identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AF8798FFC6FF99FF6EFD26FD0216A3.text	03AF8798FFC6FF99FF6EFD26FD0216A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius billitonensis Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius billitonensis Valeton (1909a: 59)</p>
            <p>Type: — INDONESIA. Bilitoeng, Mangar, sin. dat. (♂), Teijsmann s.n. (lectotype BO [No numerical identifiers found on the sheet], designated here).</p>
            <p> Notes: — Three syntypes are indicated in the protologue: Teijsmann s.n. and Ham s.n. from Billiton Island, and Hallier 2444 from Mount Kelam in West Kalimantan (“Borneo Centralis in monte Klam” fide Valeton (1909a)). Unlike the Billiton collections, the calyx morphology of Hallier 2444 from Borneo is inconsistent with the protologue—the length of the calyx and its lobes are much shorter than that stated in the protologue. Moreover, it is different from the Billiton collections by its subglabrous calyx and non-glaucous, glabrous lower leaf surfaces, and is here excluded from the circumscription of  T. billitonensis . Only the staminate inflorescence was described in Valeton’s protologue, with a note indicating that the pistillate counterpart was unknown. We identified four relevant sheets from Billiton Island in BO (where Valeton worked from 1892 to 1913): a fruiting collection with both “ Ham 19 ” and “ Burger 15 ” on its label, a sterile sheet labelled only as “ Ham 19 ”, a sterile sheet with both “ Ham 19 ” and “ Teysm. ” written on the label, and a collection by Teijsmann (without collection number) that represents the only staminate specimen. Thus,  T. billitonensis is here lectotypified by the Teijsmann collection. </p>
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	https://treatment.plazi.org/id/03AF8798FFC6FF99FF6EFD26FD0216A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC6FF99FF6EFB43FCCC1477.text	03AF8798FFC6FF99FF6EFB43FCCC1477.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius clementis Merrill 1917	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius clementis Merrill (1917b: 244)</p>
            <p>Type: —BORNEO. Malaysia, Sabah, Mount Kinabalu, Marai Parai Spur, 2 Dec 1915 (♀), Clemens 11078 (lectotype A [A00312911], designated here).</p>
            <p>Notes: —Only a single specimen of the original material is extant in A; the duplicate in PNH must have been destroyed during World War II. Merrill (1917a) mentioned that the type material of his Bornean species were deposited in PNH. Although Chen et al. (2015) correctly stated that the holotype in PNH was destroyed, a lectotypification was not performed. Therefore, we here designate the A sheet as lectotype.</p>
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	https://treatment.plazi.org/id/03AF8798FFC6FF99FF6EFB43FCCC1477	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC6FF9AFF6EF9F7FE88134D.text	03AF8798FFC6FF9AFF6EF9F7FE88134D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius esherianus W. W. Smith 1915	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius esherianus W.W. Smith (1915: 327)</p>
            <p> Type: —   BORNEO. Malaysia, Sarawak,  Mount Santubong , 14 Mar 1914 (♀), Native Collector D 200 (lectotype E [E00143855], designated here; isolectotypes E [E00438196], K [K000763577])  . </p>
            <p>Notes: —Smith cited a single collection (Native Collector D200) in his protologue and stated that “there is in the type specimen only one corolla, which I have not dissected”. There is indeed an E sheet [E00143855] with a single corolla in the packet that was annotated “only one flower that [was] not dissected” by Smith, but a K duplicate [K000763577] also has a single intact corolla. Another E duplicate [E00438196] is a sterile specimen with Smith’s annotation: “further material of this new sp. is much needed, flowers and ripe fruits especially”. We assume that the two sheets at E correspond to two separate specimens rather than a two-part specimen as there is no indication of “sheet1” and “sheet2” on them. Since Smith worked in E since 1911, the material annotated by him are likely to be his types. However, it is possible that Smith examined the K sheet prior to distribution and/or used multiple sheets when preparing his description; therefore, we here designate E00143855 as lectotype.</p>
