taxonID	type	description	language	source
03A387C13013870C11F2FC250C8FF881.taxon	diagnosis	Diagnosis Neopachylinae can be differentiated from the other DRMNmembers (Discocyrtus s. s., Mitobatinae, and Roeweriinae) due to: (1) apical portion of the stylus covered by spines; (2) ventral process of the glans with flabellum ‘ hand-shaped’; (3) flabellum armed with spines; (4) Cx IV prodorsal apophysis with the distal portion forming a 90 ° angle in relation to the body; (5) Fe IV dorsal face with one or three spines on the proximal half.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870C11F2FC250C8FF881.taxon	distribution	Distribution ARGENTINA: provinces of Buenos Aires, Córdoba, Ciudad Autónoma de Buenos Aires, Corrientes, Entre Ríos, Jujuy, Misiones, Salta, and Tucumán. BRAZIL: Distrito Federal [Federal District], states of Espírito Santo, Mato Grosso do Sul, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo. URUGUAY: departments of Artigas, Canelones, Colonia, Lavalleja, Paysandú, Río Negro, Salto, and San José.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870B11ABF8930A80FEE5.taxon	description	ZooBank LSID: urn: lsid: zoobank. org: act: 2 A 2 B 15 CD-AD 30 - 43 E 2 - B 64 D- 7 E 70 C 2 F 1 F 25 D.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870B11ABF8930A80FEE5.taxon	type_taxon	Type species: Discocyrtus rectipes Roewer, 1913.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870B11ABF8930A80FEE5.taxon	etymology	Etymology: The genus name is an honor to the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq), which has been financing scientific research for several decades in Brazil. The acronym is based on its former name (CONselho NAcional de PESQUIsas) with a Latin suffix. Gender masculine.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870B11ABF8930A80FEE5.taxon	diagnosis	Diagnosis: Conapesquius can be distinguished from the other genera of Neopachylinae due to (1) Mesotergum with areolate spots in a ‘ diamond’ pattern, absent on the lateral margins of the scutal areas I – II and IV (Fig. 7 A, D); (2) Scutal areas I (posterior margin) and II (anterior margin) parallel each other (except in C. heteracanthus; Figs 5 A, 14 C); (3) Scutal area II central portion with a transverse row of prominent tubercles (Fig. 7 A, D); (4) Fe III with a well-developed apical spur on retrodorsal face (with more than a half of the Cx II size) (Fig. 16 A); (5) Cx IV prodorsal distal apophysis ‘ scythe-shaped’ (except in C. spinifemur) (Figs 5 A 10 K, 14 A); (6) MS B 1 evident, with the central portion visible (Figs 6 B, 11 C); (7) Females’ scutal area III with a paramedian dorsal elevation (Figs 9 E, 13 E).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13013870B11ABF8930A80FEE5.taxon	distribution	Distribution: BRAZIL: states of Paraná and Santa Catarina (Fig. 3).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	description	(Figs 4 – 6)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	materials_examined	Type data Discocyrtus brevifemur: ♂ lectotype (MHNCI 3620!, examined), ♀ paralectotype (MHNCI 3621!, examined), from BRAZIL, Paraná, Piraquara, Banhado. Records Without further literature records.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	diagnosis	Diagnosis Conapesquius brevifemur can be distinguished from C. heteracanthus, C. rectipes, and C. spinifemur due to (1) Ch bulla anterior mesal margin with spines (as in C. rectipes) (Fig. 5 A); (2) mesotergum thickest at area II (Figs 4 E, 5 B); (3) Scutal area III with a paramedian pair of prominent domed-shaped tubercles (as in C. rectipes) (Figs 4 A, C – E, 5 A – B, D); (4) Fe IV dorsal face with two spines on the distal third (as in C. heteracanthus) (Figs 4 A, 5 A); (5) females’ ocularium armed with a pair of parallel spines (Figs 4 C, 5 C); (6) females’ Tr IV retrolateral face with a central apophysis (as in C. heteracanthus).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	materials_examined	Non-type material examined BRAZIL: state of Paraná: Piraquara: 1 ♂ (MNRJ 18926)!, Banhado, [- 25 458 °, - 48 990 °], vii. 1947, Gofferjé, C. N. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	distribution	Distribution BRAZIL: state of Paraná: Piraquara (Fig. 3).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130148707111BFEB6082DFA06.taxon	description	Redescription MNRJ 18926! (male) for the external body illustrations and description (Figs 4 – 5); DS, measurements: CW 2.8, CL 1.9, AW 5.1, AL 2.8; Leg I – IV measurements in Table 6; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 8 (3) / 9 (3) - 7 / 7 - 7 / 7. MNRJ 18926! for genitalic illustrations (Fig. 6). Dorsum: DS gamma-pyriform, as long as wide, with AS lateral margins strongly convex, widest at scutal area III and thickest at scutal area II, with sinuous posterior margin (Figs 4 A, E, 5 A – B). DS anterior margin divided by a small central projection in the center and a pair of shallow cheliceral sockets (Fig. 5 A). Carapace with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Figs 5 A – B). Ocularium elliptical (in dorsal view), high (c. 3 × the eye diameter), perpendicularly placed in the middle of the carapace (Figs 4 A, C, E, 5 A – C). Ocularium with a pair of parallel spines (c. 2 × the eye diameter), inclined frontwards (Figs 4 A, C, E, 5 A – C). AS lateral borders with two rows of tubercles: one external, composed of four or six prominent tubercles at areas II – IV (Fig. 5 A – B); another internal one with ordinary tubercles from the posterior corner of the carapace to the posterior margin (Fig. 5 A). Mesotergum divided into four clearly defined areas (Figs 4 A, 5 A). All areas tuberculate, with almost all tubercles individually covered / surrounded by light-colored spots (Fig. 5 A – B). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 4 A, 5 A). Scutal area I with three pairs of prominent tubercles (c. 2 × the ordinary tubercles) (Fig. 5 A – B). Scutal area II with ordinary tubercles diffusely distributed on all of its extension (Fig. 5 A – B). Scutal area II posterior-lateral margin embracing the scutal area III (Fig. 5 A). Scutal area III with a pair of paramedian prominent domed-shaped tubercles (c. 7 × the ordinary tubercles) (Figs 4 A, D – E, 5 A – B, D). Scutal area IV with two transversal rows of ordinary tubercles (Fig. 5 A – B). DS posterior margin and free tergites I – III each with a transversal row of prominent tubercles interpolated by unarmed spaces (Figs 4 D, 5 A). Anal operculum tuberculate (Fig. 4 D). Venter: Cx I – III sub-parallel to each other, each with ventral longitudinal rows of 7 – 12 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (Fig. 4 B). Cx II with a retroventral distal row of four acuminated tubercles (Fig. 4 B). Cx III with a retroventral distal row of 10 acuminated tubercles (Fig. 4 B). Cx IV much larger than the others, directed obliquely (Fig. 4 B). Intercoxal bridges are well marked (Fig. 4 B). Stigmatic area Y-inverted-shaped, clearly sunken concerning the Cx IV’s distal part (Fig. 4 B). Cx IV covered by ordinary tubercles (Fig. 4 B). Cx IV posterior border and stigmatic area each with a transversal row of ordinary tubercles. Stigmata are visible (Fig. 4 B). Free sternites with a transverse row of ordinary tubercles. Chelicera: Basichelicerite elongate, bulla well-marked (Fig. 4 A), with marginal setiferous tubercles — one anterior mesal, two lateral ectal; hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles (Fig. 4 B). Fe with a ventral basal and a mesal distal setiferous tubercle (Fig. 4 B – C). Pa unarmed. Ti with two rows (ventro-mesal and ventro-ectal) of four spines (IiII) (Fig. 4 C). Ta with two rows of three spines — ventro-mesal, (IIi) ventro-ectal. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles (Fig. 4 B). Fe I – II straight; Fe III sinuous (Fig. 4 A – B). Fe and Ti I – III with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles (Fe III proventral and retroventral tubercles larger and sharper than others) (Fig. 4 A – B). Fe II – III with an outstanding apical retrodorsal spur (Fig. 4 A). Cx IV reaching the DS posterior margin (Figs 4 A, 5 A). Cx IV tuberculate between prodorsal and ventral faces (Figs 4 B, 5 A). Cx IV with a prodorsal apophysis ‘ scythe-shaped’ (subapically curved to posterior), bearing a small accessory blunt branch on its central posterior third (Figs 4 A – B, D, 5 A – B, E). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 4 A – B, 5 A). Tr IV rectangle-shaped (in dorsal view) (Figs 4 A – B, 5 A). Tr IV distal half tuberculate on dorsal face (Fig. 5 A). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (retrolateral largest; prolateral slightly curved to dorsal on the central portion) (Figs 4 A – B, 5 A). Tr IV distal portion with a conical apophysis on prolateral and retrolateral faces (retrolateral largest; prolateral slightly curved to dorsal on the central portion) (Figs 4 B, 5 A). Tr IV distal third with a subconical prominent tubercle on prodorsal and retrodorsal faces (Fig. 5 A). Tr IV ventral face tuberculate (Fig. 4 B). Fe IV sub-straight, arched on the proximal portion towards retrodorsal face (Figs 4 A – B, 5 A). Fe IV dorsal face with four spines (iIiI, the second basalmost spine outstanding and curved to the retrolateral portion) (Figs 4 A, 5 A). Fe IV prodorsal face with four prominent tubercles (the basalmost acuminated) on the distal half and a reduced apical spur (Figs 4 A, 5 A). Fe IV proventral face with a row of four prominent acuminated tubercles on basal two-thirds, and two conical spines on the distal third (Fig. 4 B). Fe IV retroventral face with two prominent tubercles on the basal third, with four conical spines on distal two-thirds (Fig. 4 B). Fe IV retrolateral and retrodorsal faces with a longitudinal row of ordinary tubercles (Figs 4 B, 5 A). Fe IV with a sizeable apical retrodorsal spur (larger than the prodorsal spur) (Figs 4 A, 5 A). Pa IV dorsally covered by few prominent subconical tubercles (Fig. 4 E). Pa IV proventral and retroventral faces with a row of three spines. Ti IV with all faces containing longitudinal rows of spines (proventral, retroventral, and retrodorsal larger than others). Mt IV with all faces containing longitudinal rows of small spines. Mt IV with proventral and retroventral apical spurs. Coloration (in ethanol) (Fig. 4): Ch, ocularium, and carapace background Brilliant Greenish Yellow (98). Posterior portion of the carapace and central area of the mesotergum background Deep Greenish Yellow (100), with the tubercles (and their surrounding spots) Light Yellow Green (119). Paramedian armature of scutal area III, lateral portions of the carapace and AS margins Strong Greenish Yellow (99). Pp and legs I – III background Light Yellow Green (119). Leg IV background Moderate Greenish Yellow (102). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (Fig. 6 A, C). VP ventral surface entirely covered with microsetae of type 1 (Fig. 6 B – C). All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated, on the proximal part of VP (Fig. 6). MS B 1 short, inserted ventrally, and close to A 2 (Fig. 6 B – C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 6). MS D 1 short, closer to C 3 than A 1 (Fig. 6 A – B). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 beside C 1, E 2 placed between C 2 – C 3 (Fig. 6 B – C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 6 A – B). Stylus and its ventral process axis fused basally, forming a prominent trapezoidal-shaped pedestal above the glans (Fig. 6 A). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 6). Stylus without any expansion or flattening, in situ reaching the distal margin of VP (Fig. 6). Ventral process sigmoid, thinner, and as long as the stylus, with an apical flabellum (Fig. 6 B). Flabellum slightly bent ventrad, hand-shaped (with the main branch provided by short spines) (Fig. 6). Female (MHNCI 3621)!: Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 9 (3) / 10 (3) - 7 / 7 - 7 / 7. Remark: Other measurements were not accessed before its lost. Cx IV narrower than the male, with the prodorsal distal apophysis reduced to a single spine and without a retrolateral distal apophysis. Fe IV straight. Fe IV dorsal face with two spines on the proximal third. Fe IV proventral and retroventral faces with a longitudinal row of spines. Fe IV retrodorsal face with two spines on the distal third. Intraspecific variation: It was not detected relevant intraspecific variation among the minor morph / major morph males or among females.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	description	(Figs 7 – 11)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	materials_examined	Type data Discocyrtus guarauna: 2 ♀ syntypes (MZSP 49!, examined), from BRAZIL, Paraná, [Teixeira Soares], Guaraúna. Discocyrtus heteracanthus: ♂ ♀ syntypes (MNRJ 42278!, examined), from BRAZIL, Paraná, ‘ Cachoeirinha’ [= currently Arapoti, now abandoned Railway Station, - 24.149 342 °, - 49.822 454 °]. Records BRAZIL: state of Paraná: Morretes, Porto de Cima [nowadays a district of Morretes] (H. Soares 1945).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	diagnosis	Diagnosis Conapesquius heteracanthus (Mello-Leitão, 1936) can be distinguished from C. brevifemur, C. rectipes, and C. spinifemur due to (1) ocularium and its pair of spines almost forming a right angle (Figs 9 B, 10 C); (2) scutal area III with a pair of outstanding cylinders with a broad base (Figs 8 A, D, F, 9 A – B, 10 A, 10 C); (3) Fe III straight (Figs 8 A, 9 A, C); (4) flabellum hand-shaped, provided with five branches with short spines (Fig. 11 D); (5) females’ Tr IV prolateral face unarmed on the distal portion (Fig. 9 D – E); (6) females’ Fe IV retrolateral face unarmed.