taxonID	type	description	language	source
D86686AF155DD98871E6163DE3E98349.taxon	diagnosis	Diagnosis. Distinct from all allied taxa by a combination of sequence data (nrITS and nrLSU), reddish- or brownish-grey to purplish-grey, then brown to reddish-brown pileus in basidiomata, absence of olive or violet tinges on stipe surface, presence of angular pores, stipe without reticulum or rarely with a faint reticulum restricted to the very apex, bitter taste of the context, positive reaction of the stipe context with KOH (dark orange) and FeSO 4 (dark green) and medium sized (10.9 - 14.4 x 3.9 - 4.9 µm) basidiospores.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
D86686AF155DD98871E6163DE3E98349.taxon	etymology	Etymology. Referring to Indian Himalaya, the type locality.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
D86686AF155DD98871E6163DE3E98349.taxon	description	Description. Pileus 71 - 120 mm diam., initially convex then plano-convex to applanate, surface dry, matte to somewhat subvelvety, faintly areolate, brownish-grey, dull red, reddish-grey to purplish-grey or greyish-magenta (11 - 13 B-C 2 - 3) when young, gradually brown to reddish-brown (7 E 4 - 9 D 4) or darker, paler greyish-yellow (4 C 4) towards margin, pale yellow (2 A 3) at margin; margin entire, decurved to plane with a narrow flap of tissue, blond (4 C 4). Pore surface greyish-yellow (3 C 4) when young, pinkish-white (8 A 2) with age, turning greyish-brown (6 D 3) on bruising; pores angular, 2 / mm. Tubes adnexed to subdecurrent, 5 - 6 mm long, whitish-brown to brownish, light brown to brown (26 B 2 - 3) on bruising. Stipe 95 - 155 x 20 - 32 mm, mostly subclavate, hollow, pale yellow (1 - 2 A 3) at apex, brownish towards base but never violaceous; surface usually without any reticulum, but sometimes faintly reticulate at apex (1 / 8 th from the juncture), the rest longitudinally striate. Context up to 16 mm thick in pileus, milk white (1 A 2), unchanging when exposed. Stipe context turning dark green with FeSO 4, dark orange with 5 % KOH, orange with 10 % NH 4 OH. Taste bitter. Spore print not obtained. Basidiospores 10.9 - 12.5 - 14.4 x 3.9 - 4.5 - 4.9 µm, (n = 30; Q = 2.51 - 2.75 - 3.25), elongated to fusiform, inequilateral, thin-walled, smooth under light microscope. Basidia 30 - 40 x 9 - 10 µm, four-spored, clavate. Pleurocystidia 27 - 54 x 8 - 10.5 µm, emergent up to 30 µm, fusoid to ventricose, appendiculate. Tube edge sterile; cheilocystidia 32 - 48 x 5.2 - 8 µm, common, clavate to cylindrical, subfusoid to ventricose. Hymenophoral tra ma divergent, hyphae septate, gelatinous, up to 6 µm wide. Pileipellis a trichoderm, up to 150 µm thick, composed of erect hyphae, somewhat interwoven, encrusted, brown pigmented; terminal elements 20 - 50 x 5 - 10 µm, cylindrical to subcylindrical, sometimes subfusoid, content brown pigmented. Stipitipellis a cutis, made up of sub-parallel repent hyphae; caulocystidia not observed; caulobasidia similar to that of hymenial basidia.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
