identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A714762562FF882A9DE9EAFAA7FF2E.text	03A714762562FF882A9DE9EAFAA7FF2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius jacksonii	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius jacksonii complex </p>
            <p> The  Amphilius jacksonii complex differs from all other species of the genus (  A. atesuensis ,  A. brevis ,  A. caudosignatus ,  A. dimonikensis ,  A. grammatophorus ,  A. kakrimensis ,  A. korupi ,  A. lamani ,  A. longirostris ,  A. maesii ,  A. mamonekenensis ,  A. nigricaudatus ,  A. opisthophthalmus ,  A. platychir ,  A. pulcher ,  A. rheophilus ) except  A. lentiginosus by its mottled body coloration that includes dark saddles that are joined to one another laterally (vs. body mottled, dark saddles not joined laterally). It differs from  A. lentiginosus by having the head and body without spots (vs. head and body heavily spotted) and by having fewer total gill rakers on the first gill arch (6–11, rarely 5 or 12 vs. 14–16). The  A. jacksonii complex further differs from  A. longirostris and  A. opisthophthalmus by having 6+7 principal caudal-fin rays (vs. 7+8) and a well developed crenelated epidermal fold (vs. rudimentary fold), and further differs from  A. brevis ,  A. dimonikensis ,  A. korupi , and  A. maesii by having distinct crenelations on the epidermal fold (vs. crenelations absent or very weak). It further differs from  A. grammatophorus ,  A. kakrimensis ,  A. platychir , and  A. rheophilus by having the lobes formed by epidermal fold crenelations rounded, Fig. 1 A (vs. lobes formed by epidermal fold crenelations elongated and pointed, Fig. 1 B) and from  A. brevis ,  A. caudosignatus ,  A. dimonikensis ,  A. korupi ,  A. lamani ,  A. maesii ,  A. mamonekenensis ,  A. nigricaudatus , and  A. pulcher by having fewer branchiostegal rays (usually 6–8, rarely 5 or 9 vs. 9–10 in  A. pulcher , more than 9 in the other species). </p>
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	https://treatment.plazi.org/id/03A714762562FF882A9DE9EAFAA7FF2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A714762561FF8C2A9DEF44FE45FE0F.text	03A714762561FF8C2A9DEF44FE45FE0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius jacksonii Boulenger 1912	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius jacksonii Boulenger 1912</p>
            <p>(Fig. 2, Table 1)</p>
            <p> Amphilius jacksonii Boulenger 1912: 602 , Original description, Type locality: Hima R. flowing into Lake George (Ruisamba), eastern Uganda, elev. 3500 ft. [Lake George drainage, Nile basin], holotype: BMNH 1912.10.15.47; Boulenger, 1916: 307, fig. 182, description based on type; David &amp; Poll, 1937: 255, description, Rutshuru River [Lake Edward drainage, Nile basin]; Poll, 1939: 17, 62, records from Albert National Park [Lake George and Lake Edwards drainages]; Harry, 1953: 190, synonymy; Greenwood, 1957: 77, figs. 49, 80, description, size, habitat, distribution [Lake George and Lake Edwards drainages]; Copely, 1958: 154, Hima R.; Greenwood, 1958: 89, fig. 49, description, size, habitat, distribution [Lake George and Lake Edwards drainages]; Whitehead, 1958: 198, western Uganda rivers; Corbet, 1961: 81 [not examined]; Greenwood, 1966: 93, fig. 49, description, size, habitat, distribution [Lake George and Lake Edwards drainages]; Seegers, 1996a: 188, figs. 133–134 (in part), type information, holotype figured, Seegers, 1996b: 251 (in part), type information, distribution; Walsh et al., 2000: 166, redescription, diagnosis, description, life-history aspects, habitat, distribution [Lake George drainage]. </p>
            <p>Material examined. Lake George drainage: AMNH 97419, Uganda, Dura River, Dura station, south end of Kibale Forest Reserve near Queen Elizabeth National Park, ca. 0°12'00”N, 30°22'00"E (6: 41.9–79.5); BMNH 1912.10.15.47, Uganda, Hima River, ca. 0°17'19”N, 30°10'23"E (1: 86.7, holotype); BMNH 1971.1.5.37–38, Uganda, Mapanga River, about 8 km upstream just below rapids, ca. 0°05'10.6”N, 30°23'03.8"E (2: 58.7–60.1); BMNH 1971.1.5.39, Uganda, Sibwe River, ca. 0°10'01.4”N, 30°12'36.8"E (1: 87.0); BMNH 1971.2.19.14–15, Uganda, Sibwe River, ca. 0°10'01.4”N, 30°12'36.8"E (1: 87.0); CU 97331, ex. UF 110746 (4: 37.6–92.7); MCZ 100587, Uganda, Middle Dura River, Kanyanchu, Kibale Forest Reserve, ca. 0°27'00”N, 30°22'00"E (1: 76.7); MRAC 90–046–P–68–78, Uganda, Ruimi (Rwimi) River, road between Fort Portal-Kasese [Route