            <p> The inclusion of  T. esherianus in the flora of Mount Kinabalu (Beaman &amp; Anderson 2004) is based on the misidentification of a single cited specimen that corresponds to  T. pannosus (Chen et al. 2015) . The specimens recognised as  T. esherianus in Coode et al. (1996) are also misidentified and actually correspond to  T. hirsutus Merrill (1917b: 242) . </p>
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	https://treatment.plazi.org/id/03AF8798FFC6FF9AFF6EF9F7FE88134D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC5FF9BFF6EFEE0FC74134C.text	03AF8798FFC5FF9BFF6EFEE0FC74134C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius flavescens (Jack 1823) Baker 1877	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius flavescens (Jack 1823: 127) Baker (1877: 144)</p>
            <p> Basionym:  Helospora flavescens Jack</p>
            <p>Lectotype: —[Published illustration] (♀) Jack (1823: 130, t. 4, fig. 3), designated in Wong et al. (2019).</p>
            <p>Epitype: — INDONESIA. Sumatra, Palembang, Ogan Oeloe, sin. dat. (♀), Teijsmann HB3877 (epitype U [U.1577430], designated in Wong et al. (2019)).</p>
            <p> =  Guettarda brunonis Wall. ex G. Don (1834: 552) ,  syn. nov.</p>
            <p>Type:— SINGAPORE. Sin. loc., 1822 (♀), Wallich s.n. [EIC 6220] (holotype K-W [K001123335]).</p>
            <p> =  Bobea sericantha f. glabrior Miquel (1861: 545)</p>
            <p>Type:— INDONESIA. Sumatra, Palembang, Ogan Oeloe, sin. dat. (♀), Teijsmann HB3877 (holotype U [U.1577430]).</p>
            <p> =  Timonius flavescens var. brevipetiolata Valeton (1909a: 35)</p>
            <p>Type:—BORNEO. Bukit Singkadjang, 1905 (♀), Teijsmann 8348 (lectotype BO [acc 1435253], designated here; isolectotypes BO [acc 1570710; acc 1570711; acc 1570712]).</p>
            <p> =  Timonius peduncularis (Wall. ex G. Don 1834: 551) Ridley (1923: 113)</p>
            <p> Basionym:  Guettarda peduncularis Wall. ex G.Don</p>
            <p>  Type :— SINGAPORE. Sin. loc., 1822 (♀), Wallich s.n. [EIC 6222] (lectotype K-W [K000763583], first step designated in Wong et al. (2019), second step designated here to include only four detached leaves (see notes); isolectotype K-W [K001123337], explicitly excluding the left element which represents another gathering)  . </p>
            <p> Notes: —Although originally described from Sumatra,  T. flavescens has been recorded in Borneo (Merrill 1921a, Masamune 1942, Sidiyasa et al. 1999). Jack cited no specimen for  Helospora flavescens , merely stating “Native of Sumatra”, so it has to be assumed that his taxon is based on a collection by himself from Sumatra, which has not been found. Baker’s description of the plant in the Seychelles was brief, but had effectively recombined Jack’s name because he cited Jack’s binomial under his  T. flavescens . Baker cited collections by Pervillé, Wright and Horne from the Seychelles, which have been subsequently recognised as a separate species,  T. sechellensis Summerhayes (1928: 391) ; it is different from  T. flavescens because of its much shorter (less than 1 cm long) pistillate and fruiting peduncles. Teijsmann HB3877 is one of the specimens identified as  T. flavescens in an early taxonomic treatment of  Timonius (Valeton 1909a) ; because this specimen is from Sumatra, it was selected as the lectotype of  T. flavescens by Wong (1988). Darwin (1997) designated the same Sumatran specimen as the neotype of  T. flavescens , correcting Wong’s earlier use of the term lectotype, given that this specimen is not part of the original material cited in Jack’s protologue. Both authors overlooked the presence of a published illustration in the protologue, however. Wong et al. (2019) therefore selected the published illustration as lectotype and designated the Sumatran specimen as epitype because the illustration is inadequate for identification purposes. </p>
            <p> Timonius flavescens var. brevipetiolata Valeton was described based on two of Teijsmann’s collections, but only one of them (Teijsmann 8348) can be located and has a collection number. One of the duplicates in BO [acc 1435253] is the best preserved and is therefore here designated as lectotype.  Timonius flavescens var. grandiflora Valeton (1909a: 35) is excluded from the definition of this species here. </p>
