identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A287D5D803FFFA7A68FEFC1D4862DB.text	03A287D5D803FFFA7A68FEFC1D4862DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pectinodrilus ningaloo Pinder & Eberhard & Humphreys 2006	<div><p>Pectinodrilus ningaloo Pinder n. sp. (Fig. 2)</p> <p>Holotype. WAM V 4442. Specimen whole­mounted under right coverslip on same slide as paratype WAM V 4443, from groundwater in Ningaloo Homestead Well, about 500 m from the ocean, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.683334&amp;materialsCitation.latitude=-22.7" title="Search Plazi for locations around (long 113.683334/lat -22.7)">Ningaloo Station</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.683334&amp;materialsCitation.latitude=-22.7" title="Search Plazi for locations around (long 113.683334/lat -22.7)">Cape Range</a>, Western Australia (Fig. 1), 22º42’S 113º41’E, 11 June 1993. Temperature 22.8°C, conductivity 0.66 mS cm ­1, total dissolved solids 360 mg L ­1. Coll. R. D. Brooks, WAM collection # BES 2230).</p> <p>Paratypes. WAM V 4443 and 4444. One mature with head and tail missing, wholemounted under left coverslip on same slide as holotype and 1 mature with tail missing, whole­mounted on separate slide, collection details as for holotype.</p> <p>Other material. WAM V 4445. Two immature of uncertain identity in alcohol, collection details as for holotype.</p> <p>Description. Length and number of segments of holotype 8.2 mm and 61 respectively, maximum width of slide­mounted worms 0.14–0.20 mm at segment XI. Clitellum from posterior 1/3 of X to 12/13, more glandular, but not thicker, than epidermis of pre­clitellar segments. Post­clitellar segments with epidermis thinner than anterior segments. Male pores ventro­lateral on XI about 4/5 distance between 10/11 and 11/12. Spermathecal pores ventro­lateral immediately behind 9/10. Female funnels ventro­lateral at 11/12.</p> <p>Prostomium elongate oval, length:height at base 1.5, with large round clusters of cells with nuclei on the periphery, lying beneath the epidermis and projecting into the coelom. Pharynx in II/III. Pharyngeal glands, consisting of large irregular cells, mostly lateral and dorsal to the gut in IV–VI. Rest of ciliated digestive tract of variable width but not enlarging in pre­genital segments. Large dorsal and smaller ventral blood vessels connected by commissural vessels in II–V and by a plexus of capillaries surrounding the gut from VI. From VI or VII onwards, gut surrounded dorsally and laterally by large chlorogogue cells with non­staining cytoplasm occupying up to half of the coelom in many segments. Coelomocytes not observed. Chaetae 3(4–5)/bundle anteriorly, reduced to 2 posteriorly, 32–40 µm long x 1.5–1.8 µm wide at nodulus, bifid with sharp teeth, the upper tooth about half as long as lower, the nodulus distinctly distal. Straight penial chaetae 10–19/bundle in XI, protruding from body wall on a rounded papilla anterior to the male pores. Penial chaetae 32–36 µm long x ~ 1–1.5 µm wide at the base, tapering slightly towards their tips which appear to be slightly swollen and minutely hooked, but tips difficult to see as chaetae facing outwards.</p> <p>Genitalia paired. Testes antero­ventral in X with sperm sacs from IX to XIII. Ovaries antero­ventral in XI with egg sacs extending to XIV. Asymmetrical sperm funnels ventrolateral on 9/10. Vasa deferentia ciliated and tripartite: a short, thin (7–8 µm wide) ental section with thin muscle layer, a longer, thicker (maximum width 20–27 µm) middle section with thick (2–3 µm) muscle layer and thicker lining tissue and an ectal section (11–17 µm wide) with thinner muscle layer and lining tissue intermediate in width between the ental and middle sections. Atria short (50–58 µm), upright and moderately muscular, leading to simple male pore in a shallow invagination (at least in preserved specimens) behind penial chaetae. Two small ovoid prostates associated with each set of male ducts, one ventral to the middle part of vasa deferentia and one posterior to atrium, connections with the male ducts not seen. Spermathecae variable in shape, size and expansion of the lumen, but ampullae about 3 times longer than maximum width (about 50 µm in 2 mated specimens), with a short constriction between the ampulla and a small ectal vestibule. Sperm in loose masses in ampullae.