D86686AF155DD98871E6163DE3E98349.taxon	materials_examined	Specimens examined. India. Sikkim: Upper Chandmari, 1977 m alt., N 27 ° 23 ' 13.7 ", E 88 ° 46 ' 42.9 ", under Pinus sp. in temperate mixed forest, 26 Aug 2017, D. Chakraborty & K. Das, DC 17 - 31 (CAL); Uttarakhand: Champawat district, Abbot mount, 1933 m alt., N 29 ° 25.465 ', E 80 ° 06.422 ', under Cedrus deodara in temperate coniferous forest, 18 Jul 2017, A. Ghosh, KD B- 03 (CAL).	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2C7C7010F4EF5F893205097FBC105C20.taxon	materials_examined	Type. INDIA. Sikkim, South District, Maenam WLS (Maenum 3), 2136 m alt., N 27 ° 15 ' 34.7 " E 88 ° 21 ' 25.7 ", 23 Aug 2017, Quercus spp., D. Chakraborty & K. Das, DC 17 - 30 (CAL 1651)	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2C7C7010F4EF5F893205097FBC105C20.taxon	diagnosis	Diagnosis. Distinct from all allied taxa by sequence data (nrITS and nrLSU) and morphologically by its sticky orange-yellow pileus surface, pale yellow pore surface which turns to pale orange or greyish-orange when bruised and absence of reticulation on stipe surface.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2C7C7010F4EF5F893205097FBC105C20.taxon	etymology	Etymology. referring to its being a look-alike of T. balloui, a North American species.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2C7C7010F4EF5F893205097FBC105C20.taxon	description	Description. Pileus 60 - 150 mm diam., initially convex then plano-convex, surface sticky, orange-yellow to brownish-yellow (5 B-C 8), paler at margin; margin entire, plain, without any sterile flap of tissue. Pore surface pale yellow (3 A 3), turning pale orange to greyish-orange (6 A-B 3) on bruising; pores angular, 5 - 8 / mm. Tubes subdecurrent, 6 - 10 mm long, yellowish-white, brownish on bruising. Stipe 55 - 110 x 20 - 40 mm, mostly subclavate, solid, concolorous with pileus; surface pruinose, never reticulate; basal mycelium white. Context 20 mm thick in pileus, chalky white (1 A 1), unchanging on exposure but turning turquoise grey (24 D 2 - D 1) with FeSO 4 (chalk), pale yellow (4 A 3) with 5 % KOH, yellowish-grey (4 B 3) in Guaiacol. Pileus surface brownish-red (8 C 8 - 7) on bruising, turning dark green to greenish-grey (25 E-F 3 - 2) in FeSO 4, vivid yellow (3 A 8) in KOH, unchanging in NH 4 OH. Stipe 55 - 110 x 20 - 40 mm, mostly subclavate, solid, concolorous with pileus; surface pruinose, never reticulate; basal mycelium white. Odour pleasant. Taste slightly pungent. Spore print not obtained. Basidiospores 6.4 - 7.4 - 9.9 x 3.8 - 4.5 - 5.7 µm (n = 30, Q = 1.5 - 1.73 - 2.04), ellipsoidal, thin-walled, smooth under light microscope. Basidia 22 - 30 x 8 - 10 µm, four-spored, clavate. Pleurocystidia 40 - 54 x 7 - 10 µm, emergent up to 30 µm, fusoid to ventricose, appendiculate, yellow pigmented or hyaline, mostly with dense globular to oily content. Tube edge fertile; cheilocystidia 33 - 55 x 7 - 10 µm, common, clavate to cylindrical, subfusoid to ventricose. Hymenophoral trama divergent, hyphae septate, gelatinous, up to 5 µm wide. Pileipellis an ixocutis up to 150 - 280 µm thick, composed of subparallel to suberect, somewhat interwoven hyphae; terminal elements 20 - 70 x 6 - 11 µm, cylindrical to subcylindrical, sometimes subfusoid, content orange-brown pigmented. Stipitipellis up to 150 µm thick, fertile, composed of basidia and cystidia in several clusters; caulobasidia similar to that of hymenial basidia; caulocystidia 40 - 76 x 10 - 12 µm, broadly clavate to subclavate or appendiculate.