A 109], ca. 0°22'23.5”N, 30°12'40.8"E (11: 44.0–84.2); MRAC 90–046–P–79–87, Uganda, Ruimi (Rwimi) River, road between Fort Portal-Kasese [Route A 109], ca. 0°22'23.5”N, 30°12'40.8"E (11: 47.2–83.0); SAIAB 187245, ex. UF 110746 (42.3–97.0); UF 110743, Uganda, Middle Dura River, Kanyanchu, Kibale Forest Reserve, ca. 0°27'00”N, 30°22'00"E (12: 48.6–104.3); UF 110744, Uganda, Middle Dura River, Kanyanchu, Kibale Forest Reserve, ca. 0°27'00”N, 30°22'00"E (19: 23.6–115.9); UF 110745, Uganda, Middle Dura River, Kanyanchu, Kibale Forest Reserve, ca. 0°27'00”N, 30°22'00"E (5: 54.6–88.6); UF 110746, Uganda, Middle Dura River, Kanyanchu, Kibale Forest Reserve, ca. 0°27'00”N, 30°22'00"E (36: 34.1–102.0); UF 110747, Uganda, Dura River-Mainaro, ca. 0°37'00”N, 30°16'00"E (34: 54.4–111.1); UF 110748, Uganda, Ruimi River at bridge on road between Fort Portal and Hima, ca. 0°22'22.2”N, 30°12'41"E (41: 42.7–92.8); UF 110749, Uganda, Ruimi River at bridge on road between Fort Portal and Hima, ca. 0°22'22.2”N, 30°12'41"E (1: 47.6); UF 110750, Uganda, Mubuku River at bridge on road between Fort Portal and Kasese, ca. 0°15'30.8”N, 30°07'9.7"E (7: 52.8–85.1). Lake Edward drainage: AUM 47138, Uganda, Munyage River, below Butogota town, DRC-Uganda border foot path, elev. 1190m, 0°53'00”S, 29°37'50"E (3: 60.1–69.9); AUM 47139, Uganda, Tributary of Ishasha at bridge between Karangara and Butogota. Outside BINP, sources in park, elev. 1180m, 0°52'18”S, 29°39'59"E (2: 73.41–80.8); AUM 47143, Uganda, River Ishash, below bridge in cattle farm on Butogota-Kanungu road. elev. 1300m, 0°49'50”S, 29°38'21"E (2: 65.7–73.4); AUM 47146, Uganda, Bwindi Impenetrable Forest National Park, ca. 0°1'26”S, 29°41'23"E (4: 58.8–86.6); MRAC 22551, Uganda, Rutshuru, Buseregenyi, ca. 1°04'00”S, 29°26'00"E (1: 80.0); MRAC 23233, Uganda, Rutshuru, Buseregenyi, ca. 1°04'00”S, 29°26' 00"E (1: 82.2); MRAC A5–019– P–0048–0056, Uganda, Munyage River, below Butogota town, DRC-Uganda border foot path, elev. 1190m, ca. 1°04'00”S, 29°26'00"E (9: 39.4–97.3); UF 169241, ex. AUM 47143 (3: 65.9–70.7); UF 169242, ex. AUM 47139 (3: 80.7–87.3); UF 169243, ex. AUM 47138 (2: 67.1–86.3); UF 169257, ex. AUM 47146 (3: 60.7–75.1). Kagera River drainage: MRAC 164876, Rwanda, Mwogo River, 18 km NE of Butare, ca. 2°29'00”S, 29°38'00"E (1: 77.2); MRAC 71783–850, Rwanda, Mwogo River, the source of the Nile, ss-tributary of Kagera River, ca. 2°22'52”S, 29°41'48"E (61: 38.5–107.0); MRAC 71851–925, Rwanda, Mwogo River, the source of the Nile, sstributary of Kagera River, ca. 2°22'52”S, 29°41'48"E (74: 38.5–106.4); MRAC 83417, Burundi, Urigoli, Nyalugogo [Nyabugogo?] River, Urigoli territory, ca. 3°25'26”S, 29°53'30"E (1: 34.7); MRAC 86–27–P–0088– 0 0 95, Rwanda, Rubondo River, tributary of Kiryango River, near town of Mukugi, ca. 2°12'00”S, 29°41'00"E (8: 51.8–86.1); MRAC 91–30–P–0296–0299, Burundi, Nyakijanda River, tributary of Ruvubu River, the bridge on Road of General Interest 4, 38 km from Kinyinya, ca. 3°36'00”S, 30°07'00"E (4: 49.5–65.0); MRAC 91–34–P– 0147–0150, Burundi, Kavuruga River, tributary of Ruvubu River, near town of Buhinyuza, ca. 3°02'00”S, 30°21'00"E (4: 49.4–74.9); MRAC 91–34–P–0151–0152, Burundi, Kinyanderama River, tributary of Ruvubu River, near town Buhinyuza, ca. 3°02'00”S, 30°21'00"E (2: 66.6–71.1); MRAC 91–34–P–0160–0164, Burundi, Nyabiko River, on Muyinga-Gitega road, small tributary, rocky and sandy bottom, ca. 5°29'00”S, 30°14'00"E (5: 35.1–92.3); MRAC 91757–763, Rwanda, Nyabugogo River at confluence of Lusine River, ca. 1°47'41”S, 30°07'23"E (7: 58.9–96.8); MRAC 91764–836, Rwanda, Nyabugogo River at confluence of Lusine River, ca. 1°47'41”S, 30°07'23"E (72: 34.9–79.8); MRAC 93097–120, Rwanda, Nyabugogo River, at outlet of Lake Mohasi, ca. 1°46'45”S, 30°07'58"E (23: 28.2–77.0); MRAC 94272–276, Rwanda, ca. 2°32'15”S, 29°40'16"E (5: 40.1– 82.9); MRAC 96031.1611–1614, Burundi, Karuzi, small tributary stream of the fish-farming ponds, Ndurumu system, ca. 3°06'05”S, 30°09'53"E (4: 29.3–78.4).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  Amphilius jacksonii is diagnosed from all other species of the  Amphilius jacksonii complex by its more slender caudal peduncle (4.8–7.9% vs. 8.1–12.3% SL). It is further distinguished from  A. pedunculus ,  A. frieli , and  A. crassus by its longer caudal peduncle (caudal-peduncle length 16.7–20.6 vs. 13.3–18.8% SL), and from  A. frieli ,  A. crassus , and  A. lujani by its more slender body (body depth at anus 9.6–13.2% vs. 13.5–17.4% SL). It is further distinguished from  A. frieli by having fewer total gill rakers on the first gill arch (6–9, rarely 10 vs. 10–11, rarely 9 or 12) and from  A. ruziziensis by having a wider interorbital width (26.7–32.0 HL vs. 23.4–25.1% HL). </p>
