taxonID	type	description	language	source
03A2878BDA0AFFA675F0F88FFCC30342.taxon	diagnosis	Diagnosis (Figs. 1 A, 2 A, 3, 4). This species is similar to S. amurensis, but can be distinguished by the color of the wings and the genital morphology as follows. The coloration of the wing markings is darker than that of S. amurensis. In the male, width of the base of the uncus is narrower, height of the valva is narrower, the valval apodeme is slightly longer and projection of the carina of the phallus is weaker than in S. amurensis. In the female, the paired lamellae of the genital plate are angular than that of S. amurensis.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0AFFA675F0F88FFCC30342.taxon	description	Description. Length of forewing 12.5 – 17.9 mm; forewing expanse 25.6 – 36.5 mm. Head: Vertex and Frontoclypeus roughly covered with long brownish white hairs. Antenna filiform and strongly ciliated ventrally; each flagellomere covered with a row of dark brown scales. Maxillary palpus covered with grayish brown scales. Labial palpus extending anteriorly, first segment generally covered with brownish white scales plus few dark brown scales at middle; second segment covered with brownish white scales on inner side, but, dark brown on outer side. Thorax: mesonotum covered with brownish white scales; metanotum covered with brownish white scales on scutellum and posterior part of scutum; tegula dark brown anteriorly and brownish white posteriorly. Legs completely covered with grayish brown scales. Abdomen: covered with glassy pale grayish brown scales. Eighth abdominal segment without coremata. Wing markings (Fig. 1 A): Forewing upperside with dark brown scales in ground color, bordered with brownish white scales along dorsum and termen; blackish area with several weak brownish white speckles. Fringe consisting of brownish white scales and dark brown scales, the apex of which has 7 – 8 dentations. Hindwing upperside with pale grayish brown scales; basal 1 / 2 of costa covered with gray scales. Fringe consisting of grayish white scales and grayish brown, the apex of which has 6 – 7 dentations. Wing venation (Fig. 2 A): Forewing: all free veins present; R 1 arising from about 1 / 4 of discoidal cell; 1 A + 2 A about 2.5 times as long as 1 A. Hindwing: all free veins present; M 1 ending just posterior to the apex; 1 A + 2 A arched; M branched within discoidal cell. Male genitalia (Fig. 3): Tegumen and vinculum fused laterally to form a ring. Tegumen and uncus fused at base forming a dorsum. Uncus with a pair of elongate acute lobes and a small membranous protuberance between them; width of base of uncus about 1 / 2 length of uncus; gnathos and subscaphium absent. Vinculum broad; saccus broad with a rounded apex. Valva 1 / 3 as wide as height of ring, tightly fused with vinculum on ventral half; inner surface of valva setose on distal half, subapical part bulbous; middle of the base with a rounded protuberance, on which 6 – 7 setae arising; transtilla absent; valval apodeme 1 / 3 length of valva. Phallus as long as height of ring; slightly curved dorsally; vesica with sparse minute spicular cornuti; carina weakly projecting. Juxta with a pair of strongly sclerotized and elongate projections with setae apically; inner with thick and round conical process. Female genitalia (Fig. 4): Eighth abdominal tergum with a setose patch posterior to the tergum; eighth abdominal sternum complex, with a pair of long lateral digitiform processes with setae apically, and overlaid by a genital plate with a pair of triangular lamellae; about apical 1 / 3 of digitiform process extending beyond posterior margin of lamellae of genital plate. Bursa copulatrix about 2 / 3 length of apophysis posterioris. Ductus bursae 2 / 3 length of apophysis anterioris. Corpus bursae without signum.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0AFFA675F0F88FFCC30342.taxon	distribution	Distribution. Japan: Honshu (Shizuoka Pref.) and Kyushu (Kumamoto and Miyazaki Prefs.).	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0AFFA675F0F88FFCC30342.taxon	biology_ecology	Biology. The larvae feed on Fomes fomentarius [Japanese name: Tsuriganetake] (Moriuti, 1982).	