</p> <p>Remarks. Members of the genus Pectinodrilus are united by the presence of small upright atria and the large number of penial chaetae, both characters that are clearly present in the new species. All previously described Pectinodrilus are marine meiobenthic and most have simple male ducts with a thin vasa deferentia leading directly into the atria. The new species, however, resembles a small number of species from the Southern Indian Ocean, Western Australia and south­east Asia (Hong Kong) that Erséus (1992a; 1997a) considered to constitute a monophyletic group within Pectinodrilus. Members of this group share ‘multiple club­shaped and hooked penial chaetae’ and ‘various modifications and elaborations of the vasa deferentia’. Of these, Pectinodrilus multiplex (Erséus 1990) and Pectinodrilus hoihaensis Erséus 1992 have a tripartite vas like the new species but with the middle muscular section even more developed and forming a broad sperm storage organ and with P. multiplex also possessing another ampulla (atrium sensu Erséus 1992a) between the vas and a copulatory sac. The male ducts of the new species are not this complex but are more differentiated than some other species in the group such as P. marionensis (Erséus 1979) and P. nervosus (Erséus 1990), which have a small non­ muscular section of vas followed by a longer extremely muscular section leading directly into a muscular atrium. Other members of this subgroup are P. glomeriductus Erséus 1997, which has highly coiled vasa and P. vitreus Erséus 1993 which has bi­partite vasa that are not as muscular as those of P. marionensis and P. nervosus).</p> <p>Etymology. Named for the type locality: a bore next to the homestead on Ningaloo Station.</p></div> 	https://treatment.plazi.org/id/03A287D5D803FFFA7A68FEFC1D4862DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pinder, Adrian M.;Eberhard, Stefan M.;Humphreys, William F.	Pinder, Adrian M., Eberhard, Stefan M., Humphreys, William F. (2006): New phallodrilines (Annelida: Clitellata: Tubificidae) from Western Australian groundwater. Zootaxa 1304 (1): 31-48, DOI: 10.11646/zootaxa.1304.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1304.1.3
03A287D5D805FFF87A68FDC11F08627B.text	03A287D5D805FFF87A68FDC11F08627B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus parvithecatus (Erseus 1978)	<div><p>Aktedrilus parvithecatus (Erséus 1978) (Fig. 3)</p> <p>Bacescuella parvithecata Erséus, 1978: 264</p> <p>Bacescuella pilicrepus Erséus, 1984b: 153</p> <p>Aktedrilus parvithecatus (Erséus, 1978); Erséus (1987: 117); Erséus (1992: 21)</p> <p>Material examined. WAM V 4446. Two specimens whole­mounted on same slide, from a small anchialine karst window (#C­510) about 500 m from the sea, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.816666&amp;materialsCitation.latitude=-22.333334" title="Search Plazi for locations around (long 113.816666/lat -22.333334)">Cape Range</a>, Western Australia (Fig. 1), 22°20'S 113°49'E, 8 Aug 1993. Conductivity 3.30 mS cm ­1, total dissolved solids 1800 mg L ­1. Coll. W.F. Humphreys and R. D. Brooks (WAM collection # BES 2502).</p> <p>Description of new material. Length and number of segments 6.3–6.6 mm and 48–53 respectively, width of slide­mounted worms 0.15–0.21 mm at segment XI. Clitellum from posterior 1/3 of X to 12/13, more glandular, but not distinctly thicker than epidermis of other segments. Male pores ventro­lateral on XI, about one third of the distance between 10/11 and 11/12. Single spermathecal pore mid­dorsal behind 9/10. Female funnels ventrolateral on 11/12.