            <p>Description. Morphometric data are in Table 1. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising gently from tip of snout to origin of dorsal fin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital opening located at midpoint of adpressed pelvic fin, closer to pelvic-fin insertion than to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately joined at isthmus forming a U -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, slightly papillate. Rictal lobe large and papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming V -shaped band with anterior broad protrusion. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth to pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to pectoral-fin origin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to edge of branchiostegal membrane.</p>
            <p>Branchiostegal membrane with 6 (5), 7 (85), 8 (147), or 9 (1) rays. Gill rakers on first epibranchial 1 (4), 2 (178) or 3 (47); rakers on first ceratobranchial 4 (22), 5 (88), 6 (99), 7 (15), or 8 (1); total gill rakers on first arch 6 (25), 7 (74), 8 (83), 9 (38), or 10 (6).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit, covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over tip of adpressed pectoral fin. Dorsal fin with i,6 (243) rays, margin straight. Pectoral fin with i,7 (1), i,8 (104), or i,9 (137) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior to dorsalfin insertion. Pelvic fin with i,5 (243) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base; fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, not deeply incised posteriorly. Caudal fin deeply forked with tips of lobes rounded; fin with i,5,5,i (8), i,5,6,i (230), or i,6,6,i (1) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,6 (62), ii,7 (107), ii,8 (9), iii,6 (48), iii,7 (16), or iv7 (1) rays. Anal-fin margin almost straight.</p>
            <p> Character  Amphilius A. ruziziensis A. pedunculus A. frieli A. crassus A. lujani</p>
            <p> jacksonii</p>
            <p>Coloration. Body variably mottled and with dark saddles. First saddle immediately posterior of head, second saddle under dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddles connected laterally by broad stripe. Venter light brown with fourth and fifth saddles meeting those of opposite side. Dorsal and anal fins light brown with dark medial band. Adipose fin dark brown, cream-colored distally. Pectoral and pelvic fins positioned horizontally with upper surfaces brown and lower surfaces light yellow, each with dark medial band. Caudal fin brown with medial cream patch and tips of upper and lower lobes cream. Caudal coloration asymmetrical, lower lobe with more pigment than upper lobe.</p>
            <p>Distribution. Lake George, Lake Edwards, and Kagera River drainages, Nile basin, eastern Uganda, Rwanda, and Burundi (Fig. 3).</p>
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	https://treatment.plazi.org/id/03A714762561FF8C2A9DEF44FE45FE0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A714762564FF832A9DEC93FC47F979.text	03A714762564FF832A9DEC93FC47F979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius ruziziensis Thomson & Page	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius ruziziensis Thomson &amp; Page ,  new species</p>
            <p>(Fig. 4, Table 3)</p>
            <p> Amphilius platychir (non Günther).— Marlier, 1953: 194, distribution in Ruzizi drainage; Distribution mapped. </p>
            <p>Holotype. MRAC 93294, Rwanda, Lufiro River, upper reaches of Matchuza, Ruzizi River drainage, ca. 2°44'00”S, 29°02'99"E (82.8 mm SL)</p>
            <p>Paratypes. CU 97332, ex. MRAC 93300, same data as holotype (1: 47.7); MRAC 91529–531, Democratic Republic of the Congo, Kiliba River, ca. 3°14'30”S, 29°09'36”E (3: 31.2–64.9); MRAC 93295–297, same data as holotype, (3: 41.3–45.3); MRAC 93126–128, Burundi, Nyakagunda River, ca. 2°47'S, 29°04'E (3: 37.5–66.5); MRAC 93129–130, Burundi, Nyamagana River, ca. 2°55'S, 29°08'E (2: 116.1–120.6); MRAC 93131, Burundi, Lua River, ca. 2°46'S, 29°02'E (1: 79.5); SAIAB 187262, ex. MRAC 93298, same data as holotype (1:64.2); UF 184236, ex. MRAC 93299, same data as holotype (69.8).</p>