            <p> The type material of  T. peduncularis (Wallich s.n. [EIC 6222]) is represented by two sheets in K-W, both with mixed elements of  T. finlaysonianus (Wall. ex G. Don 1834: 552) Hook.f. (1880: 127) and  T. flavescens . The sheet [K000763583] designated as lectotype by Wong et al. (2019) comprises largely elements of  T. finlaysonianus , with only four detached leaves of  T. flavescens (marked with ‘X’ in pencil) mounted together with an intact twig of  T. finlaysonianus . The other sheet [K001123337] that is not cited in Wong et al. (2019) has three separate branches: two intact twigs of  T. flavescens with several attached fruits (middle and right) and one intact twig of  T. finlaysonianus (left). According to Art. 9.14 of the International Code of Nomenclature for algae, fungi, and plants (Turland et al. 2018), when a type consists of elements belonging to more than one taxon, “the name must remain attached to the part that corresponds most nearly with the original description or diagnosis”. Therefore, we here designate K000763583 (explicitly including only four detached leaves of  T. flavescens ) as second-step lectotype. We also here designate K001123337 (excluding the left element) as isolectotype of  T. peduncularis . </p>
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	https://treatment.plazi.org/id/03AF8798FFC5FF9BFF6EFEE0FC74134C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC4FF9BFF6EFEE0FD3E11E1.text	03AF8798FFC4FF9BFF6EFEE0FD3E11E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius hirsutus (Elmer) Merrill 1917	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius hirsutus Merrill (1917b: 242)</p>
            <p> Type: —   BORNEO. Malaysia, Sarawak,  Baram , Miri, 20 Apr 1895 (♂), Hose 651 (lectotype A [A00312908], designated here; isolectotype L [L0001476])  . </p>
            <p>Notes: —Although Merrill mentioned a single collection (Hose 651) in his protologue, a few duplicates of this collection exist, and apparently there was also a PNH specimen that must have been destroyed during the war. The holotype of Merrill’s Bornean species was deposited in PNH (Merrill 1917a). Chen et al. (2015) stated that the holotype in PNH was destroyed, but did not select a lectotype among the extant type material. Therefore, we here designate a well-preserved A sheet as lectotype.</p>
            <p> The name “  Timonius hirsutus ” has been used three times. The name was first used by Merrill (1917b) for the Bornean species above. The second name  Timonius hirsutus Ridley (1918: 81) postdates the Bornean name and is a synonym of the Malayan species  Antirhea atropurpurea (Craib 1932: 132) Chaw in Wong (1988: 493). Subsequently, Merrill (1921b: 480) transferred  Greenea hirsuta Elmer (1913: 1857) into  Timonius , generating the illegitimate combination  Timonius hirsutus (Elmer) Merr. for a species from the Philippines. Therefore,  Timonius hirsutus Merr. as used for the Bornean species takes precedence. </p>
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	https://treatment.plazi.org/id/03AF8798FFC4FF9BFF6EFEE0FD3E11E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC4FF9BFF6EFC04FEBA1735.text	03AF8798FFC4FF9BFF6EFC04FEBA1735.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius involucratus Merrill 1917	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius involucratus Merrill (1917b: 241)</p>
            <p> Type: —  BORNEO. Sarawak, sin. dat. (♀), Native Collector 1252 (lectotype K [K000763574], designated here; isolectotypes A [A00312909], US [US 00138422]) . </p>
            <p>Notes: —Merrill specified that the original material was deposited in “Bur. Sci.”, referring to the PNH herbarium that was destroyed during World War II. Although A was actually Merrill’s main herbarium apart from PNH, the best-preserved specimen is from K, with intact fruits and associated calyx lobes and peduncular bracts. We therefore designate the well-preserved K specimen as lectotype.</p>
            <p> Timonius involucratus Valeton (1927: 41) is an illegitimate name as it postdates the Bornean name above. Valeton’s later homonym refers to a New Guinea species and has been appropriately replaced by  Timonius papuanus Merrill (1937: 132) . </p>
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	https://treatment.plazi.org/id/03AF8798FFC4FF9BFF6EFC04FEBA1735	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC4FF9BFF6EFA38FB3D14C9.text	03AF8798FFC4FF9BFF6EFA38FB3D14C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius lasianthoides Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius lasianthoides Valeton (1909a: 48)</p>
            <p>Type: —CENTRAL BORNEO. Jelaeh Kedanim, Jul 1896 (♀), Jaheri (Exp Nieuwenhuis) 172 (lectotype BO [acc 1437765], designated here; isolectotype BO [acc 1437766]).</p>