</p> <p>Prostomium bluntly oval, length:height at base 1.1–1.3, with large round clusters of cells below the epidermis, projecting into, and occupying up to half of the prostomial coelom. Pharynx in II/III. Pharyngeal glands, consisting of large irregular cells, associated with gut in IV–V (or anterior of VI). Rest of ciliated digestive tract narrow anteriorly, wider from XIV–XVII. Dorsal and ventral blood vessels connected by commissural vessels in at least IV and V and by a plexus of capillaries surrounding the gut from about VI. Large chlorogogue cells with non­staining cytoplasm almost filling coelom from VI or VII. Coelomocytes not observed. Chaetae bifid with sharp teeth, the upper tooth about half as long as the lower, 4–7/bundle in pre­genital segments and 3–5/bundle posteriorly, located at about two–thirds of the distance between anterior and posterior septa, 33–48 µm long x 1.3–1.5 µm wide at nodulus, largest in pre­genital segments, the nodulus slightly distal. Ventral chaetae present but not modified on X, absent on XI.</p> <p>Genitalia paired, except for spermatheca. Testes antero­ventral in X with sperm sacs extending posteriad to XIV. Ovaries antero­ventral in XI, egg sacs not present. Sperm funnels ventro­lateral on 9/10 leading to very short thin (8 µm) vasa deferentia (ciliation uncertain) joining apical end of tubular atria. Atria about 300 µm long, width about 17 µm with sperm present in the lumen (not shown on Fig. 3). Cytoplasm of atrial lining cells with numerous vesicles, much more so in ectal 2/3 rds, of a similar size and shape to those in the prostate glands. Muscle layer of atrium mostly thin, but in one specimen this is somewhat thicker around the ental part of the atrium with less abundant cellular vesicles. Atria joining ovoid penes apically, leading to a pore on the inner side of the penes. Penis sacs opening to exterior ventro­laterally. Two large prostate glands associated with each set of male ducts. Anterior prostates lying medial to the atria but connection to male duct not visible. Posterior prostates lying medial to penis sacs in front of 11/12, almost as tall as the segment and connected to the male duct where the atria join the penis sacs, sometimes protruding into XII. One spermatheca present with a small ampulla (28–31 x 16 µm) connected to mid­dorsal pore by thick duct (25–32 µm long, 13–14 µm wide) narrowing towards pore with narrow lumen. Loose sperm present in ampulla.</p> <p>Remarks. Apart from lacking external spermatophores, the new specimens conform well to descriptions and illustrations of A. parvithecatus by Erséus (1978; 1984a; 1984b; 1987) and Erséus and Cantú­Martínez (1984). The spermatophores are placed on the exterior body wall during reproduction in some species of Aktedrilus, including A. parvithecatus, although their absence in these three specimens is not of great significance: they were absent on 4 of 8 specimens from Rottnest Island collected by Erséus (1993). This species has mostly been collected from intertidal and barely subtidal sands (Erséus 1987,1993), although Erséus (1992b) also recorded it from brackish anchialine waters in the Cabo Verde Islands off north­west Africa. This species is obviously capable of living in a wide range of salinities as the population described above was collected in water with tds only 1800 mg L ­1. Occurrence in both marine littoral and brackish hypogean water (a mineral spring) has similarly been reported for Aktedrilus cuneus Erséus, 1984. Aktedrilus parvithecatus is circumtropical in distribution, with records from Pacific Mexico, Galapagos, France, Canary and Cabo Verde Islands, Saudi Arabia, Hong Kong, Bermuda and south­western Australia (Rottnest Island).</p> </div>	https://treatment.plazi.org/id/03A287D5D805FFF87A68FDC11F08627B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pinder, Adrian M.;Eberhard, Stefan M.;Humphreys, William F.	Pinder, Adrian M., Eberhard, Stefan M., Humphreys, William F. (2006): New phallodrilines (Annelida: Clitellata: Tubificidae) from Western Australian groundwater. Zootaxa 1304 (1): 31-48, DOI: 10.11646/zootaxa.1304.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1304.1.3