            <p>Non-types. Mutimbuzi River drainage: MRAC 91330–389, Burundi, Murago River, ca. 3°17'S, 29°23'E (59: 29.2–71.4); MRAC 93076–096, Burundi, Musazi River [=Muzazi River], ca. 3°17'S, 29°25'E (20: 33.1–91.0); MRAC 126266–267, Burundi, Murago River, ca. 3°17'S, 29°23'E (7: 41.3–82.8). Ruzizi River drainage: MRAC 91034.0153–0159, Burundi, Nyamagana River near town of Mabayi, ca. 2°42'S, 29°15'E (7: 47.7–79.9); MRAC 91390, Burundi, Nyakagunda River, Bugarama, ca. 2°47'S, 29°04'E (1: 25.6); MRAC 93051–054, Democratic Republic of the Congo, Muniowe River, Luvungi, ca. 2°56'S, 28°57'E (4: 43.9–95.2); MRAC 93055–075, Democratic Republic of the Congo, Luvubu River, ca. 2°52'S, 29°02'E (20: 27.3–108.5); MRAC 93121–125, Burundi, Nyamagana River, ca. 2°55'S, 29°08'E (5: 44.1–97.6), MRAC 96031.1571–1576, Burundi, Nyamagana River, 3 km after Mabayi, center Kivogero, ca. 2°42'56”S, 29°14'40”E (5: 46.2–88.0); MRAC 96031.1590–1595, Ikibenga River, 26 km on Cibitoke-Mabayi road, ca. 2°43'03”S, 29°10'05”E (6: 50.5–69.3). Unknown drainage: MRAC 91392–395, Burundi, Kikoma River, ca. 3°20'S, 29°18'E (4: 45.3–86.6).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  A. ruziziensis is diagnosed from  A. pedunculus ,  A. frieli , and  A. crassus by its more slender caudal peduncle (caudal peduncle depth 8.6–9.5% SL vs. 9.7–12.3% SL) and from  A. jacksonii ,  A. pedunculus ,  A. crassus , and  A. lujani by its narrower interorbital width (23.4–25.1% HL vs. 26.7–35.8% HL). It is further diagnosed from  A. frieli ,  A. crassus , and  A. lujani by its more slender body (body depth at anus 11.1–12.9% SL vs. 13.5–17.4% SL).  A. ruziziensis is further diagnosed from  A. frieli by having fewer branchiostegal rays (6–7 vs. 8–9), fewer total gill rakers on the first gill arch (6–8, rarely 5 or 9 vs. 10–11, rarely 9 or 12), and a longer caudal peduncle (caudal peduncle length 18.7–20.3% SL vs. 14.4–16.4% SL). It is further diagnosed from  A. crassus by a longer caudal peduncle (caudal peduncle length 18.7–20.3% SL vs. 13.3– 15.5% SL) and shorter dorsal-fin insertion to adipose-fin insertion length (38.2–41.5% SL vs. 42.2–44.6% SL). It is further diagnosed from  A. jacksonii by its deeper caudal peduncle (caudal peduncle depth 8.6–9.5% SL vs. 4.8– 7.9% SL). </p>
            <p>Description. Morphometric data are in Table 3. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising gently from tip of snout to dorsal-fin origin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital openings located at midpoint of adpressed pelvic fin, much closer to pelvic-fin insertion than to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt to rounder when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately to broadly joined at isthmus forming a U -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, slightly papillate. Rictal lobe large and slightly papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming crescent shaped band. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth, not reaching pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to just short of origin of pectoral-fin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to about two-thirds of distance to edge of branchiostegal membrane. Branchiostegal membrane with 6 (7), or 7 (8) rays. Gill rakers on first epibranchial 2 (15) or 3 (1); rakers on first ceratobranchial 3 (1), 4 (8), 5(2), 6 (4), or 7 (1); total gill rakers on first arch 5 (1), 6 (8), 7 (1), 8 (5), or 9 (1).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit; covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over or just posterior to tip of pectoral fin. Dorsal fin with i,6 (16) rays, and fin margin straight. Pectoral fin with i,8 (2), or i,9 (14) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior of dorsal-fin insertion. Pelvic fin with i,5 (16) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base, fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, deeply incised posteriorly. Caudal fin moderately forked with tips of lobes rounded; fin with i,5,6,i (15) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,5 (3), ii,6 (2), iii5 (1), iii,6 (7), or iii,7 (3) rays. Anal-fin margin almost straight.</p>