            <p>Notes: —Of the four collections cited in the protologue (Nieuwenhuis 172, de Vriese s.n., Hallier s.n. and Teijsmann s.n.), only “ Nieuwenhuis 172 ” has a collection number indicated. One of the duplicates in BO (bearing the labels “leg. Jaheri” and “Exp. Nieuwenhuis”) is the best preserved and is therefore selected as lectotype.</p>
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	https://treatment.plazi.org/id/03AF8798FFC4FF9BFF6EFA38FB3D14C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC4FF9BFF6EF97CFF691A61.text	03AF8798FFC4FF9BFF6EF97CFF691A61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius matangensis Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius matangensis Valeton (1909a: 58)</p>
            <p>Type: —BORNEO. Malaysia, Sarawak, Matang, 1907 (♂), Hewitt Rb 46.2 (holotype BO [acc 1435130]).</p>
            <p> Notes: —The name  Timonius matangensis was validly published but the only specimen cited in the protologue (Hewitt Rb 56.2) is untraceable. However, there is a specimen in BO with a very close match to the protologue (Hewitt Rb 46.2): it was collected in the same locality (Matang), possesses only staminate flowers (consistent with the protologue) and was determined by Valeton himself in 1908. Thus, the original type citation seems to be erroneous and is corrected here. </p>
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	https://treatment.plazi.org/id/03AF8798FFC4FF9BFF6EF97CFF691A61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC3FF9CFF6EFF2CFC1A13DD.text	03AF8798FFC3FF9CFF6EFF2CFC1A13DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius ovalis (Korthals 1851) Boerlage 1891	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius ovalis (Korthals 1851: 216) Boerlage (1891: 132)</p>
            <p> Basionym:  Polyphragmon ovale Korth.</p>
            <p>Type: —BORNEO. Sin. loc., sin. dat. (♂), Korthals s.n. (lectotype L [L0001448], designated here; probable isolectotype L [L0057539]).</p>
            <p>Notes: —Although Korthals (1851) cited only a single locality (‘montem Pamatton’) in the protologue, there are two L sheets from Borneo without locality details labelled as types. Consistent with the protologue, both specimens possess staminate flowers. Hence, one of the sheets is here selected as lectotype.</p>
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	https://treatment.plazi.org/id/03AF8798FFC3FF9CFF6EFF2CFC1A13DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC3FF9CFF6EFE50FBAB1751.text	03AF8798FFC3FF9CFF6EFE50FBAB1751.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius palawanensis Elmer (1912: 1360	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius palawanensis Elmer (1912: 1360)</p>
            <p> Type: —   PHILIPPINES. Palawan, Brooks Point,  Addison Peak , Feb 1911 (♂), Elmer 12705 (lectotype A [A00094665], designated here; isolectotypes, BISH [BISH1004781], BO, E [E00504406], F, GH [GH00094664], HBG [HBG520961], K [K000763549], L [L0001450], NY [NY00133423], U [U0227797], US [US 00138428])  . </p>
            <p> =  Timonius villamilii Merrill (1917b: 240)</p>
            <p>  Type:—BORNEO. British North Borneo, “ Nov 1916 and Feb 1917 ” (♀),  Villamil 252 bis (lectotype K [K000763562], designated here; isolectotypes A [A00312915], US [US 00138439])  . </p>
            <p> =  Timonius heterophyllus Merrill (1917b: 243)</p>
            <p>Type:—BORNEO. Malaysia, Sabah, Khota Balud to Kibaya, trail to Mount Kinabalu, 28 Oct 1915 (♂), Clemens 9811 (lectotype A [A00312907], designated here).</p>
            <p> Notes: —Although a single collection (Elmer 12705) is cited as the type material of  T. palawanensis , many duplicates in various herbaria exist and a specimen in PNH must have been destroyed during the war. Because Elmer has worked in A (apart from PNH) and the A specimen has a well-preserved corolla, the A sheet is here designated as lectotype. </p>
            <p> For  T. villamilii, Merrill cited the collection “ Villamil 252 bis ” as type material, which comprises three extant duplicates in various herbaria and a specimen in PNH that must have been destroyed during World War II. Merrill (1917a) stated that the type material of his Bornean species was preserved in PNH. The holotype is thus destroyed, and we here designate the K sheet with well-preserved corolla and fruit as lectotype. </p>