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0AFFA675F0F88FFCC30342.taxon	etymology	Etymology. The name fusca, “ dark ” in Latin, refers to the wing color, which is darker than in any other Scardia species. Specimens examined. HOLOTYPE ♂, (Type No. 3456. ELKU) JAPAN: [Kyushu] (Miyazaki Pref.) Mt. Mukozaka, Gokase-machi, 30. vii. 2013 em., Host: Fomes fomentarius, 13. vi. 2013, Y. Osada leg, deposited in Entomological Laboratory, Kyushu University (ELKU). PARATYPES [Honshu] (Shizuoka Pref.) 1 ♂ 1 ♀, Nishimata, Tashiro, Aoi-ku, Shizuoka-shi, 15. vi. 2014 em., Host: Fomes fomentarius, 8. vi. 2014, K. Iwashita leg. [Kyushu] (Kumamoto Pref.) 1 ♂ 1 ♀, Mt. Ichifusa, Mizukami-mura, 6. vii. 1972 em., Host: Fomes fomentarius, 11. vi. 1972, S. Hisamatsu leg.; 1 ♂, Shiibagoe, Izumi-mura, 20. iv. 1996, M. Sakai leg.; (Miyazaki Pref.) 1 ♂, Mt. Mukozaka, Gokase-mura, 11. x. 1996, M. Sakai leg.; 2 ♂ 1 ♀, Mt. Mukozaka, Gokase-machi, 30. vii. 2013 em. Host: Fomes fomentarius, 13. vi. 2013, Y. Osada leg.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0AFFA675F0F88FFCC30342.taxon	discussion	Remarks. Sakai (2013) illustrated the male genitalia as S. amurensis, but we regarded it as S. fusca sp. nov. because the genital morphology agrees with those of the latter.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0FFFA575F0FF6AFEDC0376.taxon	diagnosis	Diagnosis (Figs. 1 B, 2 B, 5, 6). For morphological differences between S. fusca sp. nov. and S. amurensis, see ‘ Diagnosis’ for S. fusca sp. nov..	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0FFFA575F0FF6AFEDC0376.taxon	description	Redescription. Length of forewing 14.0 – 18.0 mm; forewing expanse 26.0 – 36.7 mm. Head, thorax and abdomen resembling those of S. fusca sp. nov., but different as follows: Vertex and face roughly clothed with yellowish white long hairs. Wing markings (Fig. 1 B): Forewing and hindwing resembling those of S. fusca sp. nov., but different as follows: Forewing upperside bordered with yellowish white along dorsum and termen. Hindwing upperside pale brownish white. Wing venation (Fig. 2 B): Wing venation resembling that of S. fusca sp. nov. Male genitalia (Fig. 5): Male genitalia resembling those of S. fusca sp. nov., but different as follows: Width of base of uncus similar to length of uncus. Gnathos and subscaphium absent. Vinculum broad; saccus broad with rounded apex. Valva 1 / 2 as wide as height of ring, valval apodeme 1 / 4 times as long as length of valva. Carina of phallus projecting. Female genitalia (Fig. 6): Female genitalia resembling those of S. fusca sp. nov., but different as follows. Genital plate with a pair of rounded quadrate lamellae; apical 1 / 4 of digitiform process extending beyond posterior margin of genital plate.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0FFFA575F0FF6AFEDC0376.taxon	distribution	Distribution. Japan: Hokkaido, Honshu (Aomori, Iwate, Nagano and Nara Prefs.), Russia (Amur, Primorsk).	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0FFFA575F0FF6AFEDC0376.taxon	biology_ecology	Biology. The larvae feed on Fomes fomentarius [Japanese name: Tsuriganetake] and Laricifomes officinalis [Japanese name: Eburiko] (Sohn 2007; Sakai 2013). Specimens examined. [Hokkaido] 1 ♂, Nukabira, Kamishihoro-cho, 29. vii. 1971, Host: Laricifomes officinalis, H. Ono leg.; 1 ♂ 1 ♀, Aizankei, Kamikawa-cho, 15. viii. 1996, M. Sakai leg.; 2 ♀, same locality and collector 28. vi. 1997; 2 ♂ 1 ♀, Mitsumata, Kamishihoro-cho, 3. vii. 1997, M. Sakai leg.; 2 ♂ 1 ♀, Shikaribetsulaleside, Shikaoi-cho, 23. viii. 1995, M. Sakai leg.; same locality and collector, 13. viii. 1996. [Honshu] (Aomori Pref.) 1 ♂, Sukayu, Towada-cho, 22. viii. 1996, M. Sakai leg.; (Iwate Pref.) 1 ♂ 1 ♀, Nakamura, Isawa-cho, 21. v. 1997, M. Sakai leg.; (Nagano Pref.) 1 ♂, Kayanotaira, Kijimadaira-mura, 13. viii. 1999, Y. Miyamoto leg.; (Nara Pref.) 1 ♂, Odaigahara, Kamikitamaya-mura, 6, viii, 2005, GH. Huang, T. Hirowatari. K. Tateiwa, BW. Lee and K. Ikeuchi leg. [Shikoku] (Ehime Pref.) 1 ♂ 1 ♀, Saragamine, Kumakogen-cho, 8. v. 1954, Host: Fomes fomentarius, S. Hisamatsu leg.; 1 ♂, Odamiyama, Oda-machi, 7. v. 1 986, Host: Fomes fomentarius, S. Hisamatsu leg. Sampling localities of Scardia species are shown in Fig. 7.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
03A2878BDA0FFFA575F0FF6AFEDC0376.taxon	discussion	Remarks. The male genitalia of Scardia amurensis and S. fusca sp. nov. are similar to those of S. boletella and S. caucasia, in lateral view, but the uncal lobes are slenderer, and not stout as in S. boletella and S. caucasia. In addition, in the two Japanese species, the digitiform processes of the female are slenderer than those of other Scardia species.	en	Osada, Yohei, Sakai, Makoto, Hirowatari, Toshiya (2016): A review of the Japanese species of the genus Scardia Treitschke (Lepidoptera, Tineidae), with description of a new species. Zootaxa 4079 (3), DOI: 10.11646/zootaxa.4079.3.4