            <p>Coloration. Body variably mottled with dark saddles. First saddle posterior of head, second saddle at dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddle connected laterally by broad stripe. Venter light brown with fourth saddle meeting that of opposite side Dorsal, and anal fins brown with dark medial band (band on anal fin often indistinct on small specimens). Adipose fin dark brown, cream-colored anteriorly, posteriorly and on distal edge. Pectoral and pelvic fins positioned horizontally with upper surfaces brown with cream-colored distal edge. Lower surfaces light yellow. Caudal fin cream-colored with medial dark band.</p>
            <p>Distribution. Ruzizi River drainage, eastern Rwanda and Burundi; western Democratic Republic of the Congo; northeastern tributaries of Lake Tanganyika in Burundi (Fig. 3).</p>
            <p>Etymology. Named for the Ruzizi River drainage in eastern Rwanda and Burundi, where type specimens of this species were collected and the species is primarily distributed.</p>
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	https://treatment.plazi.org/id/03A714762564FF832A9DEC93FC47F979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A71476256AFF842A9DEAD1FE6FFB86.text	03A71476256AFF842A9DEAD1FE6FFB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius pedunculus Thomson & Page	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius pedunculus Thomson &amp; Page ,  new species</p>
            <p>(Fig. 5, Table 4)</p>
            <p> Amphilius jacksonii .—David, 1937: 418, Malagarasi River; De vos et al., 2001: 131, Malagarasi River. </p>
            <p>Holotype. CU 97334, ex. CU 95207, Tanzania, Malagarasi River at Lower Igamba Falls (Kasagwe), Malagarasi River drainage, 5°10'48”S, 30°03'03"E (52.3 mm SL).</p>
            <p>Paratypes. Malagarasi River drainage: CU 90414, Tanzania, downstream of bridge across Malagarasi River at Uvinza, 5°07'03.7”S, 30°22'27.1"E (1: 50.8); CU 95207, same data as holotype, (16: 41.3–69.8); CU 95208, Tanzania, Malagarasi River up river 12 km E of Uvinza near village of Kanzibwe, 5°08'23.5”S, 30° 29' 21.6"E (1: 55.3); CU 95209, Tanzania, Malagarasi River downriver 4 km W from Uvinza; village of Nkwasa, 5°05'52.6”S, 30°21'16.2"E (5: 38.1–74.7); CU 95210, Tanzania, Malagarasi River near fork in road to Ngutu, 5°06'56.3”S, 30°17'48.1"E (1: 67.6), CU 95211, Tanzania, Malagarasi River in first site in gorge, 5°13'44.2”S, 30°13'26.5"E (6: 41.2–52.9); CU 95212, Tanzania, Malagarasi River in gorge, 5°12'10.2”S, 30°10'11.1"E (1: 44.8); MRAC B3–06– P–1–3, ex. CU 95209 (3: 46.3–54.6); SAIAB 187275, ex CU 95209 (3: 45.6–52.6); UF 184234, ex CU 95209 (3: 41.8–64.7).</p>
            <p>Non-types. Malagarasi River drainage: CU 90426, Tanzania, Malagarasi River at bridge between Kafuru and Makere, 4°01'32.5”S, 30°33'06.5"E (3:24.6–29.3), CU 90469, Tanzania, Malagarasi River at bridge between Kafuru and Makere, 4°01'32.5”S, 30°33'06.5"E (26: 24.7–46.7); MRAC 47343–362, Burundi, Malagarazi River and its tributaries, (19: 38.7–96.2), MRAC 91030.0217, Burundi, Nyesasa River, trib of Muyovozi River (which is a tributary of the Malagarazi), at bridge on RN8 [now RN11] just before Gihofi, ca. 3°59'S, 30°09'E (1: 51.4); MRAC 91030.0218–0225, Burundi, Kinywa River, Trib of Malagarazi, at the bridge on Route Pr85, near town of Buyaga, ca. 3°55'S, 30°10'E (7: 16.2–69.7); MRAC 91030.0226–0248, Burundi, Nyankanda River, Trib of Malagarazi, in the Fault of the Germans, on Route Pr 85, ca. 3°54'S, 30°13'E (23: 33.5–65.4); MRAC 91030.0249– 0 295, Burundi, Ntanga River, Trib of Malagarazi, at the bridge on Road of General Interest 4, 7 km from Kinyinya, ca. 3°37'39.3”S, 30°17'54.5”E (47: 26.4–76.6); MRAC 91030.0300–0309, Burundi, Musasa River, Trib of Muyovozi, at the bridge on RN8, ± 5 km Rutana, ca. 3°58'44.9”S, 30°01'56.0”E (10: 45.6–71.5); MRAC 91030.0310–0315, Burundi, Mutsindozi River, trib of Malagarazi at bridge on Route RN12, km 39 from Lake Nyanza, ca. 4°05'S, 29°57'E (6: 54.7–67.5); MRAC 91061.0106, Burundi, Musasa River, Trib of Muyovozi, at the bridge on RN8, ± 5 km Rutana, ca. 3°58'44.9”S, 30°01'56.0”E (1: 55.3); MRAC 91062.0896–0923, Burundi, Mutsindozi River, near the sugar refinery, Gihofi, ca. 4°02'S, 30°09'E (28: 29.6–73.1); MRAC 91062.0924–0929, Burundi, Musasa River, Trib of Muyovozi, at the bridge on RN8, ± 5 km Rutana, ca. 3°58'44.9”S, 30°01'56.0”E (6: 36.4–85.9); MRAC 91062.0930–0932, Burundi, Mashuro River, about 5 km from Gihofi towards Giharo, 3°57'37”S, 30°09'54.6”E (3: 28.9–48.7); MRAC 91062.0933, Burundi, Ruru River, trib of Rumpungwe River, on the road towards Cendajuru, ca. 3°19'S, 30°34'E (1: 79.2); MRAC 91062.0950–0957, Burundi, Kabingo, Mazimero