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	materials_examined	Non-type material examined BRAZIL: state of Paraná: Antonina: 1 ♂ (IBSP 8679), 3 ♂ 1 ♀ (IBSP 8809), [- 25.4286 °, - 48,7119 °], Rio Cachoeira, 15 – 19. iv. 2004, Hofer, H. leg.; 1 ♀ (UFPR), RPPN Reserva Natural Guaricica (SPVS), 10. iii. 2017, Pinto, A. P. leg.; 2 ♂ (UFPR), idem, 23 – 27. x. 2017, Pinto, A. P. leg.; 2 ♀ (UFPR), idem, Alojamento Bom Jesus, 16 – 20. iv. 2018, Pinto, A. P. leg.; 11 ♂ 6 ♀ 1 juv (MNRJ 377), idem, Trilha dos Fornos, - 25.30 252 °, - 48.66 005 °, 125 m, 16. xi. 2021, Carvalho, R. N. et al. leg.; 1 ♂ 1 ♀ (MNRJ 60568), idem, Trilha da Rede, - 25.302 599 °, - 48.672 713 °, 112 m, 07. xi. 2019, L. N. Ázara, R. N. Carvalho & A. B. Kury leg.; 3 ♂ 2 ♀ 1 juv (MNRJ 60569), idem, Trilha do Corvo, - 25.325 467 °, - 48.675 008 °, 40 m, 08. xi. 2019, L. N. Ázara, R. N. Carvalho & A. B. Kury leg.; 3 ♂ 2 ♀ (MNRJ 60571), idem, Trilha do Ferro, - 25.304 346 °, - 48.680 888 °, 68 m, 10. xi. 2019, L. N. Ázara, R. N. Carvalho & A. B. Kury leg. Guaraqueçaba: 1 ♂ 1 ♀ (MNRJ 60570), Reserva Biológica Bom Jesus, - 25.296 235 °, - 48.613 820 °, 178 m, 09. xi. 2019, L. N. Ázara, R. N. Carvalho & A. B. Kury leg. Morretes: 1 ♂ 1 ♀ (MZSP 1010)!, [- 25.4397 °, - 48.9048 °], ix. 1946, Gofferjé, C. N. leg.; 1 ♂ 1 ♀ (MZSP 1033)!, [- 25.4814 °, - 48.829 °], 1946, Hatschbach, G. G. leg.; 6 ♂ 4 ♀ (MNRJ 5442)!, Parque Estadual do Marumbi, [- 25.433 °, - 48.9166 °], 27. i. 1990, Baptista, R. L. C. leg.; 5 ♂ 2 ♀ (MZSP 18757)!, 1 ♂ 1 ♀ (MZSP 18774)!, idem, 09 – 09. iv. 1999, Pinto-da-Rocha, R. & Chagas Jr., A. leg.; 1 ♂ 1 ♀ (MZSP 36237), idem, x. 1946, Imaguirei, K. leg.; 1 ♂ (MHNCI 6325), Porto de Cima, 08. ix. 1986, Bérnils, R. S. leg.; 1 ♂ (MHNCI 174)!, 1 ♀ (MHNCI 175)!, idem, Prainha, [- 25.4369 °, - 48.8764 °], i. 1944, Leprevost leg.; 1 ♂ 2 ♀ (MHNCI 6341), Véu da Noiva, 18. vi. 1988, Wosiack & Bornschein leg.; 1 ♂ 1 ♀ (MZSP 18807)!, idem, - 25.4166, - 48.933, 08. iv. 1999, Pinto-da-Rocha, R. & Chagas Jr., A. leg.; 1 ♂ 1 ♀ (MNRJ-HS 149)!, Vista Cavalcanti, 24. iii. 1946, Hatschbach, G. leg. Quatro Barras: 1 ♂ (MHNCI 6333), Alto da Serra, [- 25.35 °, - 48.9167 °], 26. v. 1987, Segalla, M. V. leg.; 2 ♂ 2 ♀ (MHNCI 6612 A), 12. viii. 1989, Pinto-da-Rocha, R. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	distribution	Distribution (new records with an asterisk) BRAZIL: state of Paraná: Antonina *, Arapoti, Guaraqueçaba *, Morretes, Quatro Barras *, Teixeira Soares * (Fig. 3).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	description	Redescription MNRJ 42278! (male syntype) and MNRJ 60568 (male) for the external body illustrations and description (Figs 8 – 10); DS, measurements: CW 2.7, CL 1.9, AW 4.9, AL 2.7; Leg I – IV measurements in Table 7; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 12 (3) / 10 (3) - 7 / 7 - 7 / 7. MNRJ-HS 149! for genitalic illustrations (Fig. 11). Dorsum: DS gamma-pyriform, as long as wide, with AS lateral margins strongly convex, widest at scutal area II and thickest at scutal area III, with sinuous posterior margin (Figs 8 A, D, 9 A – B, 10 A, C). DS anterior margin divided by a small central projection in the center and a pair of shallow cheliceral sockets (Figs 8 A, 9 A, 10 A). Carapace anterior portion with two transversal rows of five prominent subconical tubercles, and centrally covered by prominent and ordinary tubercles (Fig. 10 A). Carapace posterior portion with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Fig. 10 A, C). Ocularium elliptical (in dorsal view), high (c. 3 × the eye diameter), perpendicularly placed on the middle of the carapace (Fig. 10 A – C). Ocularium with a pair of parallel spines (c. 2.5 × the eye diameter) (Figs 8 A, D – E, 9 A – B, 10 A – C). AS lateral margins with two rows of tubercles: one external, composed of four-five prominent subconical tubercles at areas II – IV; another internal one with ordinary tubercles from the posterior corner of the carapace to the posterior margin (Fig. 8 A). Mesotergum divided into four clearly defined areas (Figs 8 A, 9 A, 10 A). All scutal areas tuberculate, with almost all tubercles individually covered / surrounded by light-colored spots (Figs 7 A, 9 A, 10 A). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 8 A, D, 9 A, 10 A). Scutal area I with two pairs of prominent tubercles (c. 1.5 × the ordinary tubercles) (Fig. 10 A, C). Scutal area II with a transversal row of eight prominent tubercles (centrally arched to proximal margin) (Fig. 10 A). Scutal area II with anterior-lateral margin slightly embracing the scutal area I, and with posterior-lateral margin embracing the scutal area III (Figs 8 A, 9 A, 10 A). Scutal area III with a pair of remarkable cylindrical structures with a broad base (c. 7 × the ordinary tubercles), with its surroundings covered by ordinary tubercles (Figs 8 A, D, F, 9 A – B, 10 A, C). Scutal area IV with two transversal rows of four prominent subconical tubercles (c. 1.5 × the ordinary tubercles) (Figs 8 D, 10 A). DS posterior border and free tergites I – III each with a transversal row of prominent tubercles (larger ones on the medial portion) (Figs 8 A, F, 9 A, 10 A). Anal operculum tuberculate (Figs 7 C, 8 F). Venter: Cx I – III sub-parallel to each other, individually with ventral longitudinal rows of 7 – 12 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (Fig. 9 C). Cx II with a retroventral distal row of four acuminated tubercles. Cx III with a retroventral distal row of nine acuminated tubercles. Cx IV much larger than the others, directed obliquely (Fig. 9 C). Intercoxal bridges are well marked (Fig. 9 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Fig. 9 C). Cx IV covered by ordinary tubercles. Stigmata are visible (Fig. 9 C). Free sternites with a transverse row of ordinary tubercles. Chelicera: Basichelicerite elongate, bulla well marked (Fig. 10 A), with four marginal setiferous tubercles on the ectal face (Fig. 10 A); hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles. Fe with a ventral basal and a mesal distal setiferous tubercle. Pa unarmed (Figs 8 A, D – E, 9 A – C). Ti with a row of four spines (IiII) on ventro-mesal (Fig. 8 A) and ventro-ectal faces. Ta with two rows of spines — three (IIi) ventro-mesal (Fig. 8 A) and four (IIIi) ventro-ectal. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles. Fe I – III straight (Fig. 9 A – C). Fe and Ti I – III with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles (Fe III proventral and retroventral tubercles larger and sharper than others). Fe II – III with an outstanding apical retrodorsal spur (Fig. 9 A). Cx IV reaching the free tergite I (Figs 8 A, 9 A, 10 A). Cx IV tuberculate between prodorsal and ventral faces (Fig. 8 D). Cx IV with a prodorsal distal apophysis which is detected in two different shapes: (1) ‘ scythe-shaped’ (subapically curved to posterior), bearing a small accessory blunt branch on its central posterior third (Figs 8, 10 K – L), or (2) subconical and thin, centrally curved to posterior (Figs 9 A – C, 10 A, C). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 8 A – B, 9 A, C). Tr IV rectangle-shaped (in dorsal view) (Figs 8 A – C, 9 A – C, 10 A). Tr IV tuberculate on dorsal and ventral faces (Fig. 10 A, F – H). Tr IV distal portion with a transversal apophysis (similar to a hook) on prodorsal and retrodorsal faces (retrodorsal attenuated, with two prominent subconical tubercles above it) (Fig. 10 A, E – F, H). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (retrolateral largest) (Figs 8 B, 10 A). Tr IV prolateral face with two central and one distal prominent subconical tubercles (Figs 8 B, 10 A). Tr IV distal portion with a subconical prominent tubercle on retrodorsal and retrolateral distal faces (Fig. 10 A, E – H). Fe IV sub-straight, arched on the proximal portion towards the prodorsal face (Figs 8 B, D, F, 9 B, 10 E – H). Fe IV dorsal face with six conical spines (IiIiII), all centrally bent to the retrolateral portion (Fig. 10 E – F, H). Fe IV prodorsal, prolateral and retrodorsal faces with a longitudinal row of ordinary tubercles (Fig. 10 E – H). Fe IV proventral face with seven prominent tubercles (interpolated by ordinary ones) and a distalmost conical spine (Fig. 10 F – G). Fe IV retroventral face with two outstanding tubercles and a subconical spine (bent retrolaterad) on proximal half, and a subconical spine (bent retrolaterad) and two conical spines on distal half (Fig. 10 G – H). Fe IV retrolateral face with a prominent tubercle on the proximal half and three conical spines on distal half (Fig. 10 E, G – H). Fe IV apical portion with a sizeable spur on prodorsal and retrodorsal faces (Fig. 10 E – H). Pa IV dorsally covered by prominent subconical tubercles (some are outstanding, mainly on dorsal and retrodorsal faces) (Fig. 10 F). Pa IV proventral and retroventral faces with a row of three spines (Fig. 10 J). Ti IV with all faces containing longitudinal rows of spines (proventral and retroventral larger than others) (Fig. 10 I – J). Mt IV with all faces containing longitudinal rows of small spines. Mt IV with proventral and retroventral apical spurs. Coloration (in vivo) (Fig. 7): DS anterior margin and carapace background Dark Yellowish Brown (78). DS lateral margins (between the anterior portion and the scutal area II) Strong Yellow (84). Mesotergum background, DS posterior margin and free tergites I – III Dark Grayish Olive (111). Mesotergum central diamond layer and the DS posterior margin Strong Greenish Yellow (99). DS ordinary tubercles Light Yellow Green (119). DS and free tergites I – III prominent tubercles Vivid Greenish Yellow (97) or Pale Greenish Yellow (104). Scutal area III paramedian armature Blackish Red (21), with apex Dark Reddish Orange (38). Ch glossier background Deep Yellow Green (118). Pp glossier background Strong Yellow Green (117), irregularly covered by Dark Grayish Olive Green (128) spots. Tr I – III background Strong Greenish Yellow (99). Fe – Mt I – III background combines Deep Greenish Yellow (100) and Dark Grayish Olive (111). Fe II – III retrodorsal apical spur Deep Yellow (85). Cx – Tr IV background Dark Reddish Brown (44). Cx – Fe IV with the apophyses’ apex Strong Reddish Brown (40). Fe – Mt IV background Blackish Red (21). Ti IV distal portion Vivid Yellow (82). Coloration (in ethanol) (Fig. 9): DS anterior margin and carapace background Dark Olive Brown (96). DS lateral margins (between the anterior portion and the scutal area II) Dark Yellow (88). Mesotergum background Olive Black (114). Mesotergum central diamond layer, DS posterior margin and free tergites I – III Light Olive (106). Mesotergum grooves between areas I – IV combines Moderate Yellow (87) and Olive Black (114). DS prominent tubercles Pale Yellow (89). Scutal area III paramedian armature Olive Black (114), with apex Strong Yellowish Brown (74). Ch glossier Dark Grayish Olive Green (128), with honeycombed details Moderate Greenish Yellow (102). Pp and Tr I – III background Light Yellow Green (119) covered by Olive Black (114) spots. Fe – Mt I – III background combines Olive Black (114) and Strong Greenish Yellow (99). Ti III distal portion Light Yellow Green (119). Cx – Tr IV background Dark Yellowish Brown (78). Fe – Mt IV background Dark Olive Brown (96). Ti IV distal portion Vivid Greenish Yellow (97). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (Fig. 11 A – B). VP ventral surface entirely covered with microsetae of type 1 (Fig. 11 B – C). All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated, on the basal half of VP (Fig. 11 A, C). MS B 1 short, inserted ventrally, showing longitudinal asymmetry (proximal to A 2 on the right side, to A 3 on the left side) (Fig. 11 B – C). MS C 1 – C 3 thick and acuminated, forming a triangle on the distal half of VP (Fig. 11 A – C). MS D 1 short, closer to C 3 than A 1 (Fig. 11 C). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 above C 1, E 2 placed between C 1 and C 2 (Fig. 11 C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 11 A, C). Stylus and its ventral process axis fused basally, forming a pedestal above the glans (Fig. 11 A, D). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 11 A – B, C). Stylus without any expansion or flattening, in situ reaching the distal margin of VP (Fig. 11 A). Ventral process bent dorsad, as long and thinner than the stylus (Fig. 11 B, D). Flabellum slightly bent ventrad, hand-shaped (with three branches provided by short spines) (Fig. 11 D). Female (MNRJ 60568) (Fig. 9 D – F): DS, measurements: CW 2.4, CL 1.7, AW 4.1, AL 2.6; Leg I – IV measurements in Table 8; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 9 (3) / 9 (3) - 7 / x - 7 / 7. DS posterior margin concave (Fig. 9 D). AS margins less concave than detected on males (Fig. 9 D). Scutal area III with a pair of paramedian conical spines (c. 20 × the ordinary tubercles) (Fig. 9 D – E). