taxonID	type	description	language	source
03AE3108F800FFB0FF74FEC708DF942A.taxon	type_taxon	Type genus. Burmoptarthrus gen. nov.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FEC708DF942A.taxon	description	Description. Body large (6.0 mm), subflatened dorsally, moderately sclerotized and covered with dense semierect hairs; head capsule strongly constricted behind eyes; rostrum thin and long; labrum subrectangular, transverse, free; mandibles small, not exodontous; place of antennal insertion not extended; antennae not geniculate, inserted laterally in basal part of rostrum; club not compact, its three segments free; eyes elliptical, strongly protruding from head; temples short; gular suture double; pronotum bell-shaped, weakly flattened, coarsely punctate and granulated; lateral margins of pronotum granulated; scutellum large and transverse; elytra quite wide, coarsely punctate and granulated, with greatest width behind middle; weakly punctostriate; striae indistinct; scutellar striole distinct; epipleura wide; precoxal portion of prosternum shorter than procoxal cavity length; postcoxal portion longer than precoxal portion; procoxal cavities located closer to apical margin, rounded and contiguous; mesocoxal cavities closed, rounded and separated; abdomen flattened, with ventrites free; ventrite 1 subequal to ventrite 2; trochanters not separating femora and coxae; femora without teeth; tibiae almost straight, without mucro and with two apical spurs; tarsi pseudoquadrisegmented; tarsomere 1 wide-conical; tarsomeres 2 and 3 conical-bilobed; tarsomere 5 long and narrow; claws free, strongly divergent, without teeth at base.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FEC708DF942A.taxon	diagnosis	Diagnosis. The subfamily Brenthorrhininae consists of three Mesozoic tribes Eccoptarthrini, Brenthorrhinini and Distenorrhinini (Gratshev & Legalov 2014). The new tribe differs from three others in having the head capsule strongly constricted behind the eyes, eyes protruding strongly, the coarsely punctate and granulated pronotum and head, as well as granulated lateral margins of the pronotum. The procoxae are located closer to the apical margin of the prosternum similar to these in the tribe Eccoptarthrini. However, the tribe Burmoptarthrini also differs from the Eccoptarthrini in having small mandibles and bilobed tarsomeres 2 and 3. It can also be distinguished from the tribes Brenthorrhinini and Distenorrhinini by the location of the procoxae and the antennal insertion in the basal part of the rostrum.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FEC708DF942A.taxon	discussion	Remarks. The new tribe has been placed in the family Nemonychidae based on the free labrum, the double gular suture, non-geniculate antennae, free abdominal ventrites and tibiae with spurs. It belongs to the subfamily Brenthorrhininae based on the procoxae being located closer to the apical margin of the prosternum.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FA0B0DF697E9.taxon	type_taxon	Type species. Burmoptarthrus storozhenkoi sp. nov.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FA0B0DF697E9.taxon	etymology	Etymology. The name of the new genus is formed from the old name of Myanmar — “ Burma ” and part of the name “ Eccoptarthrus ”, because of the relative similarity of the new genus to the aforementioned tribe members.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB0FF74FA0B0DF697E9.taxon	diagnosis	Diagnosis. As for tribe.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB7FF74F94B09A49089.taxon	description	(Figs 1 – 4)	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB7FF74F94B09A49089.taxon	materials_examined	Material. Holotype, female, MA 2024 / 1, Burmese amber, Myanmar, Cenomanian, late Cretaceous. Paratype, male, MA 2018 / 18, Burmese amber, Myanmar, Cenomanian, late Cretaceous. The paratype is damaged. Only the front part of its body is well preserved.