taxonID	type	description	language	source
162FACF8A3F98CB77CF8B2D2127CB7FD.taxon	distribution	Distribution. Palaearctic, north Oriental.	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
54D7AC4681C3B93813F505921369B53E.taxon	distribution	Distribution. Holarctic, north Oriental, north Neotropical.	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
7513778A635B0D2C6579764837A26A5E.taxon	materials_examined	Material examined. Type material. Holotype of Ptomaphagus kuntzeni: ♂, [JAPAN] Hagi [ca. 34 ° 24 ' N, 131 ° 23 ' E] / Hiller [R. leg., probably collected during 1872 - 1875 (Esaki 1935)] // Type // 59051 // Ptomaphagus / kuntzeni Type / det. K. Sokolowski (ZMHB). Holotype of Ptomaphagus amamianus: ♂, HOLOTYPE // [JAPAN] NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E] / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // HOLOTYPE / Appended label by / N. Kobayashi / 2008 // 0000005731 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM). Allotype of Ptomaphagus amamianus: 1 ♀, ALLOTYPE // [JAPAN] NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E] / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005732 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM). Paratypes of Ptomaphagus amamianus: 1 ♂, PARATYPE // [JAPAN] NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E], / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus amamianus Nak. / Det. T. Nakane // MHNG / ENTO / 00003333 (MHNG); 1 ♀, same data as previous except: 00003334 (MHNG); 1 ♂, PARATYPE // [JAPAN] NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E] / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005733 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM); 1 ♀, [JAPAN] PARATYPE // // NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E] / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005734 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM); 1 ♂, PARATYPE // [JAPAN] NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E], AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus amamianus Nak. / Det. T. Nakane // 37 - 1 [Pl. 37, fig. 1 in Nakane 1963] // NAKANE Coll. / SEMU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005747 / Sys. Ent Hokkaido Univ. / Japan [SEHU] (HUM).	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
7513778A635B0D2C6579764837A26A5E.taxon	materials_examined	Additional material. JAPAN: Honshu: 3 ♂♂ 1 ♀, Botanic Garden [ca. 38 ° 15 ' N, 140 ° 51 ' E] / Sendai, Miyagi [Pref.] / 1. IX. 1966 / A. NARITA // Ptomaphagus / kuntzeni / Sokolowski / Det. T. Nakane (HUM); 2 ♂♂, Yoshi-ga-hira [ca. 37 ° 26 ' N, 139 ° 7 ' E] / Niigata Pref. / 14. VIII. 1987 / leg. M. NISHIKAWA // Trap // alt. 600 m (CMNE); 1 ♂, ASAMIGAWAKEI / KOKU [ca. 37 ° 13 ' N, 140 ° 57 ' E] HIRONOMACHI / FUKUSHIMA [Pref.] 1990. VIII. 2 / A. IZUMI leg. (CMNE); 1 ♀, Mt. TORINOKO-YAMA [ca. 36 ° 41 ' N, 140 ° 14 ' E] / 馬頭町 [Bato-machi], 栃木県 [Tochigi Pref.] / 30. APL. 1996 / H. OHKAWA leg. // 14 (CMNE); 1 ♂, NAGAEDANI [ca. 36 ° 34 ' N, 136 ° 41 ' E] / KANAZAWA [Ishikawa Pref.] / 13. V. 1948 / S. TAKABA [leg.] (CYHK); 1 ♂, Mt. TAKAO [- YAMA: ca. 36 ° 27 ' N 136 ° 45 ' E] / KAGA [= Ishikawa Pref.] / 2. VII. 1961 / Y. Hayashi [leg.] // Ptomaphagus / kuntzeni / Sokolowski? / Det. Y. Hayashi, 1979 (CYHK); 1 ♂ 1 ♀, Tohbu [ca. 36 ° 23 ' N, 138 ° 21 ' E], Chiisagata / Nagano [Pref.], JAPAN / 1 st, June, 1986 / legit. T. Abe (CMNE); 3 ♀♀, ISK. [= Ishikawa Pref.] Ishikawa Co. / Shiramine Vil. Shiramine [ca. 36 ° 09 ' N, 136 ° 37 ' E] / N. of Mt. Ohnadare / 13. VI- 6. VIII. 1988 // (rotten chicken trap) / 630 m, alt. / K. Katsura & / Y. Nishikawa leg. // ♀ (CJRZ); 4 ♂♂, ISK. [= Ishikawa Pref.] Ishikawa Co. / Shiramine Vil. Katarashi [ca. 36 ° 09 ' N, 136 ° 38 ' E] / NW. of Mt. Arigata-yama / 13. VI- 6. VIII. 1988 // (rotten chicken trap) / 580 m, alt. / K. Katsura & / Y. Nishikawa leg. // ♂ // Ptomaphagus kuntzeni SOKOLOWSKI / Det. K. Harusawa, 1993 (CJRZ); 1 ♂ 1 ♀, 石川県 [Ishikawa Pref.] 