03A287D5D807FFF77A68FE211E1C66EA.text	03A287D5D807FFF77A68FE211E1C66EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus podeilema Pinder & Eberhard & Humphreys 2006	<div><p>Aktedrilus podeilema Pinder n. sp. (Fig. 4)</p> <p>Holotype. WAM V 4447. Incomplete specimen whole­mounted, from Tombstone Rocks Cave, about 9 km from the ocean, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.23333&amp;materialsCitation.latitude=-30.683332" title="Search Plazi for locations around (long 115.23333/lat -30.683332)">Nambung region</a>, Western Australia (Fig. 1), 30°41'S 115°14'E, 9 June 1998. Collected from sand and fine black silt with organic matter in a deep clear pool of freshwater. Coll. S. Eberhard (collection BES 5984).</p> <p>Description. Body&gt; 4.9 mm, number of segments&gt; 40, width of slide­mounted worm 0.19 mm at segment XI. Body wall of genital segments not thicker or more glandular than that of somatic segments. Male pores ventro­lateral, each antero­medial to a small papilla, located about 2/3 of the distance between 10/11 and 11/12. Single mid­dorsal spermathecal pore located within lobes of the body wall (which may be partially everted walls of a vestibule). Female funnels not seen.</p> <p>Prostomium bluntly conical, length:height at base 1.1, with thin lining tissue. Pharynx in II/III. Pharyngeal glands, consisting of irregularly shaped cells, dorsal to gut on posterior of 3/4 and lateral to gut on posterior of 4/5 and 5/6. Rest of ciliated digestive tract narrow with thin walls in III to X, slightly wider with thicker walls in genital region and much wider with even thicker lobed walls from XIV. Circulatory system not discernable. Chlorogogue tissue consisting of a single layer of cells with non­staining cytoplasm unevenly along the gut from VI. Coelomocytes not observed. Chaetae 4–6/bundle anteriorly, reduced to 3–4/bundle posteriorly, 33–48 µm long x 1–1.5 µm wide at nodulus, all bifid with sharp teeth, the upper tooth 1/2 to 3/4 the length of the lower and the nodulus slightly distal. Chaetal bundles of first few segments located 1/2 to 2/3 of the distance between anterior and posterior intersegmental furrows but most segments with bundles closer to the posterior septa. Ventral chaetae present but not modified on X, absent on XI.</p> <p>Genitalia paired except for spermatheca. Testes antero­ventral in X with sperm sacs from IX to XIII. Ovaries antero­ventral in XI with egg sacs to XIV. Sperm funnels ventrolateral on 9/10. Ciliated, non­glandular vasa deferentia about 340 µm long x 7–11 µm wide with thin muscle layer. Vasa deferentia joining atria subapically. Atria broad (50 x 95 µm) with muscle layer (1.5 µm). Numerous muscle fibres attach atria to dorso­lateral body wall (especially the ectal portion of atria) and with muscle layer of atria binding atrium to penis sac walls. Atria opening apically into stout and asymmetrical penes which are covered by a distinct cuticular sheath with a pointed antero­ventral projection. Penis sacs opening to exterior ventro­laterally. Two prostate glands associated with each set of male ducts. Posterior prostates smaller than anterior ones, lying posterior to penis sacs and communicating with male duct where the atria join the penes. Anterior prostates lying anterior to atria in vicinity of vasa deferentia but connection to male duct not visible. One spermatheca present in X, consisting of a swollen vestibule with a narrow pore ectally and connected entally to an elongate ampulla via a constriction. Loose sperm in ampulla.