River, on Rutana-Kinyinya road, 3°53'01.5”S, 30° 11'54.2”E (8: 37.1–63.1); MRAC 91062.0958–1056, Burundi, Rugoma River, trib of Rumpungwe River, 4 km from Kinyinya, ca. 3°38'S, 30°23'E (99: 24.6–100.9); MRAC 91062.1057–1081, Burundi, Rugaragara River, on Kinyinya-Gisuru road, about ± 20 km from Gisuru, ca. 3°27'S, 30°24'E (25: 25.9–67.7); MRAC 91062.1082–1094, Burundi, Kiruhura River, trib of Mwambu River, zone Muyange, ENE of Cendajuru, ca. 3°16'S, 30°40'E (13: 32.6–71.8); MRAC 91062.1095–1107, Burundi, Nyabigosi River, trib of Rumpungwe River, near Gisuru, ca. 3°29'S, 30°30'E (13: 28.3–68.5); MRAC 91062.1139–1168, Burundi, Gitinwa River, trib of Ruru River, trib of Rumpungwe River, near Cendajuru, before village of Gusiana, ca. 3°20'S, 30°34'E (30: 24.6–83.9); MRAC 91062.1169–1245, Burundi, Mukitcha River, ± 8 km towards Cendajuru from Gasenyi, ca. 3°17'S, 30°36'E (77: 28.7–74.6); MRAC 91079.0055–0058, Burundi, Musasa River, (local name Uruhuzi) 78 km SSE of Gitega, ± 10 km SE of Rutana, 4°00'00”S, 30°06'47.3”E (5: 25.9–71.9); MRAC 93150.0184–0186, Burundi, Mazimero River, on road from Rutana-Kinyinya, ca. 3°54'S, 30°13'E (3: 63.5– 75.9); MRAC 93150.0187–0238, Burundi, Ruru River, about 9 km from Muyaga towards Cendajuru, ca. 3°18'S, 33°13'E (52: 35.4–80.4); MRAC 93150.0239, Burundi, Ntanga River, Trib of Malagarazi, at the bridge on Road of General Interest 4, 7 km from Kinyinya, ca. 3°38'S, 30°18'E (1: 52.7); MRAC 93150.0240–0260, Burundi, Mukazye River (Nyamabuye), 10 km from Giharo towards Kinyinya, ca. 3°47'S, 30°18'E (21: 26.9–72.9); MRAC 93150.0261–0333, Burundi, Idumaniro River, on Cendajuru-Gitwenge road, ca. 3°15'S, 30°39'E (73: 25.8–81.1); MRAC 93150.0351–0359, Burundi, Rumpungwe River, near Gisuru, on road from Kinyinya-Gisuru, ca. 3°27'S, 30°29'E (9: 26.2–74.3); MRAC 93150.0360–0378, Burundi, Kiruhura River, on Cendajuru-Gitwenge road, ca. 3°15'S, 30°39'E (18: 35.8–65.8), MRAC 93150.0379–0382, Burundi, Nyanzari River, on Cankuzo-Mushiha road, near Gishungo, 3°04'25”S, 30°39' 22”E (4: 48.3–68.6); MRAC 93150.0383–0392, Burundi, Mutsindozi River, near the sugar refinery, Gihofi, ca. 4°00'S, 30°40'E (12: 44.2–80.0); MRAC 93152.0613–0622, Tanzania, Malagarazi River, rapids at Uvinza, near salt mine, 5°06'07”S, 30°21'56”E (11: 37.1–59.9); MRAC 93152.0623– 0 643, Tanzania, Nyamgongo River, trib of Malagarazi River, km 46 from Uvinza to Kasulu, 4°48'34”S, 30°12'51”E (21: 28.3–69.0); MRAC 96031.1548–1549, Burundi, Nyarugunga River, km 30 after Kinyinya, on Kinyinya-Gisuru road, ca. 4°52'S, 29°50'E (2: 53.7–58.6); MRAC 96031.1550–1562, Burundi, Nyarugunga River, km 30 after Kinyinya, on Kinyinya–Gisuru road, ca. 4°52'S, 29°50'E (13: 47.6–86.6); MRAC 96031.1563–1570, Burundi, Nyarubare River, km 15 on Kinyinya-Gisuru road, ca. 3°34'S, 30°26'E (8: 45.7–81.8); MRAC 96083.1141, Tanzania, Ruchigi River, 7 km after Kasulu to Kibando, trib of Malagarasi, 4°32'01.4”S, 30°08'58.6”E (1: 71.5); MRAC 96083.1142–1145, Tanzania, Mgandazi River, near Kasulu, ca. 4°37'S, 30°06'E (4: 48.7–72.6). Luiche River drainage: MRAC 93152.0644–0649, Tanzania, Mukuti River, trib. of Luiche River, on road from Kigoma to Kasulu, 4°53'12.1”S, 29°52'12”E (6: 35.0–118.8); MRAC 93152.0650–0651, Tanzania, Mungonya River, trib of Luiche River, ± 10 km from Kigoma, route Kigoma-Kasulu, 4°52'28”S, 29°49'52”E (2: 65.2–90.9); MRAC 93152.0652–0661, Tanzania, Kidahwe River, km 34 on Kigoma-Uvinza road, trib of Luiche River, ca. 4°53'S, 29°48'E (12: 68.6–112.0). Lake Rukwa Basin: MRAC 191055 –057, Zambia, Saisi and Kalambo River, Tunduma road, ca. 9°06'S, 31°29'E (3: 30.6–39.1); MRAC 191092 –113, Zambia, Saisi River, at crossing of Abercorn-Tunduma road, 9°05'38.1”S, 31°29'20.5”E (22: 27.5–104.8); MRAC 191423–424, Zambia, Lumi river, Kawimbe Road, ca. 8°50'S, 31°32'E (2: 31.6–35.4); MRAC 94034.0822–0831, Tanzania, Piti River, 63 km south of Rungwa River on road to Makongolosi, 7°26'48.5”S, 33°25'22.1”E (10: 32.9–66.6); SAIAB 37397, Zambia, Near Mbala, Saisi River at Tunduma Road near Neilsons Farm, ca. 9°05'S, 31°32'E (1: 55.4); SAIAB 38096, Zambia, Saisi River at Tunduma Road bridge, 9°05'39”S, 31°29'22”E (1: 35.8); SAIAB 39557, Zambia, Saisi River at Tunduma Road bridge, ca. 9°05'39”S, 31°29'22”E (1: 57.0); SAIAB 38108, Zambia, Chitungulu stream, Nsunzu Farm, trib of Saisi River, ca. 9°05'S, 31°32'E (2: 54.3–70.3); SAIAB 39543, Zambia, Chitungulu stream, Nsunzu Farm, trib of Saisi River, ca. 9°05'S, 31°32'E (1: 37.6); SAIAB 50333, Tanzania, Lupa River at Lupatingatinga, on Makongolsi-Rungwa road, 8°01'48.9”S, 33°16'18.9”E (1: 42.0). Rufiji basin, Upper Great Ruaha River drainage: SAIAB 59388, Tanzania, Bridge near Chinata on Mbeya-Iringa road, 8°51'35.7”S, 34°01'33”E (31: 49.2–111.7); SAIAB 59397, Tanzania, Great Ruaha River at Route A104 crossing just east of Chimala, 8°51'16.4”S, 34°05'7.0”E (32: 30.3–72.6).