白山市 [Hakusan-shi] / 白山系 [Mts. Hakusan] / 猿壁堰堤 [Sarukabe-entei: ca. 36 ° 06 ' N, 136 ° 42 ' E] pit fall traps / 8 月 2 - 22 日 2002 年 [2 - 22. VIII. 2002] / 保科英人 [Hoshina, Hideto] 採集 [leg.] (CHHF); 1 ♂, GOZAISHI SPA [ca. 35 ° 43 ' N, 138 ° 21 ' E] / YAMANASHI [Pref.] / 12. VIII. 1989 / TATEO ITO [leg.] (CYHK); 1 ♂, 1981 - 7 - 4 [= 4. VII. 1981] / 日川林道上部 [Upper area of Hikawa-rindo: ca. 35 ° 43 ' N, 138 ° 50 ' E] // 大ボサツ [Mt. Daibosatsu-rei, Yamanashi Pref.] / 腐敗オサトラップ [decayed bait of carabid trap] // K. HAGA [leg.] (CMNE); 2 ♂♂, KAMIHIKAWA-RINDO [ca. 35 ° 43 ' N, 138 ° 50 ' E] / ([Mt.] DAIBOSATSU [- REI]) / Yamanashi Pref. / Aug. 29 - 30 th 1982 / Y. Shibata leg. (CMNE); 2 ♂♂, 東京都西多摩郡 [Tokyo, Nishitama-gun] / 檜原村本宿 [Hinohara-mura, Motoshuku: ca. 35 ° 43 ' N, 139 ° 08 ' E] // Japan: Honshu / Motoshuku, / Hinohara Mura, / Tokyo To 12. IV. 2008 / KAMEZAWA Hiromu leg. // KAMEZAWA Collection (CMNE); 1 ♂ 1 ♀, Mt. Takao [- san: ca. 35 ° 37 ' N, 139 ° 15 ' E] / Hachioji, Tokyo / 16. X. 1982 / M. Nishikawa leg. // Trap (CMNE); 4 ♂♂ 1 ♀, Mt. Mitsutoge-yama [ca. 35 ° 32 ' N, 138 ° 49 ' E] / 1200 m in alt., trap / Kawaguchiko-machi // Yamanashi Pref., C. / Japan, 19. VIII. 1993 / M. Nishikawa leg. (CMNE); 1 ♂, Dohshigawa [Riv.: ca. 35 ° 32 ' N, 139 ° 06 ' E] / Aone, Tsukui Co. / Kanagawa [Pref.], JAPAN / 18 th, May, 1986 / T. Abe & A. Sasai [leg.] (CMNE); 2 ♀♀, (Dodaira [ca. 35 ° 28 ' N, 139 ° 10 ' E]) / E. Tanzawa / Kanagawa Pref. // 11. VII. 1993 / leg. M. NISHIKAWA // carrion trap / alt. ca. 1000 m (CMNE); 1 ♂ 1 ♀, same data as previous except: 21. VIII. 1993 (CMNE); 1 ♂, [JAPAN] Honshu, / Kanagawa Pref., Atsugi City, / Shimofurusawa [ca. 35 ° 27 ' N, 139 ° 19 ' E], / 23. III. 2007. Bait trap. / Takuya FUKUZAWA leg. (CMNE); 1 ♀, same data as previous except: 22. III. 2007 (CMNE); 2 ♀♀, Idenzawa [ca. 35 ° 26 ' N, 138 ° 58 ' E] / Nishitanzawa / Kanagawa [Pref.], Japan // 31. V. - 6. VI. 2006 / leg. T. Watanabe (CMNE); 1 ♂, Mt. OHYAMA [ca. 35 ° 25 ' N, 139 ° 14 ' E] / Kanagawa [Pref.], Japan / June 15 th, 1974 / Coll. Y. Shibata (CMNE); 1 ♀, Hachikita-kogen [ca. 35 ° 24 ' N, 134 ° 32 ' E] / Muraoka-machi / [A- 1] baited-trap / Hyogo Pref. // W Honshu. Japan / 23. XI. 2004 / Shigeru Yoshida leg. // 兵庫県村岡町 [Hyogo Pref., Muraoka-machi] / ハチ 北高原 [Hachikita-kogen] A- 1 / bait-trap / 23. XI. 2004 / 吉田 茂 [Yoshida Shigeru] 採集 [leg.] (CMNE); 1 ♂, Mt. Mikuni-yama [ca. 35 ° 24 ' N, 138 ° 54 ' E] / Nishitanzawa / Kanagawa [Pref.], Japan / 3 - 7. VIII. 2003 / T. Watanabe leg. // Flight / Interception / Trap (CMNE); 1 ♀, (Near OHKURA [ca. 35 ° 24 ' N, 139 ° 10 ' E]) / Tanzawa [Mts.], Kanagawa [Pref.] / Apr. 29 th, 1973 / Coll. Y. Shibata (CMNE); 3 ♂♂ 9 ♀♀, Kitanisawa 750 m alt. [ca. 35 ° 19 ' N, 133 ° 33 ' E] / Mitsukue, Kofu / Hino-gun, Tottori // Pref., W Japan / 17. VI. 2007; b. trap / Y. Fujitani leg. // 鳥取県江府町 [Tottori Pref., Kofu-cho] / 御机 [Mitsukue] 木谷沢 [Kitanisawa] / bait-trap 750 m (CMNE); 1 ♂ 2 ♀♀, Takahachi [ca. 35 ° 19 ' N, 138 ° 43 ' E] on Mt. / Fuji (cola trap) / Shizuoka Pref. // Central Japan / 14. VII. - 24. VIII. 1996 / M. Nishikawa leg. (CMNE); 1 ♂, [Mt.] Fudohyama [ca. 35 ° 19 ' N, 139 ° 11 ' E] / Nakai-machi / Kanagawa [Pref.], Japan / 3 - 12. VI. 2006 / T. Watanabe leg. (CMNE); 1 ♂, Hisagi [35 ° 18 ' N, 139 ° 34 ' E] / Zushi c. / Kanagawa pref. / 18 - 23. V. 1983. / pit-fall Trap: No. 32 / No. 2 (CMNE); 5 ♂♂ 12 ♀♀, Yakodani [ca. 35 ° 17 ' N, 134 ° 18 ' E] / Chizu-machi / Yazu-gun, Tottori / Pref., W Honshu, Jpn / 3. VI. 2007 / Y. Fujitani leg. // 鳥取県八頭郡 [Tottori Pref., Yazu-gun] / 智頭町八河谷 [Chizu-machi, Yakodani] / bait-trap 750 m (CMNE); 6 ♂♂ 3 ♀♀, Near Nontaki waterfall [ca. 35 ° 15 ' N, 134 ° 08 ' E] / ca. 800 m in alt. / Aba-son, Tomata-gun / Okayama Pref., Japan / 8. IX. 2002; trap / Y. Fujitani leg. (CMNE); 4 ♂♂ 2 ♀♀, Mt. Yamanori-yama [ca. 35 ° 14 ' N, 133 ° 49 ' E] / Chuka-son, Maniwa- / ca. 900 m in alt. / gun, Okayama Pref. // W Honshu, W Japan / 11. VII. 2004; FIT / Y. Fujitani leg. (CMNE); 2 ♂♂ 2 ♀♀, Daruga-mine [ca. 35 ° 12 ' N, 134 ° 22 ' E] / Nishiawakura-son / 1,100 m in alt. / Aida-gun // Okayama Pref. / 7 - 20. VII. 2006; FIT / Akihiko Watanabe leg. (CMNE); 1 ♂, Akazaigawa Val. [ca. 35 ° 12 ' N, 134 ° 31 ' E] / HYOGO Pref. / 12. V. 1979 / Y. HAYASHI [leg.] // Ptomaphagus kuntzeni SOKOLOWSKI / Det. Y. HAYASHI, 2014 (CCBW); 1 ♂ 1 ♀, same data as previous except: 15. VII. 1979 (MNHA); 1 ♀, nr. Mt. Suzuga-take [ca. 35 ° 11 ' N, 136 ° 27 ' E] / Fujiwara-cho, Mie Pref. / 3. V. 2002 / Shiho Arai leg. // Ptomaphagus / kuntzeni / Sokolowski, 1957 / Det. M. Nishikawa, 2014 / # MNIC 123909 Ch 1 S (CMNE); 1 ♀, 千葉県 [Chiba Pref.] 君津市 [Kimitsu-shi] / 郷台畑 [Godaihata] / 猪の川 [Inokawa] / 滝の沢出合 [Takinosawa Deai: 35 ° 11 ' N, 140 ° 06 ' E] / malaise traps / 5 月 13 - 20 日 1997 年 [13 - 20. V. 1997] / 新田 N. [Nitta, N.] 採集 [leg.] (CHHF); 1 ♂ 2 ♀♀, MIZUHO T. [ca. 35 ° 10 ' N, 135 ° 22 ' E] / KYOTO, / 22. V. 1984 / Y. HAYASHI [leg.] // trap // Ptomaphagus kuntzeni SOKOLOWSKI / Det. Y. HAYASHI, 2014 (1 ♂ in CCBW and 2 ♀♀ in CYHK); 1 ♂ 1 ♀, same data as previous except: 26. V. 1984 (CCBW); 1 ♂ 3 ♀♀, same data as previous except: シズシ [Shizushi: 35 ° 13 ' N, 135 ° 20 ' E] // [bar code omitted] (CYHK; MNHA); 1 ♂, [Mt.] Takiyama [ca. 35 ° 09 ' N, 134 ° 09 ' E] / Nagi, Katsuta-gun / Okayama Pref. // W. Honshu, Japan / 9. V. 2002 / Y. Fujitani leg. (CMNE); 8 ♂♂ 3 ♀♀, same data as previous except: 7. VII. 2002 (CMNE); 3 ♂♂ 4 ♀♀, same data as previous except: 27. X. 2002 (CYFO); 1 ♂, same data as previous except: 10. XI. 2002 (CYFO); 1 ♀, same data as previous except: 23. XI. 2002 (CYFO); 1 ♀, same data as previous except: 29. XII. 2002 (CYFO); 19 ♂♂ 23 ♀♀, same data as previous except: 27. VI. 2003 (CYFO); 1 ♀, JAPON KYOTO / Seryo-Toge Kyoto [ca. 35 ° 09 ' N, 135 ° 45 ' E] / 500 - 600 m 6. VIII. 1980 / Cl. Besuchet [leg.] (MHNG); 1 ♀, SASAYAMA T. / 雨石山 [Mt. Amaishi-yama: ca. 35 ° 07 ' N, 135 ° 20 ' E] / HYOGO [Pref.], / 28. IV. 1984 / Y. HAYASHI [leg.] (CYHK); 2 ♂♂ 3 ♀♀, nr. Rashomon-daiichi- / do Cave [ca. 34 ° 56 ' N, 133 ° 33 ' E], Niimi-shi. / 370 m in alt. / Okayama Pref. // W Japan (carrion trap) / 18. VIII. 2001 / Y. Fujitani leg. (CMNE); 1 ♂ 1 ♀, Shiramizu [ca. 34 ° 58 ' N, 134 ° 15 ' E] / Mimasaka-shi / [carrion baited-trap] / Okayama Pref. // W Honshu, Japan / 7. V. 2005 / Y. Fujitani leg. (CMNE); 4 ♂♂ 1 ♀, Mumyodani [ca. 34 ° 56 ' N, 133 ° 27 ' E], ca. 350 m / Tetta-cho, Atetsu- / gun, Okayama Pref. // W Honshu, W. Japan / 25. IV. 2004; b. trap / Y. Fujitani leg. (CMNE); 1 ♀, 箕面市下止々呂美 [Mino-shi, Shimotodoromi: ca. 34 ° 52 ' N, 135 ° 27 ' E] / (OSAKA pref.) / 4. VIII. 1992 / leg. 齋藤琢己 [Saito Takumi] // Ptomaphagus / kuntzeni / SOKOLOWSKY / Det. Y. HAYASHI, 1993 (CYHK); 1 ♂ 4 ♀♀, Kanahira National / Forest, [ca. 34 ° 47 ' N, 133 ° 37 ' E] Takahashi- / [carrion baited-trap] / shi, Okayama Pref. // W Honshu, Japan / 16. IV. 2006 / Y. Fujitani leg. (CMNE); 1 ♂, Anatoyama-jinja [ca. 34 ° 39 ' N, 133 ° 29 ' E] / 450 m alt., Kawakami / Okayama Pref. // Honshu, W Japan / 22. IV. 2001; trap / Y. Fujitani leg. (CMNE); 1 ♀, same data as previous except: 8. IV. 2001 (CMNE); 1 ♀, same data as previous except: 16. IV. 2001 (CMNE); 1 ♂ 3 ♀♀, same data as previous except: 28. IV. 2001 (CMNE); 1 ♀, Omukai-rindo [ca. 34 ° 29 ' N, 132 ° 07 ' E] / Chojabara, Yoshiwa / 880 m in alt. / Hatsukaichi-shi // Hiroshima Pref., W / Honshu, Jpn. 28. IV. / 2007, M. Tagami