</p> <p>Remarks. Numerous other Aktedrilus possess cuticular penis sheaths. In particular, the asymmetrical sheaths of A. podeilema resembles those of two marine intertidal species: A. dentatus Erséus 1983b (from Brazil) and A. paradentatus Erséus 1984c (from Saudi Arabia). These two species, however, have other features (long tubular atria, posterior prostates attached broadly to the penis sacs and spermathecal pores posteriorly in IX or in intersegmental furrow 9/10) absent in the new species. Otherwise, A. podeilema, possessing a combination of asymmetrical penis sheaths, broad bipartite atria, long vasa deferentia and restricted connection between the posterior prostate and penis sacs, does not especially resemble any of its congeners.</p> <p>Etymology. From the Greek podos (foot) and eilema (sheath) referring to the foot–shaped penis sheath.</p></div> 	https://treatment.plazi.org/id/03A287D5D807FFF77A68FE211E1C66EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pinder, Adrian M.;Eberhard, Stefan M.;Humphreys, William F.	Pinder, Adrian M., Eberhard, Stefan M., Humphreys, William F. (2006): New phallodrilines (Annelida: Clitellata: Tubificidae) from Western Australian groundwater. Zootaxa 1304 (1): 31-48, DOI: 10.11646/zootaxa.1304.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1304.1.3
03A287D5D808FFF47A68F9B01E19670B.text	03A287D5D808FFF47A68F9B01E19670B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus leeuwinensis Pinder & Eberhard & Humphreys 2006	<div><p>Aktedrilus leeuwinensis Pinder n. sp. (Figs 5 and 6a)</p> <p>Holotype. WAM V 4448. Specimen whole­mounted, from sand and root mats (gently disturbed and caught in a net) in a pool area of a stream within <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.04722&amp;materialsCitation.latitude=-34.1" title="Search Plazi for locations around (long 115.04722/lat -34.1)">Budjur Mar Cave</a>, 34º06’00”S 115º02’50”E, 25 m asl, Leeuwin­Naturaliste Ridge, Western Australia (Fig. 1), 29 June 2002, coll. S. Eberhard.</p> <p>Paratypes. WAM V 4449 and 4450. Two mature whole­mounted, collection data as for holotype.</p> <p>Description. Length and number of segments 2.4–5.5 mm and 39–42 respectively, width of slide­mounted worms at segment XI 0.11–0.16 mm. Clitellum from posterior 1/3 of X to end of XII, epidermis of genital segments not thicker, but more glandular, than that of somatic segments. Anterior­most somatic segments with a distinct secondary annulation about 1/4 of the distance from anterior to posterior intersegmental furrow. Male pores ventro­lateral, about 1/3 of the distance from 10/11 to 11/12, in the form of crescentshaped slits medial to small papillae. Single spermathecal pore mid­dorsal behind 10/11, often with ectal part of vestibule partially protruding. Female pores not seen.</p> <p>Prostomium not much longer than tall, length:height at base 1.0–1.1, with loose clusters of cells below the epidermis. Pharynx in II/III. Pharyngeal glands, consisting of groups of large irregular cells, mostly lateral and dorsal to the gut in IV–VI. Digestive tract enlarging substantially between XI and XIV. Chlorogogue tissue present around gut but extent and distribution variable. Coelomocytes not observed. Chaetae 4–5/bundle anteriorly, reduced to 2(3)/bundle in post­clitellar segments, 25–30 µm long x 1.1–1.5 µm wide at nodulus, bifid with sharp teeth, the upper tooth 1/2 to 3/4 as long as lower, the nodulus distinctly distal.