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  Amphilius pedunculus is diagnosed from  A. jacksonii ,  A. ruziziensis and  A. lujani by its deeper caudal peduncle (caudal peduncle depth 9.9–12.3% SL vs. 4.8– 9.5% SL), and from  A. ruziziensis and  A. frieli by its wider interorbital width (28.1–35.8% HL vs. 23.4–27.7% HL). It is distinguished from  A. crassus by its longer caudal peduncle (caudal peduncle length 16.0–18.8% SL vs. 13.3–15.5% SL) and shorter dorsal-fin insertion to adipose-fin insertion length (37.5–41.7% SL vs. 42.2–44.6% SL). It is further diagnosed from  A. frieli by having fewer total gill rakers on the first gill arch (7–9, rarely 6 or 10 vs. 10–11, rarely 9 or 12). </p>
            <p>Description. Morphometric data are in Table 4. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising gently from tip of snout to dorsal-fin origin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital openings located at midpoint of adpressed pelvic fin, much closer to pelvic-fin insertionthan to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately joined at isthmus forming a V -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, slightly papillate. Rictal lobe large and slightly papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming crescent shaped band. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth, to pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to origin of pectoral-fin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to edge of branchiostegal membrane. Branchiostegal membrane with 7 (17), or 8 (24) rays. Gill rakers on first epibranchial 2 (20) or 3 (21); rakers on first ceratobranchial 4 (1), 5 (8), 6 (25), or 7 (7); total gill rakers on first arch 6 (1), 7 (4), 8 (16), 9 (16) or 10 (4).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit; covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over or just posterior to tip of pectoral fin. Dorsal fin with i,6 (41) rays, and fin margin straight. Pectoral fin with i,8 (35), or i,9 (6) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior of dorsal-fin insertion. Pelvic fin with i,5 (41) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base, fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, deeply incised posteriorly. Caudal fin deeply forked with tips of lobes rounded; fin with i,5,6,i (40) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,6 (7), ii,7 (2), iii5 (4), iii,6 (27), or iii,7 (1) rays. Anal-fin margin almost straight.</p>
            <p>Coloration. Body variably mottled with dark saddles. First saddle posterior of head, second saddle at dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddle connected laterally by broad stripe. Venter light brown with fourth and fifth saddles meeting those of opposite side. Dorsal, and anal fins light brown with dark medial band. Adipose fin dark brown to black, with posterior distal edge cream-colored. Pectoral and pelvic fins positioned horizontally with upper surfaces cream-colored with dark fin base and dark medial band. Lower surfaces light yellow. Caudal fin creamcolored with dark markings on upper and lower lobes. Caudal coloration asymmetrical, with lower lobe almost completely dark (except for small cream-colored mark at base of caudal and cream-colored tip). Upper lobe with less dark pigment, with dark blotch covering dorsal edge, but caudal base, medial ventral edge, and tip creamcolored.</p>
            <p>Distribution. Malagarasi River drainage, western Tanzania and southern Burundi; Luiche River drainage, western Tanzania; Lake Rukwa basin, Tanzania and Zambia; and upper Great Ruaha River system, Rufiji basin, eastern Tanzania (Fig. 6).</p>
            <p> Etymology. The diminutive of pes,  pedunculus , a noun, refers to the distinctively short and deep caudal peduncle of this species. </p>
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	https://treatment.plazi.org/id/03A71476256AFF842A9DEAD1FE6FFB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A71476256DFF9B2A9DE8CBFA5BF979.text	03A71476256DFF9B2A9DE8CBFA5BF979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius frieli Thomson & Page	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius frieli Thomson &amp; Page ,  new species</p>