leg. // 広島県廿日市市 [Hiroshima Pref., Hatsukaichi-shi] / 吉和長者原大向林道 [Yoshiwa, Chojabara, Omukai-rindo] / FIT 880 m / 28. IV. 2007 / 田上雅生 [Tagami Masao] 採集 [leg.] (CMNE); 1 ♂ 1 ♀, OSAKA Pr. Tondabayashi c. / Kannobi Kongo Colony [ca. 34 ° 29 ' N, 135 ° 35 ' E] / 12 - 26. IV. 1994 / leg. K. Harusawa // pit fall trap: / rotten squid // Ptomaphagus kuntzeni Sokolowski (CMPR); 6 ♂♂ 3 ♀♀, Kasayama, ca. 50 m / Hagi-shi, / ca. 34 ° 26 ' N, 131 ° 24 ' E / Yamaguchi Pref. // W Honshu, W Japan / 29. VI. 2002; trap / Y. Fujitani leg. (CMNE); 2 ♀♀, same data as previous except: 5. VII. 2002 (CMNE); 2 ♂♂ 2 ♀♀, Mt. Makio [- san: ca. 34 ° 23 ' N, 135 ° 30 ' E] / OSAKA [Pref.] / 4. VIII. 1983 / F. KIMURA [leg.] (CYHK); 1 ♂, [OSAKA Pr.] / Kawachinagano / 天見 [Amami], alt. 260 m / 6 - 14. X. 1981 / K. Harusawa / Y. Nishikawa leg. // 出合 [Deai] ~ 流谷 [Nagaredani] / 流谷八幡境内 [Nagaredani-hachiman shrine's area: ca. 34 ° 23 ' N, 135 ° 35 ' E] / 腐肉 [carrion] Trap (牛 [beef]) // Ptomaphagus / kuntzeni / SOKOLOWSKI / Det. Y. HAYASHI, 1990 (CYHK); 1 ♂, Kamisuga [ca. 34 ° 23 ' N, 136 ° 22 ' E] / Odai-machi, Taki- / gun, Mie Pref. // Honshu, Japan / 22 - 9. V. 2005 / Katsumi Akita leg. (CMNE); 3 ♂♂ 2 ♀♀, Kanoashikochi [ca. 34 ° 22 ' N, 131 ° 57 ' E] / Muikaichi-machi / 450 m in alt. / Shimane Pref. // W Honshu, Japan / 10. VI. 2005 / Y. Fujitani leg. // 六日市町 [Muikaichi-machi] / 鹿足河内 [Kanoashikochi] (CMNE); 1 ♂, HASE [ca. 34 ° 19 ' N, 135 ° 48 ' E] / YAMATO [= Nara Pref.] / 5. VI. 1966 / Y. HAYASHI [leg.] // Ptomaphagus / kuntzeni (CYHK); 1 ♂ 4 ♀♀, Nodani [ca. 34 ° 17 ' N, 131 ° 38 ' E], Tokuji-cho / Saba-gun / Yamaguchi Pref. // W Honshu, W Japan / 21. IV. 2004; b. trap / Y. Fujitani leg. (CMNE); 2 ♀♀, mouth of Nakao-do / Cave [ca. 34 ° 16 ' N, 131 ° 17 ' E]. Aokage / Shuho, Yamaguchi // Pref., W Japan / 28. VII. - 4. VIII. 2002 / Y. Fujitani leg. (CMNE); 3 ♂♂ 5 ♀♀, Akiyoshidai [ca. 34 ° 16 ' N, 131 ° 18 ' E] / Shuho-machi / Yamaguchi Pref. // W Honshu, W Japan / 12. IV. 2002; trap / Y. Fujitani leg. (CMNE); 1 ♂ 2 ♀♀, Mt. Nagano-yama [ca. 34 ° 16 ' N, 131 ° 52 ' E] / 1,010 m in alt. / b. trap / Kano-machi // Yamaguchi Pref. / Japan, 4. VI. 2005 / Y. Fujitani leg. (CMNE); Japan. / G. Lewis. / B. M. 1926 - 369. // Chiuzenji [ca. 36 ° 46 ' N, 139 ° 28 ' E]. // Ptomaphagus / sibiricus Jean. / Jeannel det. // Ptomaphagus / kuntzeni ♀ / det. WANG C. - B., 2016 (BMNH). Kyushu: 1 ♂, Hata [ca. 33 ° 48 ' N, 130 ° 46 ' E] / Yahata-City [Fukuoka Pref.] / 3. V. 1965 / coll. M. Ueda (HUM); 1 ♂, 福岡県 [Fukuoka Pref.] 添田町 [Soeda-cho] / 英彦山 [Mt. Hiko-san: ca. 33 ° 29 ' N, 130 ° 54 ' E] / 5 月 2 日 1983 年 [2. V. 1983] / 野村 S. [Nomura S.] 採集 [leg.] (CHHF); 1 ♀, (Kinsenji [ca. 32 ° 58 ' N, 130 ° 05 ' E], Mt. Tara [- dake]) / Nagasaki Pref. / Kyushu, JAPAN / 10. V. 1983 / S. Imasaka leg. // Ptomaphagus (s. str.) / kuntzeni Sokolowski, 1957 / Det. M. Nishikawa, 2010 / ♀ (CMNE); 1 ♂, Mt. Gokabaru-dake [ca. 32 ° 57 ' N, 130 ° 04 ' E] / (1,058 m in alt.) / Nagasaki Pref. // NW Kyushu, Japan / 11 - 12. V. 1991 / M. Nishikawa leg. // carrion trap / alt. ca. m (CMNE); 2 ♂♂, Unzen-bessho [ca. 32 ° 44 ' N, 130 ° 15 ' E] / Nagasaki Pref. / 1. VI. 1983 / leg. S. Imasaka (CMNE); 1 ♂, Hagi [ca. 32 ° 32 ' N, 130 ° 56 ' E], Gokanosho / Kumamoto Pref. / 18. VI. 1984 / leg. S. Imasaka (CMNE); 1 ♀, Takao [ca. 32 ° 28 ' N, 131 ° 08 ' E], Shimofukura / Shiiba, Miyazaki Pref. / Kyushu, SW Japan / 26. VII. 2015 / Takashi Watanabe leg. // Ptomaphagus (Ptomaphagus) / kuntzeni Sokolowski, 1957 / Det. M. Nishikawa, 2016 / MNIC 124950 Ch 1 S ♀ (CMNE); 1 ♂, Inao dake [ca. 31 ° 07 ' N, 130 ° 53 ' E] / Tashiro cho / Kagoshima Pref. / 14. VII. 1985 / T. TANABE leg. // alt. 300 m // Trap // Ptomaphagus / kuntzeni / Sokolowski / Det. T. Nakane (HUM); 1 ♂, same data as previous except: 575 m (HUM); 1 ♀, same data as previous except: 610 m (HUM). Ryukyus: Amami-Oshima Is.: 1 ♀, 鹿児島県 [Kagoshima Pref.] / 奄美市 [Amami-shi] / 龍郷町 [Tatsugo-cho] / 芦徳 [Ashitoku: ca. 28 ° 25 ' N, 129 ° 36 ' E] / 05. V. 2011 / 稲垣政志 [Inagaki Masashi] Leg. (CYHK); 1 ♂ 1 ♀, (JAPAN) Kagoshima pref., / Amami-Ohshima Is., / Honcha pass. [ca. 28 ° 23 ' N, 129 ° 33 ' E] / 11 - 14. IV. 2007 / T. FUKUZAWA et al. [leg.] (CMNE); 1 ♀, NAZE [ca. 28 ° 22 ' N, 129 ° 29 ' E] / AMAMI [- OSHIMA] IS. / 4. V. 1960 / T. Shibata [leg.] // Ptomaphagus / ohshimensis / NAKANE / Det. Y. HAYASHI, 1969 (CYHK); 2 ♀♀, same data as previous but no det. label (CYHK); 1 ♀, same data as previous except: // Ptomaphagus / amamianus / NAKANE / Det. Y. HAYASHI, 2009 // Collection of / Y. HAYASHI // [bar code omitted] (MNHA); 2 ♂♂ 4 ♀♀, Oganeku / 28.3598 N 129.3403 E [ca. 28 ° 21 ' N, 129 ° 20 ' E]; 260 m alt. / Yamato-son / Amami-Oshima Is., Ryukyus / Kagoshima Pref., SW Japan / 27. II. - 2. III. 2016; trap / M. Nishikawa leg. (CMNE); 4 ♀♀, nr. Hokoku-jinja, ca. 350 m / 28.362331 N 129.482009 E [ca. 28 ° 21 ' N, 129 ° 28 ' E] / Amami-shi / Amami-Oshima Is. / Kagoshima Pref., Ryukyus / SW Japan / 27. II. - 2. III. 2016; trap / M. Nishikawa leg. (CMNE); 3 ♂♂ 4 ♀♀, around Okawa Dam, ca. 28 ° 20 ' N, 129 ° 29 ' E, 130 m alt., Amami-shi, Amami-Oshima Is. / Kagoshima Pref., Ryukyus / SW Japan / 27. II. - 2. III. 2016; trap / M. Nishikawa leg. (CMNE); 1 ♂, Toen, nr. Amami ForestPolis / 28.3176 ° N 129.3297 ° E [ca. 28 ° 19 ' N, 129 ° 19 ' E]; 180 m alt. / Yamato-son, Amami-Oshima Is. / Oshima-gun, Kagoshima Pref. / Ryukyus, SW Japan / 3 - 10. III. 2015; bait trap / M. Nishikawa leg. (CMNE); 11 ♂♂ 18 ♀♀, (JAPAN) Kagoshima Pref., / Amami-Oshima Is., / Yamato vil., Odana. 310 m alt. / N 28 ° 18 ' 40.2 E 128 ° 55 ' 32.2 / 5. III. 2008. Carrion Baited trap. / Takuya FUKUZAWA leg. (CMNE); 8 ♂♂ 5 ♀♀, Naon-kengyozorin Forest [ca. 28 ° 18 ' N, 129 ° 20 ' E] / 名音県行造林 [Naon-kengyozorin Forest]; alt. 349 m / 28.3058 ° N, 129.3371 ° E / Yamato, Amami-Oshima Is. // Oshima-gun, Kagoshima Pr. / Ryukyus, SW Japan / 2 - 10. III. 2015; bait trap / M. Nishikawa leg. // Ptomaphagus (Ptomaphagus) / amamianus Nakane, 1963 / Det. M. Nishikawa, 2015 (CMNE); 1 ♂, Mt. Yuwan [- dake: ca. 28 ° 17 ' N, 129 ° 19 ' E] / Amami-Ohshima Is / 29. III. 1999 // Ptomaphagus amamianus NAKANE / Det. Y. HAYASHI, 2014 (CCBW); 1 ♀, same data as previous except: 28. IV. 2000 / 江本健- [Emoto, Ken'ichi] 採集 [leg.] // ♀ (CMNE); 1 ♀, same data as previous except: 7 - 12. V. 2006 / T. Watanabe leg. (CMNE); 1 ♂ 3 ♀♀, same data as previous except: // 650 - 680 m, / 18 - 26. III. 2010, baited / pitfall traps, forest, / Tomas Lackner leg. (CJRZ); 9 ♂♂ 9 ♀♀, (JAPAN) Kagoshima Pref., / Amami-Oshima Is., sumiyou Vil., santarou Touge Pass, 318 m alt. / N 28 ° 17 ' 03.9 E 129 ° 25 ' 16.2 / Carrion Baited Trap. 6. III. 2008. / Takuya FUKUZAWA leg. (CMNE); 2 ♂♂ 1 ♀, same data as previous except: 12 - 14. IV. 2007 (CMNE); 1 ♂ 1 ♀, same data as previous except: 13 - 16. IV. 2007. FIT / T. FUKUZAWA et al. [leg.] (CMNE); 4 ♂♂ 2 ♀♀, 鹿児島県大島郡 [Kagoshima Pref., Oshima-gun] / 宇検村 [Uken-son] ヤクガチョボシ 山麓 [foot of Mt. Yakugachoboshi-yama: ca. 28 ° 15 ' N, 129 ° 21 ' E] // Japan; Ryukyu / foot of Yakugachoboshi-yama, / Uken Son, Oshima Gun, / Kagoshima Ken // (Is. Amami-o-shima) / 27. II. 2004 / KAMEZAWA, Hiromu leg. (CMNE); 1 ♂, HATSUNO [ca. 28 ° 15 ' N, 129 ° 22 ' E] / Amami [- Oshima] Is. / 31. III. 1969 / H. NOMURA [leg.] // Ptomaphagus amamianus NAKANE / Det. Y. HAYASHI, 2014 (CCBW); 1 ♂, same data as previous except: 1. IV. 1967 (CYHK); 1 ♀, same data as previous except: 3. IV. 1967 (CYHK); 1 ♂, same data as previous except: 4. IV. 1967 (CYHK); 2 ♂♂ 1 ♀, same data as previous except: 3. IV. 1969 // K. TANIZAWA [leg.] (CCBW); 2 ♂♂ 1 ♀, same data