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine and without a retrolateral distal apophysis (Fig. 9 D, F). Fe IV straight (Fig. 9 D – F). Fe IV dorsal face with three spines (Fig. 9 E). Fe IV proventral and retrolateral faces with a row of spines (Fig. 9 E – F). Intraspecific variation: Some variations among minor morph males and major morph males were detected: (1) AS lateral margins with reduced subconical tubercles (Figs 9 A, 10 A); (2) Cx IV prodorsal apophysis not ‘ scythe-shaped’, as a conical structure bent centrally (Figs 9 A – C, 10 A); and (3) Fe – Ti IV with reduced and thinner armature (Fig. 9 B – C). Variation among major morph males: scutal area II with a transversal row of eight or ten prominent tubercles. No relevant intraspecific variation among the females was detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13018870112DDFA130B18FA82.taxon	discussion	Historical taxonomical remarks: After careful examination, it is apparent that the female holotype of Discocyrtus guarauna (MZSP 49) shows conspicuous similarities to the female syntypes of Conapesquius heteracanthus (MNRJ 42278!) and its ordinary specimens studied herein. Since the original description of Discocyrtus guarauna, no additional ordinary specimens have been reported in the literature. However, the type-locality of Teixeira Soares, Paraná, which is present within the Araucaria Forest province, does not align well with the geographical distribution of Conapesquius heteracanthus (most of records from the Atlantic Forest) (Fig. 3). It remains uncertain whether the type-locality was erroneously assigned by Piza (1940 a) or if the lack of further records of Conapesquius heteracanthus in the Araucaria Forest province is a result of limited available material. In the absence of conclusive evidence to refute the original data of Discocyrtus guarauna, it is proposed here as a junior subjective synonym of Conapesquius heteracanthus.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	description	(Figs 12 – 15)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	materials_examined	Type data Discocyrtus rectipes: 1 ♂ 6 ♀ syntypes (SMF RI 785, examined) from BRAZIL, without further locality data; 4 ♂ 3 ♀ (ZHM 1438) from BRAZIL, São Paulo, Ribeirão Pires [misidentification of D. littoralis Mello-Leitão, 1932, as reported by Carvalho 2017: 278]. Records Without further geographic records in the literature.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	diagnosis	Diagnosis Conapesquius rectipes can be distinguished from C. brevifemur, C. heteracanthus, and C. spinifemur due to (1) Ch bulla with spines on anterior mesal margin (as in C. brevifemur) (Fig. 14 A); (2) DS thickest at area I (Figs 13 B, 14 C); (3) Cx I ventral with a transversal row of juxtaposed spines; (4) Cx III retrolateral face with an apophysis (Fig. 13 C); (5) Fe IV dorsal face with four outstanding spines on the proximal half (as in C. heteracanthus) (Fig. 14 E – F, H).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	materials_examined	Material examined BRAZIL: State of Santa Catarina: Blumenau: 1 ♂ 2 ♀ (MZSP 36263), ix. 1955, Gofferjé, C. N. leg.; 1 ♂ (IBSP 5867)!, 1 ♂ (IBSP 5868), 1 ♂ (IBSP 5888)!, 1 ♀ IBSP (5897), 1 ♂ (IBSP 5899), 1 ♂ (IBSP 5930)!, 1 ♂ (IBSP 5948)!, 1 ♀ (IBSP 5956), 1 ♂ (IBSP 5960)!, 1 ♂ (IBSP 5961)!, 1 ♂ IBSP (6028), Parque Natural Municipal Nascentes do Garcia, - 27.0166 °, - 49.15 °, 21 – 28. i. 2003, Equipe Biota leg.; 3 ♂ 1 ♀ (MNRJ 2675)!, Parque Ecológico Spitzkopf, - 27.0 °, - 49.1 °, 03. ii. 1996, Bonaldo, A.; Kury, A. B. & Rocha, R. leg.; 2 ♂ (MZSP 18323)!, 1 ♂ 1 ♀ (MZSP 18332)!, idem, 29. iii. 1999, Pinto-da-Rocha, R.; Bérnils, R. S. & Lingnau, R. leg.; 6 ♂ 5 ♀ (MNRJ 2860), 1 ♂ (MNRJ 2863), Sítio Griebner, Nova Rússia, - 27.01773 °, - 49.09265 °, 362 m, 07. ii. 2022, Martins, P. H. et al. leg. Florianópolis: 1 ♂ (MNRJ 365), Morro da Aranha, 10. i. 2019, Giupponi, A. leg. Ilhota: 1 ♂ (MNRJ 2879), Parque Botânico Morro do Baú, - 26,7998 °, - 48,94 251 °, 276 m, 08. ii. 2022, Martins, P. H. et al. leg.; 1 ♂ 2 ♀ (MNRJ 6653)!, idem, - 26.80, - 48.95, 25. i. 1990, Baptista, R. L. C. & Baptista, A. R. P. leg.; 1 ♂ (MNRJ 6956)!, idem, 04. ii. 1996, Bonaldo, A., Kury, A. B. & Pinto-da-Rocha, R. leg.; 2 ♂ 2 ♀ (MZSP 18656), 01 – 02. iv. 1999, Pinto-da-Rocha, R., Bérnils, R. & Lingnau, R. leg. Indaial: 1 ♂ (MNRJ 3045), 1 ♂ (MNRJ 3050), Instituto de Permacultura Vale do Itajaí (IPEVI), - 26.96 635 °, - 49.18 029 °, 184 m, 09. ii. 2022, Martins, P. H. et al. leg. Itajaí: 1 ♀ (MNRJ 4493)!, 1 ♂ (MZSP 18156)!, [- 26 883 °, - 48,65 °], Rodovia BR 470, 09. iii. 1999, Kury, A. B.; Pinto-da-Rocha, R. & Giupponi, A. leg. Luís Alves: 4 ♂ 4 ♀ (MZSP 29868), Alto Máximo, [- 26 725 °, - 49 055 °], 12. xii. 2005, da Silva, M. B. et al. leg. Porto Belo: 1 ♂ (MNRJ 7420, APA Araçá, [- 27 137 °, - 48 542 °], Antrópica do Caixa d’Aço, 26. vii. 2011, Trivia, A. L. & Chagas-Jr., A. leg.; 9 ♂ (MNRJ 7270)!, 8 ♂ 12 ♀ 1 juv (MNRJ 7400)!, 1 ♂ 3 ♀ (MNRJ 7411)!, APA Araçá, Avançada do Refúgio, 25 - 26. iv. 2011, Trivia, A. L. & Chagas-Jr., A. leg.; 1 ♂ 2 ♀ (MNRJ 7311)!, idem, 24. vii. 48, Trivia, A. L. & Malta, L. leg.; 1 ♂ 1 ♀ 1 juv (MNRJ 7369)!, APA Araçá, Banhado do Caixa-d’aço, 24. vii. 2011, Trivia, A. L. & Malta, L. leg.; 1 ♂ (MNRJ 7383)!, APA Araçá, Banhado do Guinho, 25. vii. 2011, Trivia, A. L. & Malta, L. leg.; 1 ♂ 1 juv (MNRJ 7435)!, APA Araçá, Inicial da Bia, 25. vii. 2011, Trivia, A. L. & Malta, L.; 1 ♂ (MNRJ 7334)!, APA Araçá, Inicial do Guinho, 27. vii. 2011, Trivia, A. L. & Malta, L. leg.; 4 ♂ 1 ♀ (MNRJ 7323)!, 2 ♂ 3 ♀ (MNRJ 7388)!, APA Araçá, Média do Guinho, 25. iv. 2011, Trivia, A. L. & Chagas-Jr., A. leg. Rancho Queimado: 1 ♂ (MCN 1254)!, 1 ♂ (MCN 1255)!, 08 – 11. x. 1994, Bonaldo, A. B. leg.; 1 ♂ (MCN 1263)!, 20. x. 1994, Bonaldo, A. B. leg. São Bento do Sul: 1 ♂ 1 ♀ (MNRJ 2833), 1 ♂ (MNRJ 2844), Centro de Estudos e Pesquisas Ambientais Rugendas, - 26.3238 °, - 49.30 659 °, 639 m, 05. ii. 2022, Martins, P. H. et al. leg.; 1 ♂ (MNRJ 0041)!, 1 ♂ 7 ♀ (MNRJ 9196)!, [- 26,35 °, - 49,3 °], Rio Natal, 14. xii. 1982, Jim, J. et al. leg.; 1 ♂ (MNRJ 6956)!, Serra Alta, [- 26 266 °, - 49 383 °], 26. iii. 1993, Sachsse, R., Silva, S. P. C., Gomes, M. & Peixoto, O. leg. São Francisco do Sul: 1 ♀ (MNRJ 2823), Centro de Estudos e Pesquisas Ambientais Vila da Glória, - 26.22 655 °, - 48.68 365 °, 8 m, 05. ii. 2022, Martins, P. H. et al. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	distribution	Distribution (new records with an asterisk) BRAZIL: state of Santa Catarina: Blumenau *, Florianópolis *, Indaial *, Ilhota *, Itajaí *, Luís Alves *, Porto Belo *, Rancho Queimado *, São Bento do Sul *, São Francisco do Sul * (Fig. 3).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1301E873C12C0FA900BF4FC7A.taxon	description	Redescription IBSP 6028 (male) for the external body illustrations and description (Figs 13 A – C, 14); DS, measurements: CW 2.2, CL 1.5, AW 4.2, AL 2.3; Leg I – IV measurements in Table 9; Right / left tarsal (distitarsal) counts: 5 (3) / 5 (3) - 8 (3) / 9 (3) - 7 / 7 - 7 / 7. MNRJ 0041! for genitalic illustrations (Fig. 15 A – D). Dorsum: DS gamma-pyriform, as long as wide, with AS lateral margins strongly convex, widest at scutal areas II – III and thickest at scutal area I, with sinuous posterior margin (Figs 13 A, 14 A). DS anterior margin divided by a small central projection in the center and a pair of shallow cheliceral sockets (Fig. 14 A). Carapace anterior portion with two transversal rows of five prominent subconical tubercles, and centrally covered by ordinary tubercles (Fig. 14 A). Carapace with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Figs 14 A, C). Ocularium elliptical (in dorsal view), inclined frontwards (in lateral view), high (c. 4.5 × the eye diameter), perpendicularly placed on the anterior portion of the carapace (Figs 13 A – B, 14 A – C). Ocularium with a pair of parallel spines (c. 4 × the eye diameter), inclined frontwards (Figs 13 A – B, 14 A – C). AS lateral borders with two rows of tubercles: one external, composed of six to seven prominent subconical tubercles at areas II – IV; another internal one with ordinary tubercles from the posterior corner of the carapace to the posterior margin (Fig. 14 A). Mesotergum divided into four clearly defined areas (Figs 13 A – B, 14 A, C). All scutal areas tuberculate (Fig 14 A, C). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 13 A, 14 A). Scutal area I with two pairs of prominent tubercles (c. 1.5 × the ordinary tubercles) (Fig. 14 A). Scutal area II with a transversal row of eight prominent tubercles (centrally arched to proximal margin) (Fig. 14 A). Scutal area II posterior-lateral border embracing scutal area III (Figs 13 A, 14 A). Scutal area III with a pair of paramedian outstanding domed-shaped tubercles (c. 5 × the ordinary tubercles), with their surroundings covered with prominent tubercles (Figs 13 A – B, 14 A, C – D). Scutal area IV central portion with a transversal row of four prominent subconical tubercles (c. 1.5 × the ordinary tubercles) (Fig. 14 A, C). DS posterior margin and free tergites I – III each with a transversal row of prominent tubercles (larger on the central portion) (Fig. 14 A). Anal operculum tuberculate. Venter: Cx I – III sub-parallel to each other, each with ventral longitudinal rows of 6 – 11 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (Fig. 13 C). Cx II with a retroventral distal row of four acuminated tubercles. Cx III with a retroventral distal row of 10 acuminated tubercles. Cx IV much larger than the others, directed obliquely (Fig. 13 C). Intercoxal bridges are well marked (Fig. 13 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Fig. 13 C). Cx IV covered by ordinary tubercles. Stigmata are visible (Fig. 13 C). Free sternites with a transverse row of ordinary tubercles. Chelicera: Basichelicerite elongate, bulla well-marked (Fig. 13 A), with marginal setiferous tubercles — two anterior mesal, two anterior ectal, and one or two lateral mesal (Fig. 14 A); hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles. Fe with a ventral basal and a mesal distal setiferous tubercle. Pa unarmed. Ti with two rows of spines — four (IiII) ventro-mesal (Fig. 13 C) and five (IiIii) ventro-ectal. Ta with two rows of spines — three (Iii) ventro-mesal (Fig. 13 C) and four (IiIi) ventro-ectal. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles. Fe I – II straight (Fig. 13 C); Fe III sub-straight (Fig. 13 A, C). Fe and Ti I – III with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles (Fe III proventral and retroventral tubercles larger than others). Fe II – III with an outstanding apical retrodorsal spur (Fig. 13 A, C). Cx IV reaches the DS posterior margin (Figs 13 A, 14 A). Cx IV tuberculate between prodorsal and ventral faces (Fig. 14 A). Cx IV with a prodorsal apophysis ‘ scythe-shaped’, with a conical main branch (subapically bent to posterior), bearing a small accessory blunt branch on its central posterior third (Figs 13 B, 14 A). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 13 A, C, 14 A). Tr IV rectangle-shaped (in dorsal view) (Figs 13 A, C, 14 A). Tr IV distal portion with a transversal apophysis (screwdriver tip shaped) on prodorsal face (Fig. 14 A, E – F). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (retrolateral largest) (Fig. 14 A, G – H). Tr IV with two prominent subconical tubercles (one central and other distal) on prolateral face (Fig. 14 A, E). Tr IV tuberculate on ventral face (Fig. 14 F – H). Tr IV distal portion with a subconical prominent tubercle on retrodorsal and retrolateral faces (Fig. 14 A, E, G – H). Fe IV sub-straight, arched on the central portion towards prodorsal face (Figs 13 B, 14 E – H). Fe IV dorsal face with five conical spines (iiIii), with the four proximal most bent retrolaterad (Fig. 14 E – F, H). Fe IV prodorsal face with a row of six prominent tubercles (Fig. 14 E – F). Fe IV prolateral face with three prominent tubercles on proximal third and two prominent tubercles on distal third (Fig. 14 E – G). Fe IV proventral face with seven prominent tubercles and a distal conical spine (Fig. 14 F – G). Fe IV ventral face with one prominent tubercle on proximal third (Fig. 14 G). Fe IV retroventral with three outstanding tubercles and a distalmost subconical spine (all bent retrolaterad) on proximal half, and two conical spines on distal half (Fig. 14 G – H). Fe IV retrolateral face with a row of nine prominent tubercles on the proximal half and three prominent tubercles on the distal half (Fig. 14 E, G – H). Fe IV retrodorsal face with ordinary tubercles on proximal half, and four prominent tubercles and one conical spine on distal half (Fig. 14 E, H). Fe IV apical portion with a sizeable spur on prodorsal and retrodorsal faces, and two subconical outstanding tubercles on dorsal and retrodorsal faces (Fig. 14 E – F, H). Pa IV dorsally covered by prominent subconical tubercles (Fig. 14 E – F, H – I). Pa IV proventral and retroventral faces with a row of three spines (Fig. 14 J). Ti IV with all faces containing longitudinal rows of spines (proventral, retroventral, retrolateral and retrodorsal larger than others) (Fig. 14 I – J). Mt IV with all faces containing longitudinal rows of small spines. Mt IV with proventral and retroventral apical spurs. Coloration (in vivo) (Fig. 12): DS lateral inner margins and free tergites background Dark Grayish Olive Green (128). DS lateral outer margins between the anterior portion and scutal area II and DS posterior margin combines Strong Yellowish Brown (74) and Dark Yellow (88). Prominent tubercles on AS and free tergites Deep Yellow (85). Carapace, ocularium, and mesotergum backgrounds Dark Grayish Yellowish Brown (81), with ocularium’s pair of spines Dark Yellowish Brown (78). Carapace posterior tubercles, mesotergum central diamond layer of tubercles, and the DS posterior margin Dark Yellow (88). Mesotergum grooves between scutal areas I – IV Dark Greenish Yellowish Green (151). Scutal area III paramedian outstanding armature Dark Reddish Orange (38). Ch and Pp glossier background Deep Greenish Yellow (100), irregularly covered by Dark Grayish Olive Green (128) spots. Cx Pp and I – III background Moderate Olive Brown (95). Tr – Mt I – III background Moderate Olive (107). Fe II – III retrodorsal apical spur Deep Yellow (85). Cx – Tr IV and Fe IV basal two-thirds background combines Blackish Red (21) and Deep Reddish Brown (41). Cx IV prodorsal apophysis glossier background Reddish Black (24). Cx IV basal two-thirds, Fe IV distal half and Pa IV with tubercles’ apex Deep Yellow (85). Apex of the Cx IV retrodorsal apophysis and Tr IV retrolateral apophyses with Strong Brown (55). Cx IV prodorsal distal tubercles, apex of the Cx IV prodorsal apophysis and of Fe IV spines Vivid Reddish Orange (34). Tr IV and Fe IV proximal half with tubercles’ apex, spines and apophyses Strong Reddish Brown (40). Fe IV distal third and Pa – Mt IV background Dark Grayish Olive (111). Ti I – IV distal portion Moderate Greenish Yellow (102). Coloration (in ethanol) (Fig. 13): DS anterior and posterior margins, scutal area IV, and free tergites Dark Olive Brown (96). DS lateral outer margins (between the anterior portion and the scutal area II) and free tergites I – III posterior margins Grayish Yellow (90). DS lateral inner margins Dark Grayish Yellowish Brown (81). Carapace, scutal areas I – III, and Cx IV background Deep Yellowish Brown (75). Grooves between the carapace and scutal areas I – IV Dark Grayish Yellow (91). Carapace, mesotergum, free tergites, and Cx IV with tubercles (and their surroundings) Light Orange Yellow (70). Scutal area III paramedian outstanding armature Moderate Olive (107). Ch and Pp background Light Yellow Green (119), with honey-combed details Dark Grayish Olive Green (128) (leg I with that color pattern inverted). Cx Pp and Cx I – III background Moderate Olive Brown (95). Fe – Mt II – III background Pale Yellow (89), with details Dark Olive (108). Cx IV distal third and prodorsal apophysis background Olive Black (114). Tr – Mt IV background combines Dark Grayish Yellowish Brown (81) and Dark Yellow (88). Tr – Pa IV with the apex of the tubercles, spines and apophyses Dark Greenish Yellow (103). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (Fig. 15 A – B). VP ventral surface entirely covered with microsetae of type 1 (Fig. 14 B – C). All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated, on the basal part of the VP (A 2 and A 3 placed at the same height) (Fig. 15 A – C). MS B 1 short, inserted ventrally, and close to A 2 (Fig. 15 B – C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 15 A – C). MS D 1 short, closer to C 3 than A 1 (Fig. 15 A – C). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 beside MS C 1, E 2 between C 2 – C 3 (Fig. 15 B – C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 15 A, C). Stylus and its ventral process axis fused basally, forming a pedestal above the glans (Fig. 15 A, C). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 15 A, C – D). Stylus without any expansion or flattening, in situ reaching the distal margin of VP (Fig. 15 A – C). Ventral process bent dorsad, as long and thinner than the stylus (Fig. 15 A, C). Flabellum slightly bent ventrad, hand-shaped (with the main branch provided by short spines) (Fig. 15 A, C – D). Female (IBSP 5956) (Fig. 13 D – F): DS, measurements: CW 2.8, CL 2.0, AW 5.5, AL 3.5; Leg I – IV measurements in Table 10; Right / left tarsal (distitarsal) counts: 5 (3) / 5 (3) - 8 (3) / 7 (3) - 7 / 7 - 7 / 7. DS posterior margin concave (Fig. 13 D – E). AS margins less concave than detected on males (Fig. 13 D). Scutal area III with a pair of paramedian conical spines (c. 20 × the ordinary tubercles) (Fig. 13 D – E). Cx IV is narrower than males, with the prodorsal distal apophysis reduced to a single spine and without a retrolateral distal apophysis (Fig. 13 D, F). Fe IV straight (Fig. 13 D, F). Fe IV dorsal face with four spines (Fig. 13 D – E). Fe IV proventral and retrolateral faces with a row of spines (Fig. 13 F). Fe IV retrodorsal face with two spines on the distal third. Intraspecific variation: Some variations among minor morph males and major morph males were detected: (1) Cx IV prodorsal apophysis less developed and bent backward; and (2) Fe – Ti IV with reduced and thinner armature. No relevant intraspecific variation among the females was detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	description	(Figs 16 – 20)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	materials_examined	Type data Discocyrtus spinifemur: ♂ holotype (MHNCI 177!, examined), from BRAZIL, Paraná, Curitiba, Barigui. Records BRAZIL: state of Paraná: Curitiba (Soares and Soares 1947 a).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	diagnosis	Diagnosis Conapesquius spinifemur can be distinguished from C. brevifemur, C. heteracanthus, and C. rectipes due to (1) Ch bulla with anterior mesal margin unarmed (Fig. 17 A); (2) DS gamma type (Figs 16 A, 17 A, 18 A); (3) DS thickest at area III (as in C. heteracanthus) (Figs 16 E, 17 B, 18 B); (4) Fe IV with three outstanding spines on the proximal half (as in C. heteracanthus) (Figs 16 A, 17 A, 18 F – G, I); (5) MS A 1 on the central part of the VP (Fig. 19 A – C); (6) MS B below MS A 3 (Fig. 19 B – C); (7) Ventral process of the glans convex dorsad (Fig. 19 C); (8) Flabellum hand-shaped, provided with three branches with short spines (Fig. 19 D).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	materials_examined	Non-type material examined BRAZIL: state of Paraná: [Campina Grande do Sul]: 3 ♂ 12 ♀ 2 juv (MHNCI 6538), Represa Capivari, [- 25.177 °, - 48.997 °], 05. vi. 1989. Paranaguá: 1 ♂ (MHNCI 6359) PR, Ilha do Mel, [- 25.52 °, - 48.34 °], x. 1988, Moraes, V. leg.; 1 ♀ (MHNCI 6324), Morro da Baleia, 09. iv. 1988, Bedin, S. & Moraes, V. leg. State of Santa Catarina: Florianópolis: 5 ♂ (MNRJ 2458)!, [- 27.61 °, - 48.49 °], Mata do morro atrás dos prédios APAE, ÚNICA e SESI, 15 - 17. xii. 1999, Giupponi, A. P. L. & Pedroso, D. R. leg. Paulo Lopes: 1 ♂ 1 ♀ (MZSP 28477), [- 27.954 °, - 48.722 °], Parque Estadual do Tabuleiro, 10 – 20. i. 2003, Equipe Biota leg. Rancho Queimado: 1 ♀ (MCN 1292), [- 27.707 °, - 49.071 °], 15 - 18. xi. 1995, Bonaldo, A. B. leg. São Francisco do Sul: 5 ♀ (UFMG 10424), Centro de Estudos e Pesquisas Ambientais Vila da Glória, - 23.2204 °, - 48.688 °, 12 - 15. xii. 2011, Magalhães, I. L. F. et al. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	distribution	Distribution (new records with an asterisk) BRAZIL: state of Paraná: Campina Grande do Sul *, Curitiba, Paranaguá *. State of Santa Catarina: Florianópolis *; Paulo Lopes *, Rancho Queimado *, São Francisco do Sul * (Fig. 3).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302387371298FC280DF9FCF9.taxon	description	Redescription MHNCI 177! (male holotype) and MHNCI 6538 (male) for the external body illustrations and description (Figs 16, 17 A – C, 18); DS, measurements: CW 2.9, CL 2.0, AW 6.0, AL 3.2; Leg I – IV measurements in Table 11; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 9 (3) / 9 (3) - 7 / 7 - 7 / 7. MNRJ 2458! for genitalic illustrations (Fig. 19). Dorsum: DS gamma, as long as wide, with AS lateral margins strongly convex, widest at areas II – III and thickest at area III, with sinuous posterior margin (Figs 16 A, 17 A – B, 18 A – B). DS anterior margin divided by a small central projection in the centre and a pair of shallow cheliceral sockets (Fig. 18 A). Carapace with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Figs 17 A – B, 18 A – B). Ocularium elliptical (in dorsal view), slightly inclined frontwards (in lateral view), high (c. 3 × the eye diameter), perpendicularly placed on the proximal portion of the carapace (Figs 16 A, C, E, 17 A – B, 18 A – C). Ocularium with a pair of parallel spines (c. 1.5 × the eye diameter), slightly inclined frontwards (in lateral view) (Figs 16 A, C, E, 17 A – B, 18 A – C). AS lateral borders with two rows of tubercles: one external, composed of four – five prominent subconical tubercles at scutal areas II – IV; another internal one with ordinary tubercles from the anterior corner of the carapace to the posterior margin (Figs 17 A, 18 A). Mesotergum divided into four clearly defined areas (Figs 16 A, E, 17 A – B, 18 A – B). All scutal areas tuberculate (Figs 17 A – B, 18 A – B). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 16 A, 17 A, 18 A). Scutal area I with three pairs of prominent columnshaped tubercles (the two centralmost c. 3 × the ordinary tubercles; the lateralmost 2 × the ordinary ones) (Fig. 18 A – B). Scutal area II with two pairs of prominent tubercles (c. 2 × the ordinary tubercles) (Fig. 18 A – B). Scutal area II posterior-lateral border embracing the scutal area III (Figs 16 A, 17 A, 18 A). Scutal area III with a pair of paramedian outstanding subconical spines (c. 10 × the ordinary tubercles), with their posterior surroundings covered with prominent tubercles (c. 2 × the ordinary tubercles) (Figs 16 A, C – E, 17 A – B, 18 A – B, D). Scutal area IV central portion with a transversal row of six prominent subconical tubercles (c. 2 × the ordinary tubercles) (Fig. 18 A – B). DS posterior margin and free tergites I – III each with a transversal row of prominent tubercles (larger on the central portion) (Fig. 18 A). Anal operculum tuberculate (Fig. 16 B, D). Venter: Cx I – III sub-parallel to each other, each with ventral longitudinal rows of 8 – 12 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (Fig. 16 B). Cx II with a retroventral distal row of five acuminate tubercles. Cx III with a retroventral distal row of 10 acuminate tubercles. Cx IV much larger than the others, directed obliquely (Figs 16 B, 17 C). Intercoxal bridges are well marked (Figs 16 B, 17 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Figs 16 B, 17 C). Cx IV covered by ordinary tubercles. Stigmata are visible (Figs 16 B, 17 C). Free sternites with a transverse row of ordinary tubercles. Chelicera: Basichelicerite elongate, bulla well-marked (Figs 16 A, C, 18 A), with marginal setiferous tubercles — one lateral mesal, three lateral ectal and three anterior-posterior (Fig. 18 A); hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles. Fe with a ventral basal and a mesal distal setiferous tubercle (Figs 16 B, 17 C). Pa unarmed (Figs 16 E, 17 C). Ti with two rows of spines — four (IiIi) ventro-mesal and five (IiIii) ventro-ectal (Fig. 16 C). Ta with two rows of spines — three (IIi) or two (Ii) ventro-mesal and four (IiIi) ventro-ectal (Fig. 16 C). Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles (Fig. 16 B). Fe I – II straight (Figs 16 A, 17 A). Fe III sub-straight (Figs 16 A – B, 17 A). Fe and Ti I – II with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles (Fig. 16 C). Fe III and Ti III with prodorsal, proventral, retroventral, and retrodorsal rows of ordinary tubercles (Fe III proventral and retroventral tubercles larger than others) (Fig. 16 B). Fe II – III with an outstanding apical retrodorsal spur (Figs 16 A, 17 A – C, 18 E). Cx IV reaches the DS posterior margin (Figs 16 A, 17 A, 18 A). Cx IV tuberculate between prodorsal and ventral faces (Fig. 18 A). Cx IV with a prodorsal conical apophysis (subapically bent to posterior), bearing a small accessory blunt branch on its central posterior third (Figs 16 A – E, 17 A – C, 18 A – B). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 16 A – B, 17 A, 18 A). Tr IV rectangle-shaped (in dorsal view) (Figs 16 A – B, E, 17 A – C, 18 A). Tr IV central portion with a prominent tubercle on the dorsal and prolateral faces (Fig. 18 A, F – G). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (retrolateral largest) (Fig. 18 A, F). Tr IV distal portion with a transversal apophysis (similar to a hook) on prodorsal and retrodorsal faces, and a subconical prominent tubercle on retrolateral face (Fig. 18 A, F – I). Tr IV tuberculate on ventral face (Fig. 18 G – I). Fe IV sub-straight, arched on the central portion towards retrodorsal face (Figs 16 B – C, E, F – I). Fe IV dorsal face with six conical spines (IiIiIi) (the four proximal-most bent retrolaterad) and an apical prominent conical tubercle (Fig. 18 F – G, I). Fe IV prodorsal face with seven prominent tubercles (Fig. 18 F – G). Fe IV proventral face with 10 prominent tubercles (the three distalmost acuminated) (Fig. 18 G – H). Fe IV retroventral face with (1) one basalmost prominent subconical tubercle followed by a subconical spine (bent retrolaterad) on the proximal third, (2) a subconical spine on the medial third, and (3) two conical spines on the distal third (Fig. 18 H – I). Fe IV retrolateral face with (1) one prominent subconical tubercle on the proximal third, (2) a subconical spine on the medial third, and (3) two conical spines on the distal third (the basalmost largest) (Fig. 18 F, H – I). Fe IV retrodorsal face with a row of ordinary tubercles and a sizeable apical spur (Fig. 18 F, I). Pa IV dorsally covered by prominent and acuminated tubercles (retrolateral and retrodorsal larger and shaper than others) (Fig. 18 F – I). Pa IV proventral and retroventral faces with rows of four and three spines, respectively (Fig. 18 F – G, I). Ti IV with all faces containing longitudinal rows of spines (proventral, retroventral, and retrolateral larger than others) (Fig. 18 F – I). Mt IV with all faces containing longitudinal rows of small spines. Mt IV with proventral and retroventral apical spurs. Coloration (in ethanol) (Fig. 16): Carapace and ocularium background Brilliant Greenish Yellow (98). Scutal areas I – IV background and free tergites I – III background Light Greenish Yellow (101). AS lateral margins and the outer half of scutal areas I – II Grayish Greenish Yellow (105). Pair of paramedian spines on scutal area III Moderate Olive (107). Ch background Light Greenish Yellow (101). Pp and legs I – III background Light Yellow Green (119). Cx IV with (1) proximal third Moderate Greenish Yellow (102), (2) central third Strong Yellowish Brown (74), and (3) distal third Dark Grayish Olive (111). Tr IV background Deep Yellow (85). Fe IV with the proximal half background Dark Yellow (88). Fe IV distal half and Tr – Mt IV background Light Yellow Green (119). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (Fig. 19 A – B). VP ventral surface entirely covered with microsetae of type 1 (Fig. 19 B – C). All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated — A 1 on the central portion of the VP, A 2 beside A 3 on the basal half of VP (Fig. 19 A – C). MS B 1 short, inserted ventrally, below A 3 (Fig. 19 B – C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 19 A – C). MS D 1 short, as close to A 1 as C 3 (Fig. 19 A, C). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 beside C 1, E 2 beside C 3 (Fig. 19 C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 19 A, C). Stylus and its ventral process axis fused basally, forming a pedestal above the glans (Fig. 19 C – D). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 19 A, C – D). Stylus without any expansion or flattening, in situ reaching the distal margin of VP. Ventral process bent dorsad, as long and thinner than the stylus (Fig. 19 A, C). Flabellum slightly bent ventrad, hand-shaped (with three branches provided by short spines) (Fig. 19 A, C – D). Female (MHNCI 6538) (Figs 17 D – F, 20): DS, measurements: CW 2.8, CL 2.0, AW 5.2, AL 2.8; Leg I – IV measurements in Table 12; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 8 (3) / 9 (3) - 7 / 7 - 7 / 7. DS gamma-pyriform (Figs 17 D, 20 B). Ocularium lower than in males (Figs 17 D – E, 20 B). AS lateral margins less concave than in males (Figs 17 D, 20 B). Scutal area III with a pair of paramedian conical spines (c. 20 × the ordinary tubercles) (Figs 17 D – E, 20 B). DS posterior margin and free tergites I – III with a transversal row of outstanding tubercles (Figs 17 D – E, 20 B). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine and without a retrolateral distal apophysis (Figs 17 D, F, 20 B). Tr IV unarmed on prolateral proximal and retrodorsal distal halves (Figs 17 D, 20 B – D). Tr IV prodorsal distal face with two prominent subconical tubercles on the distal third (Fig. 20 B – C). Tr IV retrolateral distal apophysis forming a 90 ° angle in relation to the Tr IV longitudinal axis (Fig. 20 B – D). Fe IV straight (Fig. 20 C – D). Fe IV with four – five spines on the dorsal face (Fig. 20 C). Fe IV with a row of spines (iiIiI) on the retrolateral face (Fig. 20 C – D). Fe IV with a sizeable spur on the prodorsal apical face (Fig. 20 C). Ti IV (in dorsal view) with tubercles not clearly organized in longitudinal rows (Fig. 20 B). Intraspecific variation: Some variations between minor morph males and major morph males were detected: (1) Cx IV prodorsal apophysis less developed and bent to posterior; and (2) Fe-Ti IV thinner, with a smaller and thinner armature. No evident intraspecific variation among the major morph males and among females were detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302887371640FCAA0CCDF96E.taxon	description	ZooBank LSID: urn: lsid: zoobank. org: act: 7 E 36 E 340 - D 653 - 461 C- 9417 - 7410178 B 4 A 04.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302887371640FCAA0CCDF96E.taxon	type_taxon	Type species: Iamarinus pontesi gen. et sp. nov.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302887371640FCAA0CCDF96E.taxon	etymology	Etymology: Iamarinus is named in honor of biologist Atila Iamarino, who actively engaged in science communication, delivering significant contributions through his work, which have been immensely valuable to the Brazilian population in recent years. Gender masculine.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302887371640FCAA0CCDF96E.taxon	diagnosis	Diagnosis: Iamarinus can be distinguished from the other genera of Neopachylinae due to (1) DS posterior margin full extension with a transversal row of conspicuous tubercles (Figs 117, 126); (2) Tr IV long, with twice or more the Tr III longitudinal size (Figs 116, 120 – 122, 126, 131 – 134); (3) Fe IV retrolateral face with spines interpolated (by unarmed spaces) on the proximal third (Figs 131, 133 – 134); (4) Fe IV retrolateral face with a pair of outstanding spines (distalmost largest) on the central third (Figs 115 – 116, 131, 133 – 134); (5) VP basal half 1 / 3 wider than the distal half (Figs 136, 138); (6) Females’ Fe IV dorsal face with two prominent spines on the distal third (Figs 113, 141).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302887371640FCAA0CCDF96E.taxon	distribution	Distribution: BRAZIL: states of Paraná and Santa Catarina (Fig. 21).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	description	(Figs 22 – 23)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	materials_examined	Type data Discocyrtus fenax: ♂ holotype (MZSP 18158, examined) from BRAZIL, state of Santa Catarina, close to road BR 470, near Itajaí; ♂ paratype (SMF RI 812) from BRAZIL, state of Santa Catarina, Joinville. Records Without further literature records.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	diagnosis	Diagnosis Iamarinus fenax can be distinguished from I. pontesi due to: (1) Scutal area I with two pairs of paramedian conspicuous tubercles (Kury et al. 2018 b: fig. 10 A); (2) Cx IV prodorsal distal apophysis with distal portion forming a 90 ° angle in relation to the longitudinal axis (Kury et al. 2018 b: fig. 10 A); (3) Cx IV prodorsal distal apophysis elongated, bent centrad and distally sinuous (Kury et al. 2018 b: fig. 10 A); (4) Cx IV prodorsal distal apophysis without a secondary branch (Kury et al. 2018 b: fig. 10 D); (5) Fe IV sub-straight (Kury et al. 2018 b: fig. 10 E); (6) Ti IV with an apical regular spine on retroventral face (Kury et al. 2018 b: fig. 10 H).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	materials_examined	Non-type material examined BRAZIL: state of Santa Catarina: Blumenau: 5 ♂ (MZSP 36268), iii. 1987, Gofferjé, C. N. leg.; 1 ♂ 3 ♀ (MHNCI 6305), 2 ♂ 4 ♀ (MZSP 36294), i. 1988, Gofferjé, C. N. leg.; 2 ♂ 1 ♀ (MHNCI 6348), Centro, iii. 1987, Gofferjé, C. N. leg. Penha: 1 ♂ 3 ♀ (MHNCI 6822), Armação, i. 1989, Gofferjé, C. N. leg.; 8 ♂ 14 ♀ 2 juv (MZSP 36288), idem, xii. 1969, Gofferjé, C. N. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	distribution	Distribution (new records with an asterisk) BRAZIL: state of Santa Catarina: Blumenau *, Itajaí, Joinville, Penha * (Fig. 21).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130288735110BF93E0D68F923.taxon	description	Male’s redescription See the original description in Kury et al. (2018 b). Female description (MHNCI 6305) (Figs 22 – 23). DS, measurements: CW 2.9, CL 2.2, AW 5.3, AL 3.3; Leg I – IV measurements in Table 13; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 9 (3) / 9 (3) - 7 / 7 - 7 / 7. DS lambda (Figs 22 A, 23 C). Ocularium elliptical (in dorsal view), high (c. 2.5 × the eye diameter), perpendicularly placed on the central portion of the carapace (Figs 22 A, D, 23 B – C). Ocularium with a pair of divergent spines (c. 3 × the eye diameter) (Figs 22 A, D, 23 B – C). AS lateral margins with one spiniform apophysis at scutal areas II – III (Figs 22 A – B, 23 C). Scutal area III with a paramedian pair of subconical spines (Figs 22 A – B, D, 23 C). Scutal area IV with ordinary tubercles (Figs 22 A – B, 23 A – C). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine and a reduced retroventral distal apophysis (Figs 22 A, C – D, 23 C). Tr IV prodorsal distal half and prolateral entire length portions with prominent tubercles (Figs 22 A, C, 23 D). Tr IV with a proximal and a distal conical apophysis (distalmost largest) on retrolateral face, with a reduced conical apophysis on the central portion (Figs 22 C, 23 D – E). Fe IV straight and thinner than the males (Figs 22 A, C – D, 23 D – E). Fe IV dorsal face with four spines (one on proximal third, two on central third and one on distal third) (Figs 22 A, D, 23 D). Fe IV prolateral face with a row of acuminated tubercles (Fig. 23 D – E). Fe IV proventral and retroventral faces with a row of acuminated tubercles on the proximal half and three – four conical spines on distal half (Fig. 23 E). Fe IV retrolateral face with a row of conical spines (iiiI on the proximal half, Iii on the distal half) (Fig. 23 D – E). Intraspecific variation: Some variations among minor morph males and major morph males were detected: (1) DS narrower; (2) Cx IV prolateral and retrolateral apophyses reduced; and (3) Fe IV thinner, with reduced proximal spines. Variation among major morph males: the Fe IV torsion, which can be more or less prominent. No relevant intraspecific variation among females was detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	description	(Figs 24 – 28) ZooBank LSID: urn: lsid: zoobank. org: act: 5 C 224 A 36 - 7322 - 43 D 9 - B 45 A-C 354 D 9 DC 2 DDF.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	etymology	Etymology Iamarinus pontesi is designated in acknowledgment of the substantialcontributionsbyconservationistJoãoPontestothepreservation and conservation of biodiversity in the Reserva Natural Guaricica, situated in Antonina, Paraná, Brazil. Gender masculine.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	materials_examined	Type data Iamarinus pontesi: ♂ holotype (MNRJ 2789), 3 ♂ paratypes 3 ♀ paratypes 1 juv (MNRJ 60562), from BRAZIL, state of Paraná, Antonina, ReservaBiológicaBomJesus, - 25.296 235 °, - 48.613 820 °, 178 m, 09. xi. 2019, Ázara, L. N., Carvalho, R. N. & Kury, A. B. leg.; 3 ♂ paratypes ♀ paratype (MNRJ 378), from BRAZIL, state of Paraná, Antonina, Reserva Natural Guaricica, Trilha dos Fornos, - 25,30 252 °, - 48,66 005 °, 125 m, 16. xi. 2021, Carvalho, R. N. et al. leg.; ♂ paratype ♀ paratype (MNRJ 60561), from BRAZIL, state of Paraná, Antonina, Reserva Natural Guaricica, Trilha da Rede, - 25.302 599 °, - 48.672 713 °, 112 m, 07. xi. 2019, Ázara, L. N., Carvalho, R. N. & Kury, A. B. leg.; ♂ paratype (MHNCI 6429), from BRAZIL, state of Paraná, Guaraqueçaba, Casa da SEMMA, 04. iii. 1988, Bornschein & Motta leg.; 2 ♂ 2 ♀ (MHNCI 6835), from BRAZIL, state of Paraná, Piraquara, Banhado, 13. i. 1991, Pinto-da-Rocha, R. & Bérnils, R. S. leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	diagnosis	Diagnosis Iamarinus pontesi can be distinguished from I. fenax due to: (1) Scutal area I with three pairs of paramedian conspicuous tubercles (Figs 25 A, 26 A); (2) Cx IV prodorsal distal apophysis with distal portion forming an acute angle in relation to the longitudinal axis (Figs 25 A, 26 A); (3) Cx IV prodorsal distal apophysis short, slightly bent distad (Fig. 25 A – B, D); (4) Cx IV prodorsal apophysis with a second branch on central third (Figs 25 A – C, 26 A, D); (5) Fe IV sinuous (Figs 25 B, 26 F, H); (6) Ti IV with an apical bifurcated spine on retroventral face (Fig. 26 F, I).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	distribution	Distribution BRAZIL: state of Paraná: Antonina, Guaraqueçaba, Piraquara (Fig. 21).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1302A872F1198F9710C45FA32.taxon	description	Redescription MNRJ 2789 (male) for the external body illustrations and description; DS, measurements: CW 3.3, CL 2.4, AW 6.0, AL 3.1; Leg I – IV measurements inTable 14; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 10 (3) / 10 (3) - 7 / 7 - 7 / 7. MNRJ 60562 (male) for genitalic illustrations. Dorsum: DS gamma-pyriform, as long as wide, with AS lateral margins strongly convex, widest and thickest at scutal area III, with sinuous posterior margin (Figs 24 A – B, 25 A – B, 26 A, D). DS anterior margin divided by a small central projection in the center and a pair of shallow cheliceral sockets (Figs 25 A, 26 A). Carapace posterior portion with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Figs 25 A, 26 A, D). Ocularium elliptical (in dorsal view), high (c. 4 × the eye diameter), perpendicularly placed in the middle of the carapace (Figs 24 A – B, 25 A – B, 26 A – B, D). Ocularium with a pair of almost parallel spines (c. 3 × the eye diameter) (Figs 24 A – B, 25 A – B, 26 A – B, D). AS lateral margins with two rows of tubercles: one external, composed of four – five prominent subconical tubercles at areas II – IV; another internal one with ordinary tubercles from the posterior corner of the carapace to the posterior margin (Figs 25 A, 26 A). Mesotergum is divided into four clearly defined areas (Figs 24 A, 25 A, 26 A). All scutal areas tuberculate (Figs 25 A, 26 A, D). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 24 A, 25 A, 26 A). Scutal areas I – II with three pairs of conspicuous tubercles (c. 2 × the ordinary tubercles) (Figs 25 A, 26 A, D). Scutal area II posterior-lateral margin embracing the scutal area III (Figs 24 A, 25 A, 26 A). Scutal area III with a pair of paramedian outstanding subconical spines (c. 14 × the ordinary tubercles) (Figs 24 A – C, 25 A, 26 A, C – D). Scutal area IV central portion with a transversal row of five – six prominent tubercles (c. 2 × the ordinary tubercles) (Figs 24 C, 25 A, 26 A, D). DS posterior margin and free tergites I – III each with a transversal row of tubercles, growing in size towards the central portion (Figs 24 C, 26 A). Anal operculum tuberculate. Venter: Cx I – III sub-parallel to each other, each with ventral longitudinal rows of 8 – 13 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others). Cx II with a retroventral distal row of six acuminated tubercles. Cx III with a retroventral distal row of 10 acuminated tubercles. Cx IV much larger than the others, directed obliquely (Fig. 25 C). Intercoxal bridges are well marked (Fig. 25 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Fig. 25 C). Cx IV covered by ordinary tubercles (Fig. 25 C). Stigmata are visible (Fig. 25 C). Free sternites with a transverse row of ordinary tubercles. Chelicera: Basichelicerite elongate, bulla well marked (Fig. 26 A), with marginal setiferous tubercles — two or three lateral ectal, one or two posteriors, two lateral mesal (Fig. 26 A); hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles (Fig. 25 C). Fe with a ventral basal and a mesal distal setiferous tubercle (Fig. 25 C). Pa unarmed (Fig. 25 C). Ti with two rows of four (IiIi) spines on ventro-mesal and ventro-ectal faces. Ta with two rows of spines — three (IIi) ventro-mesal and four (IiIi) ventro-ectal. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles. Fe I – II straight (Fig. 25 A). Fe III sub-straight (Fig. 25 A). Fe and Ti I – II with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles. Fe III and Ti III with prodorsal, proventral, retroventral, and retrodorsal rows of tubercles (Fe III proventral and retroventral tubercles larger and sharper than others). Fe III with an apical retrodorsal spur (Fig. 25 A). Cx IV reaches scutal area IV (Figs 25 A, 26 A). Cx IV tuberculate between prodorsal and ventral faces (Figs 25 A – C, 26 A). Cx IV with a prodorsal subconical apophysis, slightly bent to posterior, bearing a small accessory blunt branch on its central posterior third (Figs 25 A – C, 26 A, D – E). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 24 C, 25 A, C, 26 A, F, H). Tr IV rectangle-shaped (in dorsal view) (Figs 24 A – C, 25 A – C, 26 A, F – H). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (Figs 24 A – C, 26 A, F – H). Tr IV distal portion with a screwdriver tip-shaped transversal apophysis, dorsally covered with four tubercles, on prodorsal face (Figs 25 A, 26 A, F – G, I). Tr IV with two proximal and one distal subconical prominent tubercles on prolateral face (Fig. 26 F). Tr IV tuberculate on ventral face (Figs 24 C, 26 G – I). Tr IV distal portion with a subconical apophysis on retrolateral face (Figs 26 F, G – I). Fe IV sinuous, arched (1) on the proximal portion towards the prodorsal face and (2) on the distal portion towards the retroventral face (Figs 24 A – C, 26 F – I). Fe IV dorsal face with four spines on the proximal half and two spines on the distal third (Figs 26 F – G, I). Fe IV prodorsal face with a row of ordinary tubercles intercalated by two subconical spines on the central third (Fig. 26 F – G). Fe IV prolateral face with a row of tubercles divided in (1) ordinary tubercles on the proximal third, (2) prominent tubercles on the central third (c. 2.5 × – 3 × the ordinary ones), and (3) outstanding tubercles on the distal third (c. 3.5 × – 4 × the ordinary ones) (Fig. 26 F – H). Fe IV proventral face with (1) a row of ordinary tubercles on proximal and central thirds and (2) outstanding subconical tubercles interpolated by ordinary ones and a distalmost spine on the distal third (Fig. 26 G – H). Fe IV ventral face with a prominent tubercle on the proximal fifth (Fig. 26 H). Fe IV retroventral face with subconical outstanding tubercles on proximal and distal thirds and prominent tubercles on the central third (Fig. 26 H – I). Fe IV retrolateral face with six spines (the two distalmost larger than the Fe IV diameter) on proximal third and with a pair of outstanding spines (distalmost largest) on the central third (Fig. 26 F, H – I). Fe IV retrodorsal face with two outstanding subconical tubercles and three prominent tubercles on the proximal third and three conical spines on the distal third (Fig. 26 F, I). Pa IV dorsal face tuberculate (Fig. 26 F – G, I). Pa IV proventral and retroventral faces with rows of four and three spines, respectively (Fig. 26 G – I). Ti IV with all faces (except ventral face) containing longitudinal rows of acuminated tubercles (Fig. 26 F – I). Ti IV retrolateral face with three-four prominent subconical tubercles on proximal half (Fig. 26 F, H – I). Ti IV proventral and retroventral faces with spines on the distal half (the retroventral distalmost spine is bifurcated) (Fig. 26 F – I). Mt IV with all faces containing longitudinal rows of small subconical tubercles (Fig. 26 J). Mt IV with proventral and retroventral apical spurs (Fig. 26 J). Coloration (in vivo) (Fig. 24): DS and Cx – Tr IV background Blackish Green (152). Ocularium spines, scutal areas II – III and Cx IV prodorsal distal apophysis Brownish Black (65). Carapace and scutal areas I – IV with tubercles Brownish Orange (54). AS lateral margins with prominent tubercles Deep Brown (56). DS posterior margin and free tergites I – III with tubercles Dark Orange Yellow (72). Ch and Pp glossier background Moderate Yellow Green (120), with honeycombed Dark Grayish Olive Green (128) reticle. Tr I – III background Strong Yellowish Brown (74), with a dorsal distal semicircle Light Yellow (86). Fe – Pa I – III and Ti I – III proximal half background Dark Grayish Brown (62). Ti I – IV distal half background Dark Greenish Yellow (103). Fe – Pa IV and Ti IV proximal half background Reddish Black (24). Tr IV with apophyses and prominent tubercles Moderate Red (15). Fe IV with spines and prominent tubercles Deep Orange (51). Pa – Ti IV with spines and tubercles Deep Orange Yellow (69). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (1.5 × wider than distal part) (Fig. 27 A, C). VP ventral surface entirely covered with microsetae of type 1. All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated, on the basal half of the VP (Fig. 27 A – C). MS B 1 short, inserted ventrally, close to A 3 (Fig. 27 C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 27 A – C). MS D 1 short, closer to C 3 than A 1 (Fig. 27 A – C). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 between C 1 – C 2, E 2 between C 2 – C 3 (Fig. 27 B – C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 27 A – B). Stylus and its ventral process axis fused basally, forming a pedestal above the glans (Fig. 27 B). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 27 A – B). Stylus without any expansion or flattening, in situ reaching the distal margin of VP (Fig. 27 A – C). Ventral process sinuous, as long and thinner than the stylus (Fig. 27 A – B). Flabellum slightly bent ventrad, hand-shaped, with a comb of straight and acuminated spines on lateral faces (Fig. 27 B). Female (MNRJ 60562) (Figs 24 D – E, 25 D – F, 28 A – C): DS, measurements: CW 3.0, CL 2.3, AW 5.3, AL 3.4; Leg I – IV measurements in Table 15; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 10 (3) / 10 (3) - 7 / 7 - 7 / 7. Ocularium longitudinally reduced than males (Figs 25 D – E, 28 A). Scutal area III with a paramedian pair of conical spines (larger than males) (Figs 25 D – E, 28 B). Scutal area IV covered with ordinary tubercles (Figs 25 D – E, 28 B). Cx IV is narrower than males, with the prodorsal distal apophysis reduced to a single spine and a reduced retroventral distal apophysis (Figs 25 D – F, 28 B). Tr IV unarmed on prodorsal and prolateral faces (Fig. 28 B – C). Tr IV retrolateral face with a conical apophysis on proximal and distal thirds (distal largest), and a prominent subconical tubercle on central third (Fig. 28 C). Fe IV straight and thinner than males (Fig. 28 C). Fe IV dorsal face with two spines on proximal and distal halves (Fig. 28 C). Fe IV retrolateral face with (1) a prominent tubercle on proximal third; (2) an outstanding spine (largest than Fe IV diameter) on central third; and (3) two prominent tubercles (distalmost largest) on distal third (Fig. 28 C). Intraspecific variation: Some variations among the major morph males were detected: (1) Cx IV prodorsal distal apophysis length, with the main branch larger or reduced in comparison to holotype; (2) Fe IV dorsal row of spines with reduced spines; and (3) Fe IV slightly thinner and more extensive than the holotype. Among the minor morph males (Fig. 11 A) (compared to major morph): (1) DS narrower than males; (2) Cx IV distal apophysis reduced on prolateral and retrolateral faces; and (3) Fe and Ti IV thinner, with reduced spines. No relevant intraspecific variation among females was detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13030872E118BFA620869FEE5.taxon	type_taxon	Type species: Oliverius jordanensis Soares & Soares, 1945, by original designation.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13030872E118BFA620869FEE5.taxon	etymology	Etymology: As communicated by Soares and Soares (1945), the name was dedicated to the ornithologist Dr Olivério Mário de Oliveira Pinto.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C13030872E118BFA620869FEE5.taxon	diagnosis	Diagnosis: Oliverius can be distinguished from the other genera of Neopachylinae due to (1) DS outline of the males gamma-pyriform ‘ Gonyleptes - like’ sensu Kury and Medrano (2016) (Fig. 32 B); (2) Scutal area IV divided into left and right halves by a longitudinal median groove (Figs 31 A, 32 B); (3) Free tergites I – III with two or three pairs of paramedian conspicuous tubercles (Figs 31 A, 32 B); (4) Females’ free tergites I – III with a row of outstanding subconical tubercles (Figs 30 F, 31 D – E, 32 I); (5) Pedestal on the glans square-shaped (Fig. 33 B, D); 6) Ventral process of the glans as wide as the stylus (Fig. 33 B, D). Distribution: BRAZIL: states of Minas Gerais, Rio de Janeiro, and São Paulo (Fig. 29).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	description	(Figs 30 – 33)	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	materials_examined	Type data Discocyrtus granulatus: ♂ holotype (MZSP 4506, examined) from BRAZIL, Rio de Janeiro, Itatiaia, Macieiras. Oliverius jordanensis: ♂ holotype (MZSP 0821, examined) BRAZIL, São Paulo, Campos do Jordão. Records Without further literature records.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	diagnosis	Diagnosis As presented for the genus.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	materials_examined	Non-type material examined BRAZIL: state of Minas Gerais: Delfim Moreira: 1 ♂ 2 ♀ (MZSP 29845), [- 22.5092 °, - 45.28 °], 01. xi. 2004, Pinto-da-Rocha, R. & daSilva, M. B. leg. Lima Duarte: 2 ♂ 2 ♀ (MNRJ 8604)!, Parque Estadual do Ibitipoca, 5 - 8. v. 2008, Oliveira, I. S. leg. Itamonte: 2 ♂ (ISLA 12817)!, x. 2009; 1 ♂ 1 ♀ (MNRJ 2238)!, Aiuruoca, 31. x. 2007, Sampaio, C. leg.; 1 ♂ (MNRJ 5599)!, 7 ♂ 2 ♀ (MNRJ 5602)!, Parque Nacional de Itatiaia, Brejo da Lapa, - 22.362 °, - 44.735 °, 2200 m, 05 – 06. ii. 1997, Kury, A. B. et al. leg.; 5 ♂ 6 ♀ (MNRJ 58980), idem, road to Brejo da Lapa, - 22,375 941 °, - 44,748 277 °, 1800 m, 27. xii. 2019, Kury, A. B. et al. leg.; 1 ♀ (UERJ A 0425), 1 ♂ (UERJ A 0426), idem, Parte Alta, road near to the Casa do Pesquisador, 05 – 08. xii. 2019, Corrêa, C. C. D. leg. State of Rio de Janeiro: Itatiaia: 1 ♂ 1 ♀ (MNRJ 58998), Parque Nacional de Itatiaia, Abrigo Lamego / Piscina do Maromba, - 22.42 995 °, - 44.62 216 °, 2000 m, 29. xii. 2019, Kury, A. B. et al. leg.; 2 ♂ (UERJ A 0043), idem, caminho da Casa de Pedra, 25. ix. 2018, Equipe Diptera leg.; 1 ♂ 1 ♀ 1 juv (UERJ A 0074), idem, Casa de Pedra, 24. ix. 2018, Equipe Diptera leg.; 1 ♂ 1 ♀ (MNRJ 5561)!, idem, Véu da Noiva, Córrego Maromba, - 22.4361 °, - 44.6244 °, 1100 m, 02 - 03. ii. 1997, Kury, A. B. et al. leg. State of São Paulo: São José do Barreiro: 1 ♂ (IBSP 2774)!, Parque Nacional da Serra da Bocaina, 28. iv – 03. v. 2002, Equipe Biota leg.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	distribution	Distribution (new records with an asterisk) BRAZIL: state of Minas Gerais: Delfim Moreira *, Itamonte *, Lima Duarte *. State of Rio de Janeiro: Itatiaia. State of São Paulo: Campos do Jordão; São José do Barreiro * (Fig. 29).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303187291285FEB60C9AFBF9.taxon	description	Redescription MZSP 29845 (male) for the external body illustrations and description; DS, measurements: CW 3.4, CL 2.6, AW 7.1, AL 3.2; Leg I – IV measurements inTable 16; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - x / 12 (3) - x / 7 - 7 / 7. MNRJ 5561! (male) for genitalic illustrations (Figs 33 A – E). Dorsum: DSgamma, moreextensivethanlong, withASlateralmargins strongly convex, widest at scutal area II and thickest at scutal area III, with sinuous posterior margin (Figs 31 A – B, 32 B – C). DS anterior margin divided by a small central projection in the centre and a pair of shallow cheliceral sockets (Fig. 32 B). Carapace anterior third with two sets of prominent subconical tubercles on two transversal rows (five tubercles each on anterior set, four tubercles each on posterior set), divided by a central portion covered by ordinary tubercles (Fig. 32 B). Carapace posterior two-thirds with two paramedian pairs of outstanding tubercles, surrounded by ordinary and prominent tubercles on lateral and posterior portions (Figs 31 A – B, 32 B – C). Ocularium elliptical (in dorsal view), high (c. 3 × the eye diameter), perpendicularly placed on the middle of the carapace (Figs 31 A – B, 32 A – C). Ocularium with a pair of sub-parallel spines (c. 2.5 × the eye diameter) inclined frontwards (Figs 31 A – B, 32 A – C). AS lateral margins with two rows of tubercles: one external, composed of three-six outstanding subconical tubercles at areas I – IV; another internal one with ordinary tubercles from the anterior corner of the carapace to the DS posterior margin (Fig. 32 B – C). Mesotergum divided into four clearly defined areas (Figs 31 A, 32 B). All scutal areas tuberculate (Figs 31 A – B, 32 B – C). Scutal areas I and IV divided into left and right halves by a longitudinal median groove (Fig. 32 B). Scutal area I with a pair of paramedian outstanding tubercles (c. 7 × the ordinary tubercles), placed between two other prominent tubercles (c. 1.5 × the ordinary tubercles) (Figs 30 A – B, 31 B, 32 B – C). Scutal areas II and IV with two pairs of paramedian outstanding tubercles (c. 2 × the ordinary tubercles) (Figs 31 A – B, 32 B – C). Scutal area II posterior-lateral margin embracing the scutal area III (Figs 30 B, 32 B). Scutal area III with a pair of remarkable cylindrical structures with a broad base, slightly bent to posterior (c. 6.5 × the ordinary tubercles) (Figs 30 A – C, 31 A – B, 32 B – C). Scutal area III with two pairs of outstanding tubercles on the medial portion (Figs 30 B, 31 A, 32 B). DS posterior border and free tergites I – III each with a transversal row of prominent and rounded tubercles (Figs 30 C, 31 A, 32 B). Anal operculum tuberculate (Fig. 30). Venter: Cx I – III sub-parallel to each other, individually with ventral longitudinal rows of 9 – 11 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (Figs 30 D, 31 C). Cx II with a retroventral distal row of four acuminated tubercles. Cx III with a retroventral distal row of 12 acuminated tubercles. Cx IV much larger than the others, directed obliquely (Figs 30 D, 31 C). Intercoxal bridges are well marked (Figs 30 D, 31 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Figs 30 D, 31 C). Cx IV covered by ordinary tubercles (Figs 30 D, 31 C). Stigmata are visible (Figs 30 D, 31 C). Free sternites with a transverse row of ordinary tubercles (Figs 30 C – D, 31 C). Chelicera: Basichelicerite elongate, bulla well marked (Figs 31 A, 32 B), with marginal setiferous tubercles — two lateral ectal, one posterior (Fig. 32 B); hand not swollen. Pedipalps: Tr ventral with a geminate setiferous tubercle. Fe with a ventral basal and a mesal distal setiferous tubercle. Pa unarmed (Fig. 31 A). Ti with a row of four spines (IiIi) on ventro-mesal and ventro-ectal faces. Ta with a row of three spines (III) on ventro-mesal and ventro-ectal faces. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles (Fig. 30 D). Fe I – II straight (Figs 30 A – B, 31 A). Fe and Ti I – II with all faces containing rows of small tubercles (Fig. 30 B). Fe III sub-straight (Fig. 30 D). Fe III andTiIIIwithprodorsal, proventral, retroventral, andretrodorsal rows of acuminated tubercles (Fig. 30 A – B, D). Pa III dorsally covered by acuminated tubercles (Fig. 30 A). Cx IV reaching the anal operculum (Figs 30 B, D, 31 A, 32 B). Cx IV tuberculate between prodorsal and ventral faces (Figs 30 A – D, 31 A – C, 32 B). Cx IV with a prodorsal distal subconical apophysis (elongate, subapically bent to posterior), bearing a small accessory blunt branch (not totally visible in dorsal view) (Figs 31 A – C, 32 B). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (Figs 30 B – D, 31 A, C, 32 B). Tr IV rectangle-shaped (in dorsal view) (Figs 30 B, D, 31 A – C, 32 B). Tr IV central portion with a prominent tubercle on the dorsal and retrodorsal faces (Figs 30 B, 32 B, D). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (both slightly bent dorsad) (Figs 31 A – B, 32 B). Tr IV distal portion with a hook-shaped apophysis on prodorsal face (Figs 31 A – B, 32 B, D – E). Tr IV tuberculate on ventral face (Figs 30 D, 31 B – C, 32 E – G). Tr IV distal portion with a reduced conical apophysis on retrolateral face (Figs 31 C, 32 B, D, F – G). Fe IV sub-straight, arched on the distal half towards dorsal and prolateral faces (Fig. 32 D – G). Fe IV dorsal face with a row of one proximal-most conical spine (I) and eight conical tubercles (Fig. 32 D – E, G). Fe IV prodorsal face with a row of 20 ordinary tubercles (Fig. 32 D – E). Fe IV prolateral face with a row of 18 slightly prominent acuminated tubercles (Fig. 32 D – F). Fe IV proventral face with a row of 22 prominent tubercles (Fig. 32 E – F). Fe IV retroventral face with a row of 16 outstanding tubercles except the four proximal-most and the two distalmost, which are prominent tubercles (Fig. 32 E – F). Fe IV retrolateral face with a row of three subconical spines (iiI) on the proximal half, and a subconical spine (I) on the distal half (all of them bent dorsad) (Fig. 32 D, F – G). Fe IV retrodorsal face with a row of 18 ordinary tubercles (Fig. 32 D, G). Fe IV apical portion with subconical spurs on prodorsal and retrodorsal faces (retrodorsal largest) (Fig. 32 D – E, G). Pa IV tuberculate on dorsal view (Fig. 32 D – E, G). Pa IV with a row of three spines on proventral and retroventral faces (Fig. 32 E, G). Ti IV with all faces (except the ventral) containing longitudinal rows of acuminated tubercles (Fig. 32 E, G). Mt IV with all faces containing longitudinal rows of small-acuminated tubercles. Mt IV with proventral and retroventral apical spurs. Coloration (in vivo) (Fig. 30): Carapace and DS anterior and lateral margins background Dark Grayish Red (20). Mesotergum background Dark Bluish Gray (192). DS posterior border and free tergites I – III background Dark Greenish Yellow (103). Spines on the ocularium and scutal area III Dark Grayish Reddish Brown (47). DS with tubercles Deep Orange Yellow (69) or Strong Yellowish Brown (74). AS lateral borders with prominent subconical tubercles Deep Reddish Brown (41). DS posterior margin and free tergites I – III with a row of tubercles varying between Vivid Orange (48) and Strong Reddish Brown (40). Ch and Pp glossier background Blackish Green (152), with setiferous tubercles and claw Brilliant Yellow Green (116) and honey-combed details Moderate Yellow Green (120). Tr I – III and Fe I – III proximal half background Dark Yellowish Brown (78). Fe I – III distal half, Pa – Mt I – IV, and Cx IV background Dark Greenish Gray (156). Tr – Fe IV background Dark Reddish Brown (44). Legs I – III with ordinary tubercles Deep Orange Yellow (69). Leg IV with tubercles, spines, and apophyses varying between Vivid Orange (48) and Deep Reddish Brown (41). Coloration (in ethanol) (Fig. 31): DS background and margins vary between Moderate Olive Brown (95) and Dark Grayish Yellowish Brown (81), with a reticulum Strong Yellow (84). DS tubercles and ocularium spines Moderate Yellow Green (120). Scutal area III with spines Dark Grayish Brown (62). Ch and Pp background Olive Black (114), with setiferous tubercles and claw Strong Yellow Green (117) and honey-combed details Brilliant Greenish Yellow (98). Legs I – III background Dark Olive (108) with details Strong Greenish Yellow (99). Leg IV background Moderate Olive Brown (95), with honey-combed details on Cx IV prolateral proximal portion Moderate Greenish Yellow (102). Fe IV distal third background Strong Greenish Yellow (99). Pa – Mt distal third background Brilliant Yellow Green (116). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (Fig. 33 A, C). VP ventral surface entirely covered with microsetae of type 1 (Fig. 33 B – C). All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 3 thick and acuminated, forming a longitudinal row on the basal third of VP (Fig. 33 A – B). MS B 1 short, inserted ventrally, close to A 3 (Fig. 33 B – C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 33 A – C). MS D 1 short, closer to C 3 than A 1 (Fig. 33 A – C). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 above C 1, E 2 beside C 2 (Fig. 33 B – C). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 33 A – B). Stylus and its ventral process axis fused basally, forming a square-shaped short pedestal above the glans (Fig. 33 A – B, D – E). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 33 A – B, D – E). Stylus without any expansion or flattening, in situ reaching the distal margin of VP (Fig. 33 B – C). Ventral process almost straight, as long and thinner than the stylus (Fig. 33 B, D – E). Flabellum bent 90 ° ventrad, hand-shaped (with short serrulations and spines) (Fig. 33 A – B, D – E). Female (MZSP 29845) (Figs 30 F, 31 D – F, 32 H – J): DS, measurements: CW 3.4, CL 2.5, AW 6.2, AL 3.6; Leg I – IV measurements in Table 17; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 10 (3) / 10 (3) - 7 / 7 - 7 / 7. Ocularium with reduced spines compared to males (Figs 31 B, 32 H). AS margins are less concave compared to males (Figs 30 F, 31 B, 32 J). DS posterior margins and free tergites I – III with a row of outstanding acuminated tubercles (Figs 30 F, 31 D – E, 32 I). Scutal area III with a pair of paramedian conical spines (with a rounded apex) (Figs 30 F, 31 D – E). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine (Figs 31 D, 32 J). Fe IV thinner and less curved in comparison to males (Fig. 31 D, F). Fe IV, Tr IV, and Ti IV with reduced armature in comparison to males. Intraspecific variation: Some variations between minor morph males (Fig. 30 E) and major morph males were detected: (1) DS with less developed armature; (2) Cx IV distal portion with reduced apophysis on prolateral and retrolateral faces; and (3) Fe IV thinner, with less developed armature. No relevant intraspecific variation among major morph males and among females was detected in the material studied. Historical taxonomical remarks: The male holotype of Discocyrtus granulatus (MZSP 4506) matches almost 100 % with the morphology of Oliverius jordanensis holotype (MZSP 0821). No additional specimens were recorded in the literature for both species. The type-locality of Oliverius jordanensis (Campos do Jordão, São Paulo) and Discocyrtus granulatus (Itatiaia, Rio de Janeiro) are part of the Atlantic province (Fig. 29). The same occurs with almost all the ordinary specimen records studied here (except Delfim Moreira, Minas Gerais, which belongs to the Paraná Forest province). Therefore, Discocyrtus granulatus is here considered a junior subjective synonym of Oliverius jordanensis.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130368728165FFBA10BFCFA1A.taxon	type_taxon	Type species: Pachylobos areolatus Piza, 1940, by original designation.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130368728165FFBA10BFCFA1A.taxon	diagnosis	Diagnosis: See the original diagnosis in Carvalho and Kury (2020: 88). An updated version, following the results obtained here, as follows: Pachylobos can be distinguished from the other genera of Neopachylinae due to: (1) Ventral process of the glans sigmoid (Carvalho and Kury 2020: fig. 15 D); (2) Outline of the male’s DS type lambda sensu Kury and Medrano (2016) (Figs 35 A, 36 B); (3) Mesotergum with an areolate pattern of stains forming circles around of the tubercles (Carvalho and Kury 2020: figs 13 A, 35 A, 36 B); (4) Fe IV retrolateral face with a spine comb growing distally in size on the proximal third (Figs 36 F, 36 H); (5) Fe IV retrolateral face with a pair of intervaled spines (with almost the same size) on the central third (Fig. 36 F, H); (6) Tr III retrolateral face of the females with the distal portion unarmed (Fig. 38 B).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130368728165FFBA10BFCFA1A.taxon	distribution	Distribution: BRAZIL: states of Paraná and São Paulo (Fig. 34).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	description	(Figs 35 – 38) ZooBank LSID: urn: lsid: zoobank. org: act: 96 D 322 FE-E 209 - 4334 - 882 B- 0 D 31 E 66 A 5 F 54.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	etymology	Etymology The species name is an adjective from the type locality.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	materials_examined	Type data Pachylobos iratiensis: ♂ holotype 2 ♀ paratypes (MNRJ 363), from BRAZIL, state of Paraná, Fernandes Pinheiro, Floresta Nacional de Irati.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	diagnosis	Diagnosis Pachylobos iratiensis: can be distinguished from P. littoralis and P. longicornis due to: (1) Scutal area III with a pair of spines with subconical apex (Fig. 36 B, D); (2) Cx IV prodorsal distal apophysis with half of the AW size (Fig. 36 B); (3) Fe IV dorsal face with a comb of spines (IIII) on the proximal half (Fig. 36 F – G, I); (4) Fe IV retrodorsal face with a row of tubercles (as in P. littoralis) and a distalmost outstanding spine (as in P. longicornis) (Fig. 36 F, I); (5) VP with proximal half elliptical, 1.5 × wider than distal part (Fig. 37 A, C); (6) MS B 1 evident, with the central portion visible (Fig. 37 C); (7) Females’ free tergites I – III with a pair of outstanding subconical tubercles (Fig. 38 B).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	distribution	Distribution BRAZIL: state of Paraná: Fernandes Pinheiro (Fig. 34).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C130378723134BFA770B3DFB5C.taxon	description	Redescription MNRJ 363 (male) for the external body illustrations and description; DS, measurements: CW 3.3, CL 2.3, AW 6.4, AL 3.1; Leg I – IV measurements inTable 18; Right / left tarsal (distitarsal) counts: 6 (3) / 6 (3) - 13 (3) / 8 (3) - 7 / 7 - 7 / 7. MNRJ 363 (male) for genitalic illustrations. Dorsum: DS lambda, wider than long, with AS lateral margins strongly convex, widest at scutal area II and thickest at scutal area III (Figs 35 A – B, 36 B, D). AS with sinuous posterior margin (Fig. 36 B, D). DS anterior margin divided by a small central projection in the centre and a pair of shallow cheliceral sockets (Figs 35 A, 36 B). Carapace anterior portion with two transversal rows of four – five prominent subconical tubercles, and centrally covered by ordinary tubercles (Fig. 36 B). Carapace with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (Fig. 36 B, D). Ocularium elliptical (in dorsal view), slightly inclined frontwards (in lateral view), high (c. 3 × the eye diameter), perpendicularly placed on the central portion of the carapace (Figs 35 A – B, 36 A – B, D). Ocularium with a pair of almost parallel spines (c. 3 × the eye diameter) fused at baseline and slightly inclined frontwards (Figs 35 A – B, 36 A – B, D). AS lateral borders with two rows of tubercles, one external, composed of three – four prominent subconical tubercles at areas I – IV; another internal one with ordinary tubercles from the posterior corner of the carapace to the DS posterior margin (Figs 35 A – B, 36 B, D). Mesotergum is divided into four clearly defined scutal areas (Figs 35 A, 36 B). All scutal areas tuberculate, with almost all tubercles individually covered / surrounded by light-coloured spots (Figs 35 A, 36 B). Scutal area I divided into left and right halves by a longitudinal median groove (Figs 35 A, 36 B). Scutal areas I – II and IV with two or three pairs of prominent tubercles (c. 1.5 × the ordinary tubercles) (Figs 35 A, 36 B, D). Scutal area II with anterior-lateral margin slightly embracing scutal area I, and with posterior-lateral margin embracing scutal area III (Figs 35 A, 36 B). Scutal area III with a pair of paramedian subconical spines with rounded apex (c. 7 × the ordinary tubercles) (Figs 35 A – B, 36 B – D). Scutal area III with posterior-central margin slightly invading scutal area IV (on paramedian portion) (Figs 35 A, 36 B). DS posterior border and free tergites I – III each with a transversal row of tubercles (growing in size towards central portion, with six prominent ones) (Figs 35 A, 36 B, D). Anal operculum tuberculate. Venter: Cx I – III sub-parallel to each other (Fig. 35 C), individually with ventral longitudinal rows of 8 – 15 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others). Cx II with two acuminated tubercles on retroventral distal. Cx III with a retroventral distal row of nine acuminate tubercles. Cx IV much larger than the others, directed obliquely (Fig. 35 C). Intercoxal bridges are well marked (Fig. 35 C). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (Fig. 35 C). Cx IV covered by ordinary tubercles (Fig. 35 C). Stigmata are visible (Fig. 35 C). Free sternites with a transverse row of ordinary tubercles (Fig. 35 C). Chelicera: Basichelicerite elongate, bulla well marked (Fig. 36 B), with marginal setiferous tubercles — two lateral mesal, two or three posteriors, three lateral ectal (Fig. 36 B); hand not swollen. Pedipalps: Tr with two geminated ventral setiferous tubercles. Fe with a ventral basal and a mesal distal setiferous tubercle. Pa unarmed. Ti with a row of four spines (IiIi) on ventro-mesal and ventro-ectal faces. Ta with two rows of spines — three (IIi) ventro-mesal and four (IiIi) ventro-ectal. Legs: All the unmentioned podomeres are unarmed or without relevant armature. Cx I – II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I – III each with several ventral tubercles. Fe I and III sub straight (Figs 35 C, 36 E); Fe II straight (Fig. 35 C). Fe and Ti I – III with all faces containing rows of small tubercles (Fig. 36 E). Fe III with a reduced apical retrodorsal spur (Figs 35 A, 36 E). Pa I – III dorsally covered by small tubercles. Cx IV reaches the DS posterior margin (Fig. 36 B). Cx IV tuberculate between prodorsal and ventral faces (Fig. 36 B). Cx IV prodorsal distal apophysis with a conical main branch (as long as the AS’ half-width, centrally bent to posterior) (Fig. 36 B, D, F – H). Cx IV retroventral distal apophysis with a blunt main branch, with a small secondary branch on its proximal half (Fig. 36 B, F, H – I). Tr IV rectangle-shaped (in dorsal view) (Fig. 36 B, F – I). Tr IV central portion with a prominent tubercle on dorsal face (Fig. 36 B, F – G). Tr IV distal portion with a hook-shaped apophysis on prodorsal face (Fig. 36 B, F – G, I). Tr IV proximal portion with a conical apophysis (slightly bent dorsad) on prolateral face (Fig. 36 B, F, H). Tr IV tuberculate on ventral face (Fig. 36 G – I). Tr IV central portion with a conical apophysis (slightly bent dorsad) and two prominent subconical tubercles on retrolateral face (Fig. 36 B, H – I). Tr IV distal portion with a subconical apophysis on retrolateral and retrodorsal faces (Fig. 36 B, F, H – I). Fe IV sub-straight, arched (1) on the proximal half towards the prodorsal face and (2) on the distal half towards the retroventral face (Fig. 36 F – I). Fe IV dorsal face with a row of seven spines (IIIIIiI) bent to retrolateral, (except the proximal-most and the two distalmost, which are straight) (Fig. 36 F – I). Fe IV prodorsal face with a row of ordinary tubercles and a prominent apical tubercle (Fig. 36 F – G). Fe IV prolateral face with a row of 13 outstanding subconical tubercles (Fig. 36 F – H). Fe IV proventral face with a row of five spines (iiiII) on distal half (Fig. 36 G – H). Fe IV ventral face with a subconical prominent tubercle on the proximal fifth (Fig. 36 H). Fe IV retroventral face with a row of eight spines (iiiiiIII) — the five proximal-most subconical, the three distalmost acuminated (Fig. 36 F – G, I). Fe IV retrolateral face with a row of seven spines (iiIIIii), with a diastema on the central portion between the four proximal-most and the five distalmost (Fig. 36 F, H – I). Fe IV retrodorsal face with ordinary tubercles on the proximal and central thirds, a spine on the distal third and an outstanding apical spur (Fig. 36 F, I). Pa IV dorsally covered by prominent acuminated tubercles (some are outstanding, mainly on dorsal and retrodorsal faces) (Fig. 36 F, I). Pa IV proventral and retroventral faces with rows of five and two spines, respectively (Fig. 36 G – I). Ti IV with all faces containing longitudinal rows of spines (except ventral; proventral, retroventral and retrolateral largest) (Fig. 36 H – I). Ti IV with a sizeable spur on proventral and retroventral apical portions (Fig. 36 H – I). Mt IV with all faces containing longitudinal rows of small tubercles (Fig. 36 J). Mt IV with a reduced spur on proventral and retroventral apical portions (Fig. 36 J). Coloration (in ethanol) (Fig. 35): DS background, free tergites I – III and Cx I – IV Dark Grayish Yellowish Brown (81). Mesotergum grooves between areas I – IV Light Olive Gray (112). Tubercles surrounded by lighter circles Grayish Greenish Yellow (105) on mesotergum, AS lateral portions and free tergites I – III. Spines on scutal area III and Cx IV prodorsal distal apophysis Brownish Black (65). Ch glossier background Dark Grayish Yellow (91), with honeycombed Dark Grayish Olive (111) reticle. Pp glossier background Very Pale Green (148), with honeycombed Greenish Gray (155) reticle. Tr I – IV background in a combination of Grayish Greenish Yellow (105) and Dark Grayish Yellowish Brown (81), with a dorsal semicircle Very Light Yellowish Green (134) on distal half. Fe – Mt I – III background Grayish Greenish Yellow (105), with honeycombed Dark Olive Brown (96) reticle. Fe – Mt IV background Dark Grayish Olive (111), with apical portions of tubercles and spines Moderate Greenish Yellow (102) or Moderate Yellow Green (120). Male genitalia: VP slightly divided into a distal half forming a trapezium (widest at the base) with latero-apical flaps and a proximal half elliptical (1.5 × wider than distal part) (Fig. 37 A, C). VP ventral surface entirely covered with microsetae of type 1. All macrosetae cylindrical, inserted on lateral of VP. MS A 1 – A 4 thick and acuminated, on the basal half of the VP (Fig. 37 A – C). MS B 1 short, inserted ventrally, close to A 4 (Fig. 37 B – C). MS C 1 – C 3 thick and acuminated, forming a longitudinal row on the distal half of VP (Fig. 37 A – C). MS D 1 short, closer to C 3 than A 1 (Fig. 37 A – B). MS E 1 – E 2 very reduced, located on the laterodistal flange of VP — E 1 above C 1, E 2 between C 2 – C 3 (Fig. 37 B). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (Fig. 37 A – B, D). Stylus and its ventral process fused basally, forming a pedestal above the glans (Fig. 37 B, D). Stylus cylindrical, bent at the subdistal part (forming a plateau) and armed with a set of ventral subapical spines (Fig. 37 A – D). Stylus without any expansion or flattening, in situ reaching the distal margin of VP. Ventral process bent dorsad, as long and thinner than the stylus (Fig. 37 A, D). Flabellum bent 90 ° ventrad, hand-shaped (with more than 11 lateral prominent spines, some of them bifurcated) (Fig. 37 B, D). Female (MNRJ 363) (Figs 35 D – F, 38): DS, measurements: CW 2.9, CL 2.1, AW 5.3, AL 3.6; Leg I – IV measurements in Table 19; Right / left tarsal (distitarsal) counts: 5 (2) / 6 (3) - 9 (3) / 9 (3) - 7 / 7 - 7 / 7. Ocularium reduced compared to males (Figs 35 E, 38 A). DS lambda, with AS margins are less concave than males (Figs 35 D, 36 B). DS posterior margin and free tergites I – III with a pair of outstanding subconical tubercles (Figs 35 D, 38 B). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine (Figs 35 D – E, 38 B – C). Tr IV distal portion unarmed on prodorsal and retrodorsal faces (Figs 35 D – E, 38 C). Tr IV distal portion with a prominent subconical tubercle on prolateral face (Fig. 38 C). Fe IV sub-straight, thinner compared to males, arched on the proximal half towards prodorsal face (Figs 35 D – F, 38 C). Fe IV with five small acuminated spines on dorsal face (Fig. 38 C). Fe IV retrolateral face with a prominent subconical tubercle on the proximal fifth and two outstanding spines on the distal half (Fig. 38 C). Intraspecific variation: No other male specimen, except the holotype, has been studied here. No intraspecific variation among females was detected in the material studied.	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872312BCFAC30A83F9CA.taxon	materials_examined	Type data Discocyrtus littoralis: ♂ holotype (no identified depository, lost) from BRAZIL, São Paulo, São Sebastião. Discocyrtus cornutus: ♀ holotype (MZSP 54, examined) from BRAZIL, São Paulo, Serra da Cantareira. Pachylobos areolatus: 2 ♂ syntypes (MZSP and IBSP, examined by photograph in original description) from BRAZIL, São Paulo, Serra da Cantareira. Discocyrtulosoma soaresi: ♂ ♀ syntypes (MZSP 103, examined), ♂ syntype (MZLQ, not examined) from BRAZIL, São Paulo, Butantan / Serra da Cantareira. Records BRAZIL: state of São Paulo: Ubatuba (B. Soares 1943); Alto da Serra, Mogi das Cruzes, Perus, São Paulo (B. Soares 1944 b, 1945 c, 1946); Ilha dos Búzios, Ilha da Vitória (H. Soares 1966); Campos do Jordão, Cocaia, Guarulhos, Ilha da Queimada Grande, Jundiaí, Juquitiba, Mairiporã, Miracatu, São Francisco Xavier (Carvalho and Kury 2020). Unverified / mistaken records ARGENTINA: Misiones: Santa María (Ringuelet 1959).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872312BCFAC30A83F9CA.taxon	diagnosis	Diagnosis The diagnosis provided here is an updated version of the one presented in Carvalho and Kury (2020: 94), as follows: P. littoralis can be distinguished from P. iratiensis and P. longicornis due to: (1) Ocularium with pair of parallel and independent armature (as in P. iratiensis) (Carvalho and Kury 2020: fig. 9 B); (2) Scutal area III with a pair of paramedian domes (Carvalho and Kury 2020: figs 8 A, C, 9 A, C); (3) Fe II unarmed on retrodorsal distal face; (4) Cx IV prodorsal apophysis with less of the AW half-size (Carvalho and Kury 2020: figs 8 A, 9 A); (5) Fe IV dorsal face with a comb of spines (iiI) on the proximal half (Carvalho and Kury 2020: figs 9 D – E, G); (6) Fe IV armed with a tiny spine on retrodorsal distal face (Carvalho and Kury 2020: figs 8 A, 9 D – E); (7) DS posterior margin and free tergites I – III of females with a row of outstanding acuminated tubercles (as in P. longicornis) (Carvalho and Kury 2020: fig. 8 D).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872312BCFAC30A83F9CA.taxon	distribution	Distribution BRAZIL: state of São Paulo: Alto da Serra, Campos do Jordão, Cocaia, Guarulhos, Ilha da Queimada Grande, Ilha da Vitória, Ilha dos Búzios, Jundiaí, Juquitiba, Mairiporã; Miracatu, Perus, São Francisco Xavier, São Paulo and Ubatuba (Fig. 34).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872312BCFAC30A83F9CA.taxon	description	Redescription See the detailed redescription in Carvalho and Kury (2020: 88 – 98).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872211C9F94209DEF9F3.taxon	materials_examined	Type data Discocyrtus guttatus: ♂ holotype (SMF 1437, examined by photograph) from BRAZIL, São Paulo, Ribeirão Pires. Discocyrtus transversalis: ♂ ♀ syntypes (MZSP 858, examined by photograph) from BRAZIL, São Paulo, Alto da Serra. Gonyleptes longicornis: 2 ♂ syntypes (MZSP 1563, examined) from BRAZIL, São Paulo, Alto da Serra. Records BRAZIL: state of São Paulo: Bertioga, Boracéia, Iguape, Ilhabela, Itanhaém, Miracatu, Paranapiacaba, Peruíbe, Salesópolis, Santo André, São Paulo, Sete Barras (Carvalho and Kury 2020).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872211C9F94209DEF9F3.taxon	diagnosis	Diagnosis The diagnosis provided here is an updated version of the one presented in Carvalho and Kury (2020: 99), as follows: P. longicornis can be distinguished from P. iratiensis and P. littoralis due to: (1) Ocularium with a pair of divergent and fused at base spines (Carvalho and Kury 2020: fig. 14 B); (2) Scutal area III with a pair of paramedian cones with rounded apex (Carvalho and Kury 2020: fig. 14 A, C); (3) Fe II retrodorsal distal with a spur; (4) Fe IV dorsal face with a comb of spines (IiiI) on the proximal half (Carvalho and Kury 2020: fig. 14 A); (5) Fe IV armed with an outstanding spine on retrodorsal distal face (Carvalho and Kury 2020: fig. 14 D, G); (6) Fe IV of females straight (Carvalho and Kury 2020: fig. 13 D – E); (7) DS posterior margin and free tergites I – III of females with a row of outstanding acuminated tubercles (as in P. littoralis) (Carvalho and Kury 2020: fig. 8 D).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872211C9F94209DEF9F3.taxon	distribution	Distribution BRAZIL: state of São Paulo: Alto da Serra, Bertioga, Iguape, Ilhabela, Itanhaém, Miracatu, Peruíbe, Ribeirão Pires, Salesópolis, Santo André, São Paulo, Sete Barras (Fig. 34).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
03A387C1303C872211C9F94209DEF9F3.taxon	description	Redescription See the detailed redescription in Carvalho and Kury (2020: 98 – 102).	en	Carvalho, Rafael N., Kury, Adriano B. (2025): Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species. Zoological Journal of the Linnean Society (zlae 023) 203 (1): 1-65, DOI: 10.1093/zoolinnean/zlae023, URL: https://doi.org/10.1093/zoolinnean/zlae023