            <p> For  T. heterophyllus , only a single specimen of the original material is extant in A; a duplicate in PNH must have been destroyed during the war. Chen et al. (2015) regarded the destroyed PNH specimen as holotype and did not perform any lectotypification. We therefore here select the A sheet as lectotype. </p>
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	https://treatment.plazi.org/id/03AF8798FFC3FF9CFF6EFE50FBAB1751	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC3FF9CFF6EFAD4FBD61431.text	03AF8798FFC3FF9CFF6EFAD4FBD61431.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius polyneurus Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius polyneurus Valeton (1909a: 59)</p>
            <p> Type: —   BORNEO. West Kalimantan, Sintang, sin. dat. (sterile),  Teijsmann 8361 (lectotype L [L0001451], designated here; isolectotype K [K000763570])  . </p>
            <p>Notes: —Although a single collection (Teijsmann 8361) is cited in the protologue, two duplicates of the original material exist. We therefore here designate a well-preserved specimen from L as lectotype.</p>
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	https://treatment.plazi.org/id/03AF8798FFC3FF9CFF6EFAD4FBD61431	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC3FF9CFF6EF934FDFC158D.text	03AF8798FFC3FF9CFF6EF934FDFC158D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius salicifolius Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius salicifolius Valeton (1909b: 273 , t. 298) </p>
            <p>Type: —BORNEO. Poeloe Madjang, 1905 (♀), Teijsmann 8330 (lectotype L [L0057844], designated here; isolectotypes BO [acc 1438665; acc 1438666; acc 1438667; acc 1438668], L [L0057845]).</p>
            <p>Notes: —Of the six collections cited in Valeton’s protologue (Teijsmann 8326, 8327, 8330; Hallier 2482, 1300, 2200), only three can be located in BO and L. The specimen Teijsmann 8330 [L0057844] is fertile, well-preserved and bears the manuscript label “Poeloe Madjang”; it is therefore here selected as lectotype.</p>
            <p> The two specimens recognised as  T. salicifolius in Coode et al. (1996: 293) were misidentified; we have not encountered any specimen of  T. salicifolius in Brunei. </p>
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	https://treatment.plazi.org/id/03AF8798FFC3FF9CFF6EF934FDFC158D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
03AF8798FFC2FF9DFF6EFF2CFD011174.text	03AF8798FFC2FF9DFF6EFF2CFD011174.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Timonius wallichianus (Korth.) Valeton 1909	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Timonius wallichianus Valeton (1909a: 36)</p>
            <p>Type: — SINGAPORE. Sin. loc., 1822 (♀ and ♂), Wallich s.n. [EIC 6217] (lectotype K-W [K001123325], first step designated in Wong (1988), second step designated in Wong et al. (2019); isolectotypes BR [BR0000005754595], K-W [K001123326], PH [PH 00023297]).</p>
            <p> Notes: —Although  T. wallichianus was excluded in early enumerations of the Bornean flora (Merrill 1921a, Masamune 1942), it has been more recently recorded in East Kalimantan (Sidiyasa et al. 1999). The name “  Bobea wallichiana ” was invalidly published by Korthals (1851: 211), lacking any description or diagnosis. Therefore, Valeton’s name was incorrectly stated in protologue as a combination “  Timonius wallichianus (Korth.) Valeton ”. Valeton (1909a) proceeded to equate  T. wallichianus with  T. rumphii sensu Hooker (1880: 127) non Candolle (1830: 461), a misapplied name that was based on the specimen Wallich s.n. [EIC 6217]. The Wallich specimen was cited in Valeton’s protologue and indeed represents  T. wallichianus , and hence Wong (1988) cited it as the holotype of  T. wallichianus . However, a Malacca collection by Ridley and collections from Banka by Teijsmann and Kobus were also cited in Valeton’s protologue. The original material of  T. wallichianus therefore comprises several elements, and the use of the term holotype in Wong (1988) is a correctable error that should be corrected to lectotype. Although Wong (1988) is considered to have effectively lectotypified this name, a second-step lectotypification was performed in Wong et al. (2019) as there are two specimens of the Wallich collection in K-W. </p>
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	https://treatment.plazi.org/id/03AF8798FFC2FF9DFF6EFF2CFD011174	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chen, Junhao;Wong, Khoon Meng	Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