as previous except: // Ptomaphagus / amamianus NAKANE / Det. Y. HAYASHI, 1993 (CMNE); 1 ♂ 1 ♀, same data as previous except: 31. III. 1969 / H. NOMURA [leg.] (CYHK); 4 ♂♂, same data as previous except: 1. IV. 1969 (CYHK); 4 ♂♂ 1 ♀, same data as previous except: 2. IV. 1969 (CYHK); 1 ♂ 1 ♀, same data as previous except: 3. IV. 1969 (CYHK); 7 ♂♂ 3 ♀♀, same data as previous except: K. Tanizawa [leg.] (CYHK); 1 ♀, same data as previous except: 5. V. 1969 / Y. MAEDA [leg.] (CMNE); 1 ♀, Nishinakama [ca. 28 ° 15 ' N, 129 ° 24 ' E] / AMAMI [- OSHIMA] Isl. / 5. IV. 1969 / K. TANIZAWA [leg.] (CYHK); 3 ♂♂ 3 ♀♀, nr. Yakugachi Tunnel / 28.2295 N 129.3599 E [ca. 28 ° 13 ' N, 129 ° 21 ' E]; 30 m alt. / Kamiyakugachi, Amami-shi / Amami-Oshima Is., Ryukyus / Kagoshima Pref., SW Japan / 27. II. - 2. III. 2016; trap / M. Nishikawa leg. (CMNE); 1 ♀, (由井岳 [Mt. Yui-dake: ca. 28 ° 11 ' N, 129 ° 18 ' E]) / 瀬戸内 [Setouchi-cho] 奄美大島 [Amami-Oshima Is.] / 25. IV. 2000 / 江 本健- [Emoto Ken'ichi] 採集 [leg.] // ♀ (CMNE); 6 ♂♂, Aminoko-toge Pass / 28.1893 ° N 129.3659 ° E [ca. 28 ° 11 ' N, 129 ° 21 ' E]; 350 m alt. / Setsuko, Setouchi-cho / Amami-Oshima Is. / Oshima-gun, Kagoshima Pref. / Ryukyus, SW Japan / 6 - 10. III. 2015; bait trap / M. Nishikawa leg. (CMNE); 1 ♂ 3 ♀♀, 鹿児島県 [Kagoshima Pref.] / 大島郡 [Oshima-gun] / 瀬戸内町 [Setouchi-cho: ca. 28 ° 08 ' N, 129 ° 18 ' E] / 05. V. 2011 / 稲垣政志 [Inagaki, Masashi] Leg. (CYHK); 1 ♂, IKARI [unlocated] / AMAMI [- OSHIMA] IS. / 18. V. 1960 / T. Shibata [leg.] (CYHK). Ryukyus: Kume-jima Is.: 3 ♀♀, Shirase-gawa [Riv.: ca. 26 ° 20 ' N, 126 ° 46 ' E] / Gushikawa-son / Kume-jima Is. // Ryukyus, SW Japan / 15 - 17. III. 1998 / M. Maruyama leg. (CMNE). Ryukyus: Okinawa-honto Is.: 1 ♂ 1 ♀, [Okinawa: JAPAN] / Uka-rindo [ca. 26 ° 48 ' N, 128 ° 14 ' E], alt. 250 - / 300 m, Kunigami vill. / 8. II. 2009 / Takashi Kurihara leg. // ベイトトラップ [bait trap] / 鳥の手羽元 [fowl wing sticks] // Ptomaphagus / kuntzeni? / Y. HAYASHI, 19 2014 (CYHK); 2 ♂♂, same data as previous except 7. II. 2009 (CYHK); 2 ♂♂, Yona [ca. 26 ° 45 ' N, 128 ° 12 ' E] (Ohkuni 5) / Okinawa-jima Is. / Ryukyus // SW Japan / 25 - 27. IV. 1996 / M. Nagano leg. (CMNE); 1 ♀, Nishime-dake [ca. 26 ° 48 ' N, 128 ° 16 ' E]. OKN [= Okinawa Pref.] / Date: 14. X. 2002 / K. MASUMOTO leg. (CMNE); 2 ♀♀, [Okinawa: JAPAN] / Hama-rindo [ca. 26 ° 43 ' N, 128 ° 09 ' E], alt. 50 - / 100 m, Kunigami vill. / 8. II. 2009 / Takashi Kurihara leg. // ベイトトラップ [bait trap] / 鳥の手羽元 [fowl wing sticks] (CYHK); 4 ♂♂, (JAPAN, Ryukyus) / Okinawa pref. / Okinawa Is. / Kunigami Vil., Hiji [ca. 26 ° 43 ' N, 128 ° 10 ' E] / 13. III. 2009. / Takuya FUKUZAWA leg. (CMNE); 1 ♂, 与那覇岳 [Mt. Yonaha-dake: ca. 26 ° 42 ' N, 128 ° 13 ' E] / 沖縄本島 [Okinawa-honto Is.] / 14. iv. 2000 / 保科英人 [Hoshina, Hideto leg.] / 腐肉トラップ [carrion trap] // 2 (CMNE); 7 ♂♂ 5 ♀♀, bank of Haneji-o-kawa / Riv. [ca. 26 ° 36 ' N, 128 ° 01 ' E], 20 - 40 m in alt. / Nago-shi, Okinawa- // jima Is., Ryukyus / (c. trap) 18. IV. 1993 / R. Yakita leg. (CMNE); 13 ♂♂ 7 ♀♀, same data as previous except: 8 - 18. IV. 1993 (CMNE); 14 ♂♂ 9 ♀♀, (JAPAN, Ryukyus) / Okinawa Pref. / Okinawa Is. / Nago City, Genka. / 26 ° 36 ' N, 128 ° 04 ' E // 10 - 13. III. 2009. / by flight Intercept Trap / T. Fukuzawa, T. Ishikawa & M. Kishi leg. (CMNE). Ryukyus: Tokuno-shima Is.: 1 ♂, (JAPAN) Kagoshima Pref. / Tokuno-shima Is., / Amagi Town, / Mt. Yamatogusuku-san [ca. 27 ° 48 ' N, 128 ° 55 ' E] / 29. II. - 4. III. 2008. (FIT) Takuya FUKUZAWA leg. (CMNE); 85 ♂♂ 67 ♀♀, (JAPAN) Kagoshima Pref. / Tokuno-shima Is., / Amagi Town, / Mt. Sankyo-dake, 185 m alt. / N 27 ° 46 ' 13.6, E 128 ° 57 ' 18.0 / Carrion Baited Trap. 4. III. 2008. / Takuya FUKUZAWA leg. (CMNE); 4 ♂♂ 3 ♀♀, same data as previous except: 27. II. - 4. III. 2008. (FIT) (CMNE). Sadoga-shima Is.: 1 ♂, Mt. Donden-yama [ca. 38 ° 07 ' N, 138 ° 22 ' E] / Sado Is., Niigata / Pref., Honshu, Japan // 30. IV. - 26. V. / 1990 / leg. M. NISHIKAWA // carrion trap / alt. ca. 200 m (CMNE). Shikoku: 1 ♀, [SHIKOKU] / Komenono [ca. 33 ° 58 ' N, 132 ° 51 ' E] / Matsuyama [Ehime Pref.] / 25. IV. 1993 / Lizhen Li leg. (EUM); 1 ♂, (Ehime: Japan) / Aonamimachi [ca. 33 ° 53 ' N, 132 ° 51 ' E] / Matsuyama-shi / 24. V. 2006 / Shoma Sejima leg. (EUM); 3 ♀♀, same data as previous except: 2. VI. 2006 (EUM); 2 ♀♀, Minokoshi on Mt. / [ca. 33 ° 51 ' N, 134 ° 05 ' E] / Tsurugi-san / ca. 1400 m in alt. / Tokushima Pref. // Shikoku, Japan / 25. VII. 2004; trap / Y. Fujitani leg. // 德島県東祖谷山村 [Tokushima Pref., Higashiiyayama-son] / 見の越 [Minokoshi] / bait- trap (CMNE); 2 ♂♂ 2 ♀♀, 瓶が森 [Mt. Kamegamori: ca. 33 ° 47 ' N, 133 ° 11 ' E] (alt. 1670 m) / 高知県吾川郡いの町 [Kochi Pref., Agawa-gun, Ino-cho] / 寺川 [Terakawa] 10. V. - 8. VIII. 2009 / 吉田 正隆 [Yoshida Masataka] 採集 [leg.] // Mt. Kamegamori (alt. 1670 / m) Terakawa Ino-cho / Agawa-gun Kochi-Pref. / 10. V. - 8. VIII. 2009 / Masataka YOSHIDA leg. // 鳥ガラトラップ [fowl bone trap] / 地中 [underground] (50 cm) (CMNE); 2 ♂♂ 1 ♀, [SHIKOKU] / Mt. Ishizuchi [ca. 33 ° 44 ' N, 133 ° 06 ' E] / Ehime Pref. / 27. VIII. 1990 / alt. 800 m / S. Takano leg. (EUM); 2 ♂♂, same data as previous except: 13. IX. 1990 / alt. 500 m (EUM); 1 ♂, EHIME [Pref.]; Japan / Mt. Saragamine [ca. 33 ° 43 ' N, 132 ° 53 ' E] / Shigenobu-Town / 22. IV. 1999 / Tatsuya Kan leg. (EUM); 4 ♀♀, 156 JAPAN, Shikoku, / Jshizuchi [Ishizuchi] Mt. Nat. Park, / OMOGO Valley, [ca. 33 ° 42 ' N, 133 ° 05 ' E] 700 m, / S. & J. Peck leg. // 158 mega carrion traps, / mixed warm temperate / forest, 18 - 25. viii. 1980 (CJRZ); 1 ♀, Nishidani [ca. 33 ° 31 ' N, 132 ° 57 ' E] / Yanadani, Ehime [Pref.] / 6 - 8. V. 1994 / Sakai, Li, Aita leg. / (bait-trap) // No. 8 (EUM); 4 ♂♂ 2 ♀♀, (SHIKOKU) / Odamiyama [ca. 33 ° 31 ' N, 132 ° 53 ' E] / Ehime Pref. / 22. VIII. 1984 / E. Yamamoto [leg.] (EUM); 1 ♂ 3 ♀♀, same data as previous except: 14. VIII. 1984 / 山本栄治 [Yamamoto Eiji] 採集 [leg.] (EUM); 1 ♂ 2 ♀♀, Komi [ca. 33 ° 30 ' N, 132 ° 57 ' E] / Yanadani, Ehime [Pref.] / 6 - 8. V. 1994 / Ohbayashi, Nishino, Okada [leg.] / (by bait-trap) // No. 2 (EUM); 1 ♂ 1 ♀, same data as previous except: No. 1 (EUM); 4 ♂♂ 5 ♀♀, Tengu-Kogen [ca. 33 ° 28 ' N, 133 ° 00 ' E] (alt. 1280 m) / Kumakogen-cho Kamiuke- / na-gun Ehime-Pref. / 9. V. - 25. VII. 2010 (Fowl trap) / Masataka YOSHIDA leg. // 愛媛県上浮穴郡 [Ehime Pref., Kamiukena-gun] / 久万高原町天狗高原 [Kumakogen-cho, Tengu-Kogen] / (alt. 1280 m) 9. V. - 25. VII. / 2010 (鶏ガラトラップ [chicken bone trap]) / 吉田正隆 [Yoshida Masataka] 採集 [leg.] (CYHK); 1 ♀, Nagano [ca. 33 ° 27 ' N, 132 ° 56 ' E] / Yusuhara-cho [Kochi Pref.] / 1 - 2. X. 1994 / Y. Utsunomiya [leg.] / (by bait-trap) // No. 9 (EUM). Shimokoshiki-jima Is.: 1 ♀, Mt. Odake [ca. 31 ° 43 ' N, 129 ° 44 ' E] / Is. Shimokoshiki [- jima] / Kagoshima-pref. / 20. VI. 1982 / S. Imasaka leg. (CMNE).	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
7513778A635B0D2C6579764837A26A5E.taxon	distribution	Distribution. China (Taiwan), Japan (Fig. 8),? Myanmar.	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
4725DC14BDA4DEC7FFCD4ED28BDCB3CB.taxon	description	Description. Male. EBL: 2.7 - 3.0 mm (2.7 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.5: 0.8: 0.8: 1.7 mm; width: HW: EW: PW: ELW = 0.7: 0.1: 1.2: 1.3 mm. Proportion of antennomeres from base to tip in µm (length x width): 116 x 61, 91 x 56, 70 x 61, 40 x 66, 43 x 76, 44 x 95, 67 x 102, 32 x 107, 59 x 112, 68 x 113, 121 x 104. Habitus (Fig. 1 D) elongated oval, regularly convex and sublustrous. Well pigmented: mostly brown; mouthparts, apical half of ultimate antennomere, protarsi, and apical parts of meso- and metatarsi more or less paler. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous. Head transverse, HW / HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes small, EW / HW = 0.1. Antennae (Fig. 5 A) slender, AL / HW = 1.1; antennomere III a little shorter than II; VI with length / width = 0.5; XI pear-shaped. Pronotum (Fig. 5 B) transverse, widest just before hind angles, PW / PL = 1.6. Sides gently arched, gradually narrowing from posterior to anterior; hind angles slightly drawn out and bluntly rounded. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles. Elytra oval, widest at approx. basal 1 / 4, ELL / EW = 1.35. Sides weakly arched, gradually narrowing from widest part to apices, which narrowly rounded (Fig. 5 G). Sutural striae present. Metathoracic wings fully developed. Prolegs robust, with basal three protarsomeres (Fig. 5 C) moderately expanded: TW / BTW = 1.5. Protibiae (Fig. 5 E) distinctly expanded towards apex. Profemora broad. Mesotibiae slightly arcuate, mesotarsi simply linear. Metatibiae much thick and straight. Abdominal ventrite VIII (Fig. 5 I) almost round at posterior edge and with a distinct median notch. Spiculum gastrale of genital segment (Fig. 5 J) with nearly 1 / 2 of length protruding beyond anterior edge of epipleurite IX. Aedeagus small, slender and narrow, with median lobe gradually narrowing towards a leaf-shaped apex and terminated by an obtusely rounded knob in dorsal view (Fig. 6 A); opening of genital orifice situated on dorsal surface, deeply cut inwards on left edge of median lobe at subapex. Ventral surface of the apex of the median lobe (Figs 6 B, D) inserted with 4 ventrally oriented setae on the left side and 5 ventrally oriented setae on the right side; parameres narrow, reaching almost apical 1 / 8 of median lobe, each apex (Fig. 6 E) with 2 lateral setae and 1 apical seta relatively shorter. In lateral view (Fig. 6 C), median lobe thin, bent in basal half and almost straight in apical half, gradually tapering apically. Endophallus with stylus quite slender, a cheliform complex just below the base of stylus, and a circular complex at the basal region. Female. Similar to male in general appearance (Fig. 1 E), but distinguished by the following characteristics: protarsi (Fig. 5 D) simply linear; protibiae (Fig. 5 F) narrower; elytral apices (Fig. 5 H) acuminate; ventrite VIII (Fig. 6 F) slightly protruded in median of posterior margin; genital segment as shown in Fig. 6 G; spermatheca (Fig. 6 G) sinuous or coiled in distal part, not coiled in proximal part.	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
4725DC14BDA4DEC7FFCD4ED28BDCB3CB.taxon	distribution	Distribution. Japan (Fig. 9).	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
4725DC14BDA4DEC7FFCD4ED28BDCB3CB.taxon	etymology	Etymology. The specific epithet is from the name of " Piccolo ", a fictional character in the Dragon Ball manga series created by Akira Toriyama, which also has an Italian origin meaning of " small " that refers to the small body size of this new species.	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
BE879D22438CB7746DFECA20D944B368.taxon	materials_examined	Material examined. Type material. Holotype: ♀, [RUSSIA, Far East] Wladiwostok [ca. 43 ° 10 ' N, 132 ° 00 ' E] // Reitter // Coll. Koltze // Pt. variicornis / Rosenh. // Ptomaphagus sibiricus Jeann. / type / R. Jeannel det. // DEI Muencheberg / Col - 07069 (SDEI).	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
BE879D22438CB7746DFECA20D944B368.taxon	distribution	Distribution. Russia (Far East).	en	Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel, Hayashi, Yasuhiko (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan. ZooKeys 607: 119-144, DOI: http://dx.doi.org/10.3897/zookeys.607.9074, URL: http://dx.doi.org/10.3897/zookeys.607.9074