</p> <p>Genitalia paired, except for spermatheca. Male funnels ventro­lateral on 10/11, feeding narrow (4 µm) ciliated vasa deferentia (ciliation not illustrated). Vasa deferentia partially obscured by anterior prostate, presumably joining narrow ental end of atria, but union not seen. Atria narrow entally, expanding abruptly into a broad sac (length 70 µm, width = 34 µm) then tapering ectally to form a short ejaculatory duct which joins penes apically. Atrial muscle layer thin, lining tissue cells particularly large but position of lumen unclear — possibly towards one side of widest part of atria in holotype but more central in some other specimens (apparent position perhaps depending on aspect of view). Penes lying in deep penis sacs and enclosed by narrow, tapering and curved cuticular sheaths (Fig. 6), with the ental ends lateral to the gut and curving inwards (ventro­medially) so full extent of curve not seen from lateral view. Diagonal length of penis sheaths 20–25 µm. Penis sheaths of holotype presumably bent during lateral compression of specimen on slide (Fig. 5). Details of penis sacs difficult to discern (Fig. 5 is the authors’ interpretation) but each seeming to have a glandular lobe at least partially enclosing the penis but with uncertain attachment to the sac walls and of similar glandular histology to the prostate tissue, especially entally. Anterior prostates voluminous but connection to male ducts uncertain. Posterior prostate glands between the penis sac and septa 11/12, joining the male duct at the atrial/penis sac union. Spermatheca single, with a long ampulla (up to 141 x 37 µm), connected to the mid­dorsal pore by a smaller vestibule (up to 37.5 µm long x 29 µm). Loose sperm present in ampulla. Female funnels not seen. Egg sacs extending to XIII, sperm sacs from VIII–XIII.</p> <p>Remarks. Details of the male genitalia of this species needs to be confirmed further when new material becomes available. Nevertheless, this species most closely resembles a number of Aktedrilus which share narrow and frequently curved and/or tapering penis sheaths: A. argatxae Giani and Rodriguez 1988, A. mortoni Erséus 1984b, A. ruffoi Sambugar et al. 1999, A. svetlovi Finogenova 1976, A. longitubularis Finogenova and Shurova 1980, A. oregonensis Strehlow 1982, A. brevis Erséus 1980 and A. martiniquensis Erséus 1983b. The last four of these also have a glandular pad on the penis sac wall (longitubularis, brevis, martiniquensis) and/or a glandular lobe of tissue within the sac (brevis, oregonensis) associated with, or even representing the full extent of (longitubularis, martiniquensis), the posterior prostate gland. The precise anatomy of the penis sac and its interaction with the posterior prostate is not clear in the new species, but the narrow curved penis sheath and the suggestion of a glandular region within the penis sac indicates that A. leeuwinensis is allied with these marine intertidal species. Numerous features of these other species, however, distinguish them from leeuwinensis: martiniquensis and longitubularis have reduced posterior prostates and the former has simple­pointed chaetae posteriorly; longitubularis, oregonensis and brevis have more tubular atria; only oregonensis has a spermatheca with a well–defined duct/vestibule ectal to an ampulla (but with a much smaller ampulla than is present in leeuwinensis) and all except brevis have sperm partly embedded within the ampulla walls. In addition, the shape and/or size of the penis sheath differs in all of these species. Aktedrilus longitubularis is widespread and has been recorded from Rottnest Island off the south–west coast of Australia whereas oregonensis, martiniquensis and brevis are known only from the coasts of the north–east Pacific, the West Indies and the south–west Atlantic respectively.</p> <p>Etymology. Named after the geographical feature close to where the type material was collected: Cape Leeuwin.</p></div> 	https://treatment.plazi.org/id/03A287D5D808FFF47A68F9B01E19670B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pinder, Adrian M.;Eberhard, Stefan M.;Humphreys, William F.	Pinder, Adrian M., Eberhard, Stefan M., Humphreys, William F. (2006): New phallodrilines (Annelida: Clitellata: Tubificidae) from Western Australian groundwater. Zootaxa 1304 (1): 31-48, DOI: 10.11646/zootaxa.1304.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1304.1.3