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB7FF74F94B09A49089.taxon	description	Description. Body black-brown, moderately sclerotized, subflattened dorsally. Integument covered with dense semierect pale hairs. MALE. Head capsule strongly constricted behind eyes. Labrum well developed, separated from rostrum, subrectangular. Maxillary palpi 4 - segmented, flexible. Rostrum elongated, weakly curved, densely punctate, slightly thickened at apex, about 6.4 times as long as wide at apex, about 7.4 times as long as wide at mid length, about 6.7 times as long as wide at base, 1.2 times as long as pronotum. Dorsal surface of rostrum without carinae. Ventral surface of rostrum with two weak carinae. Antennae long, thin, non-geniculate, inserted laterally in basal part of rostrum, with quite long pale setae. Place of antennal insertion not extended. Distance from rostrum apex to antennal insertion 6.7 times as long as distance from rostrum base to antennal insertion. Antennomeres subconical. Antennomere 1, about 2.1 times as long as wide at mid length. Antennomere 2, about 1.3 times as long as wide at apex, about 0.5 times as long as and about 0.8 times as narrow as antennomere 1. Antennomeres 2 - 6 subequal. Antennomere 3, about 2.2 times as long as wide at apex, about 1.4 times as long as and about 0.8 times as narrow as antennomere 2. Antennomere 7, 1.4 times as long as wide at apex, 0.7 times as long as and slightly wider than antennomere 6. Antennomere 8, about 1.5 times as long as wide at apex, about 1.2 times as long as and 1.1 times as wide as antennomere 7. Antennal club not compact, its three segments free. Antennomere 9, about 1.7 times as long as wide at apex, about 1.2 times as long as and about 1.1 times as wide as antennomere 8. Antennomere 10, 1.5 times as long as wide at apex, 1.2 times as long as and about 1.3 times as wide as antennomere 9. Antennomere 11, 2.8 times as long as wide at base, about 1.2 times as long as and about 0.6 times as narrow as antennomere 10. Eyes elliptical, distinctly protruding from head. Temples short, about 0.8 times as long as eye length. Forehead wide, flattened, densely punctate. Pronotum bell-shaped, with arcuate sides. Disc weakly flattened, coarsely punctate and granulated. Greatest width at mid length. Lateral margins of pronotum granulated. Scutellum large, subtrapezoid, about 0.7 times as long as wide. Elytra with slightly smoothed humeri. Scutellar striole distinct. Precoxal portion of prosternum about 0.8 times as long as procoxal cavity length. Postcoxal portion of prosternum about of same length to procoxal cavity length and about 1.2 times as long as precoxal portion. Procoxal cavities located near middle of prosternum, rounded and contiguous. Mesocoxal cavities rounded, narrowly separated. Legs long. Trochanters oblique. Femora quite slender, slightly incrassating subapically, covered with tubercles. Tibiae almost straight, widened at apex, covered with tubercles, without mucro; with two apical spurs. Tarsi wide, pseudoquadrisegmented. Tarsomeres 1 – 3 with pulvilli on lower surface. Tarsomere 1 wide-conical. Tarsomeres 2 and 3 conical-bilobed. Tarsomere 5 long and narrow. Claws free, strongly divergent, without teeth at base. Protarsi: tarsomere 1 about, 0.4 times as long as wide at apex; tarsomere 2, about 0.4 times as long as wide at apex, slightly shorter and narrower than tarsomere 1; tarsomere 3, about 0.9 times as long as wide at apex, 2.6 times as long as and about 1.3 times as wide as tarsomere 2; tarsomere 5, about 3.7 times as long as wide at apex, about 1.7 times as long as and about 0.4 times as narrow as tarsomere 3. Head and pronorum length 2.5 mm; rostrum length 2.0 mm. FEMALE. Labrum about 0.5 times as long as wide. Mandibles small, not exodontous. Rostrum 5.0 times as long as wide at apex, about 8.3 times as long as wide at mid length, about 7.1 times as long as wide at base, 1.2 times as long as pronotum. Antennae extend beyond middle of pronotum, inserted in basal part of rostrum. Distance from rostrum apex to antennal insertion 5.6 times as long as distance from rostrum base to antennal insertion. Antennomere 1, 1.8 times as long as wide at mid length. Antennomere 