shared with  C. aerolata (Trägårdh, 1908) ,  C. enghoffi Southcott, 1991 and  C. beninensis Haitlinger, 2007 (Trägårdh 1908; Southcott 1991; Haitlinger 2007b). It differs from  C. adellae in the longer PsFd (74–90 vs. 62–74) and σI placed at microseta κ, both distally located on genu I and beyond normal setae vs. σI placed relatively far from microseta κ and before three last normal setae and famulus ε on Ta I present vs. famulus ε on Ta I absent; from  C. arlettae in the shorter ML (61–69 vs. 70–86), PL (51–63 vs. 66–72), 1a (37–52 vs. 54–64), longer Ta I (169–194 vs. 160), Ti I without seta Cp vs. Ti I with seta Cp, palptarsus with 3B, 2N normal setae vs. 4N normal setae; from  C. falcata in the shorter MW (85–108 vs. 119), PW (107–124 vs. 136), ISD (64–74 vs. 89), L (100–115 vs. 133), W (112–136 vs. 140), Ti I (193–244 vs. 289), Ti III (245–296 vs. 310) and fD (65–94 vs. 44); from  C. oudemansi in the longer L (100–115 vs. 77–92), AP (45–54 vs. 33–37), Ta I (169–194 vs. 158), shorter 1a (37–52 vs. 85) and 1b (72–100 vs. 110); from  C. barbata in the longer AL (72–82 vs. 56–65), ML (61–69 vs. 47–58), 3b’’ (51–68 vs. 43), shorter Ti I (193–244 vs. 256), TiIII (245–296 vs. 317), Ti I without Cp vs. Ti I with Cp and cs nude vs cs barbed; from  C. aerolata in the longer PW (107–124 vs. 91–103), W (112–136 vs. 101–111), AL 72–82 vs. 38–55), ML (61–69 vs. 40–53) and Ta I (169–194 vs. 140–172); from  C. enghoffi in the shorter ISD (64–74 vs. 73–77), ASE (52–70 vs. 70–75), PSE (79–110 vs. 116–129), 1a 37–52 vs. 69–75), longer Ta I (169–194 vs. 156–164), Ta II (154–188 vs. 131–149), Ta III (171–195 vs. 153–167) and σI placed distally to most distal normal setae vs. σI placed proximally to most distal normal setae and from  C. beninensis in the longer AL (72–82 vs. 66–70), Ta I (169–194 vs. 156–170), Ta II (154–188 vs. 150–156), Ta III (171–195 vs. 162–174), IP (2513–2906 vs. 2346–2552), palptarsus with 3B, 2N normal setae vs. palptarsus with 4N normal setae, tarsus I with famulus ε vs. tarsus I without famulus ε, Ti I without Cp vs. Ti I with Cp, fn Ta II (23–30 vs. 20), fn Ta III (25–30 vs. 21) and anterior border straight vs. anterior border slightly concave. </p>
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	https://treatment.plazi.org/id/03AF182F1B24051723A9FA23FAF72F5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Haitlinger, Ryszard;Kekeunou, Sévilor;Łupicki, Dariusz	Haitlinger, Ryszard, Kekeunou, Sévilor, Łupicki, Dariusz (2014): Charletonia cameroonensis Haitlinger & Kekeunou sp. nov. and the first record of C. justynae Haitlinger, 1987 (Acari: Erythraeidae) from Cameroon with redescription of the species. Zootaxa 3760 (1): 39-53, DOI: 10.11646/zootaxa.3760.1.2