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2C7C7010F4EF5F893205097FBC105C20.taxon	materials_examined	Specimens examined. India. Sikkim: South District, Maenam WLS (Maenam 3), 2136 m alt., N 27 ° 15 ' 34.7 ", E 88 ° 21 ' 25.7 ", 23 Aug 2017, Quercus sp., D. Chakraborty & K. Das, DC 17 - 35 (CAL); Uttarakhand: Champawat district, Abbot mount, 1885 m alt., N 29 ° 25.466 ', E 80 ° 06.085 ', 18 July 2017, A. Ghosh, KD B- 02 (CAL). Uttarakhand: Pauri district, 1971 m alt., N 30 ° 02.874 ', E 79 ° 08.221 ', 4 Aug 2017, K. Das, KD 17 - 24 (CAL).	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2FE67B37D51B2E1CFF039B67E5019455.taxon	description	Description. Pileus 70 - 120 mm broad, convex to broadly convex; surface distinctly tomentose to subvelvety, dry, subviscid when wet; reddish-brown (8 - 9 F 4) when young, rosewood (9 C 5) to vinaceous-brown (16 C 5) with maturity, fawn (7 E 4) towards margin. Pores yellowish-white or cream with a pinkish tinge, orange-grey (6 B 2) with age; pore 2 - 3 / mm, rounded. Tubes 10 - 15 mm, adnate-sinuate, white in colour, yellow-brown or orange white with maturity. Stipe 60 - 100 x 18 - 22 mm, cylindrical, solid, surface dry, glabrous to subvelutinous, typically distinctly reticulate at apex, reticulation greyish-ruby (12 C-D 4) to dark ruby (12 F 5); surface pinkish brown to vinaceous or violaceous, dark brown to reddish-brown with maturity. Context chalky white, but pinkish-brown when exposed. Spore print not obtained. Basidiospores 10 - 11.5 - 13.5 x 4 - 4.6 - 5.2 µm (n = 30, Q = 2.05 - 2.48 - 2.76), ellipsoid to narrowly subfusoid, inequilateral, smooth, thin-walled. Basidia 30 - 36 x 10 - 11 µm, 4 - spored, clavate, thin-walled, hyaline or pale grey in KOH. Pleurocystidia 35 - 66 x 14 - 24 µm, scattering and numerous, fusoid-ventricose or subclavate, with orange brown contents. Cheilocystidia 33 - 38 x 9 - 12 µm, ventricose to fusoid, shorter and smaller than pleurocystidia thin-walled, with orange brown contents. Pileipellis 100 - 150 µm thick, an ixotrichoderm of suberect, branched, septate hyphae; terminal elements ventricose to fusoid, vaculolated, 28 - 50 x 12 - 14 µm, with granular yellowish to brown orange contents in KOH; subterminal elements mostly with incrustations. Stipitipellis 35 - 65 µm, fertile, composed of basidia and cystidia. Caulocystidia 52 - 63 x 8 - 13 µm, fusoid to subfusoid, ventricose to ventricose-rostrate or narrowly cylindrical, content granular. Clamp connection absent in all tissues.	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2FE67B37D51B2E1CFF039B67E5019455.taxon	distribution	Known distribution. Japan (Kawamura 1954; Hongo 1960; Imazeki et al. 1970, 1988; Takahashi 1986; Gelardi et al. 2015), China (Ying and Zang 1994; Li and Song 2000; Wang et al. 2004; Fu et al. 2006; Wu et al. 2011; Gelardi et al. 2015), Russia (Vasil'jeva 1978) and New Guinea (Hongo 1973).	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
2FE67B37D51B2E1CFF039B67E5019455.taxon	materials_examined	Specimens examined. India. Sikkim: East district, Fambonglo WLS, 2021 m alt., N 27 ° 21 ' 47.5 " E 88 ° 34 ' 13.2 ", 26 Aug 2016, D. Chakraborty & K. Das, DC 16 - 63 (CAL); ibid., D. Chakraborty & K. Das, DC 16 - 64 (CAL).	en	Chakraborty, Dyutiparna, Vizzini, Alfredo, Das, Kanad (2018): Two new species and one new record of the genus Tylopilus (Boletaceae) from Indian Himalaya with morphological details and phylogenetic estimations. MycoKeys 33: 103-124, DOI: http://dx.doi.org/10.3897/mycokeys.33.23703, URL: http://dx.doi.org/10.3897/mycokeys.33.23703