2, 1.5 times as long as wide at apex, about 0.7 times as long as and 0.8 times as narrow as antennomere 1. Antennomere 3, about 3.3 times as long as wide at apex, about 2.2 times as long as and of same width to antennomere 2. Antennomere 4, about 1.3 times as long as wide at apex, about 0.5 times as long as and slightly wider than antennomere 3. Antennomere 5, about 1.3 times as long as wide at apex, about 0.8 times as long as and slightly narrower than antennomere 4. Antennomere 6, about 1.6 times as long as wide at apex, slightly shorter and wider than antennomere 5. Antennomere 7, about 1.6 times as long as wide at apex, slightly longer and wider than antennomere 6. Antennomere 8, about 1.1 times as long as wide at apex, of same length and about 1.4 times as wide as antennomere 7. Antennal club not compact, its three segments free. Antennomere 9, about 1.3 times as long as wide at apex, about 1.3 times as long as and about 1.1 times as wide as antennomere 8. Antennomere 10, 1.5 times as long as wide at apex, of same length and slightly wider than antennomere 9. Antennomere 11, about 3.1 times as long as wide at base, about 1.6 times as long as and about 0.8 times as narrow as antennomere 10. Eyes distinctly protruding from head. Temples about 0.4 times as long as eye length. Forehead about 1.9 times as wide as rostrum base width, flattened, densely punctate. Pronotum bell-shaped, with arcuate sides, about 2.0 times as long as wide at apex, slightly longer than wide at mid length, about 1.2 times as long as wide at base. Disc weakly flattened, coarsely punctate and granulated. Greatest width at mid length. Lateral margins of pronotum granulated. Scutellum 0.7 times as long as wide. Elytra quite wide, about 1.6 times as long as wide at base and at apical fourth, about 1.3 times as long as wide at mid length, about 2.0 times as long as pronotum; weakly punctostriate, with slightly smoothed humeri and distinct wide epipleura. Greatest width behind middle. Disc coarsely punctate and granulated. Striae indistinct. Scutellar striole distinct. Prosternum, covered with tubercles. Precoxal portion of prosternum about 0.8 times as long as procoxal cavity length. Postcoxal portion of prosternum about of same length to procoxal cavity length and about 1.6 times as long as precoxal portion. Procoxal cavities located near middle of prosternum, rounded and contiguous. Mesocoxal cavities rounded, narrowly separated. Metacoxal cavities transverse. Metaventrite convex, long, about 2.2 times as long as metacoxal cavity. Metanepisterna about 5.7 times as long as wide at mid length, finely punctate. Abdomen weakly convex. Ventrites free. Ventrite 1, about 1.5 times as long as metacoxal cavity. Ventrite 2, slightly shorter than ventrite 1. Femora covered with tubercles, without teeth. Tibiae almost straight, widened at apex, covered with tubercles, without mucro; with two apical spurs. Tarsi wide, pseudoquadrisegmented. Tarsomeres 1 – 3 with pulvilli on lower surface. Tarsomere 1 wide-conical. Tarsomeres 2 and 3 conical- bilobed. Tarsomere 5 long and narrow. Claws free, strongly divergent, without teeth at base. Body length without rostrum 6.0 mm; rostrum length 1.9 mm.	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB7FF74F94B09A49089.taxon	etymology	Etymology. The species is named in honour of Dr. Sergei Yu. Storozhenko (Vladivostok, Russia).	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
03AE3108F800FFB7FF74F94B09A49089.taxon	discussion	Remarks. In females, the rostrum is more strongly expanded at the apex (5.0 times as long as wide at apex) and narrower in the middle (about 8.3 times as long as wide at mid length), whereas in males, it is less expanded at the apex (6.4 times as long as wide at apex) and wider in the middle (about 7.4 times as long as wide at mid length).	en	Legalov, Andrei A. (2025): First record of the subfamily Brenthorrhininae (Coleoptera, Nemonychidae) in Cretaceous Kachin amber. Zootaxa 5715 (1): 225-235, DOI: 10.11646/zootaxa.5715.1.20, URL: https://doi.org/10.11646/zootaxa.5715.1.20