            <p>(Fig. 7, Table 5)</p>
            <p> Amphilius platychir (non Günther).— Boulenger, 1920: 26, Lubumbashi à Élisabethville (MRAC P–6787) [Kafubu River system, Upper Congo basin] </p>
            <p>Holotype. CU 97335, ex. CU 91052, Zambia, Tswishi Stream at bridge under construction, Lwela River system, 11°33'26”S, 29°09'59"E (83.1 mm SL).</p>
            <p>Paratypes. Lwela River system: AUM 57570, ex. CU 91049 (3: 36.3–51.7); CU 91049, Zambia, Lwela River at bridge on Mansa-? road, 11°33'25”S, 29°10'10"E (21: 32–104.5); CU 91050, Zambia, Ngo Stream, about 40 km SW of Mansa, 11°31'33”S, 29°09'03"E (4: 34.2–46.8); CU 91052, same data as holotype, (11: 31.7–44.9); SAIAB 76634, same data as CU 91049 (12: 32.7–86.7); SAIAB 76789, same data as CU 91050 (5: 36.0–47.5); MRAC B3–06–P–4–6, ex. CU 91049 (3: 41.9–47.4); SAIAB 76799, same data as holotype (13: 27.3–57.2); UF 184235, ex. CU 91049 (3: 39.0–50.8).</p>
            <p>Non-types. Chambeshi River system: CU 91056, Zambia, Kanchibiya Stream at bridge on Kasama-Mpika road, 11°29’44.2”S, 31°16'46.6"E (28: 32.5–57.0); MRAC 96031.1540–1547, Zambia, Musombizi River, trib of Chambeshi, ± 55km on the road Mbala-Kasama, ca. 09°18’S, 31°16.5'E (8: 25.8–57.8); MRAC 96031.1620–1625, Zambia, Chambeshi River, at bridge km 40 on road Mbala-Kasama, 09°12’15.3”S, 31°20'49.8"E (6: 57.7–135.6); SAIAB 40125, Zambia, Chambeshi River, rapids near Kapoloso Stream, ca. 10°43’S, 31°15'E (1: 47.8); SAIAB 46788, Zambia, Mansha River, Shiwa Hot Springs &amp; road bridge, ca. 11°10’S, 31°35'E (16: 32.0–64.8); SAIAB 77139, Zambia, Samfa Rapids at pontoon on Chambeshsi River, 10°51’07.6”S, 31°10'02.3"E (1: 69.5); SAIAB 77142, same data as CU 91056 (28: 29.4–85.3). Kafubu River system: MRAC 6787, Democratic Republic of the Congo, Lubumbashi River, at Elisabethville [Lubumbashi], ca. 11°39’S, 27°28'E (1: 39.1); MRAC 183365–366, Democratic Republic of the Congo, at the bridge on Lubumbashi River, forested camp of the Kipopo, ca. 11°33’S, 27°22'E (2: 31.7–33.1); MRAC 183367–369, Democratic Republic of the Congo, Lubumbashi River, ± 25 km upstream of Elisabethville, ca. 11°33’S, 27°22'E (3: 33.9–39.2); MRAC 73025.0803, Democratic Republic of the Congo, Lubumbashi, in front of Lido, Lubumbashi River, ca. 11°39’S, 27°27'E (1: 118.8). Luapula River system: SAIAB 76705, Zambia, Mambilima Falls on Luapula River, 10°32’22.2”S, 28°39'40.3"E (1: 57.0). Lufira River system: MRAC 165341, Democratic Republic of the Congo, Gombela, Kafila River, Katanga, 10°46’31”S, 27°47'38"E (1: 60.8); SAIAB 77478, Democratic Republic of Congo, Diptera River near Fungurume, 10°36’20.9”S, 26°16'40.1"E (4: 43.5–77.0). Luongo River system: CU 91051, Zambia, Luongo River, below Musonda Dam, on road from Mansa to Kashiba (Route D79), 10°42’15.5”S, 28°48'03.6"E (2: 60.0–62.0); CU 91053, Zambia, Luongo River above pontoon south of Musonda Falls, Mansa-Serenje road (Route D235), 10°40’51.6”S, 28°43'09.1"E (2: 57.0–78.7); CU 91054, Zambia, Luongo River at bridge on Kashiba-Mwenda road (Route M3), 10°28’12.7”S, 31°01'28.2"E (3: 44.0–105.1); SAIAB 76663, same data as CU 91053 (3: 54.6–94.3); SAIAB 76670, same data as CU 91051 (1: 83.2); SAIAB 76729, same data as CU 91054 (5: 39.9–65.4); SAIAB 76925, Zambia, Luongo River at Mukonshi Bridge on Mwenda-Kawambwa road (Route M13), 10°08’39”S, 29°10'01.2"E (41: 32.2–53.8). Upper Congo (Lualaba) River drainage: SAIAB 81501, Democratic Republic of the Congo, Bona River [Tributary of Lac Delcommune (= Lac Nzilo)], Near Lenge Village, 10°36’34”S, 25°49'15"E (1: 78.8); SAIAB 81566, Democratic Republic of the Congo, large stream 1, Bona River system, 10°35’49”S, 25°53'23"E (1: 43.0); SAIAB 82855, Democratic Republic of the Congo, Kisanfu River, Upstream of Nayebe crossing, 10°48’01.1”S, 25°58'52.6"E (2: 46.8–85.4); SAIAB 82877, Democratic Republic of the Congo, Kisanfu River, just below bridge by hydroelectric station, 10°45’49.3”S, 25°57'49.3"E (6: 36.5–42.1); SAIAB 82898, Democratic Republic of the Congo, Kisanfu River, channel downstream of hydro station, 10°45’49.3”S, 25°57'49.3"E (6: 37.5–74.3).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  Amphilius frieli is diagnosed from all other species of the  Amphilius jacksonii complex by having more gill rakers on the first gill arch (10–11, rarely 9 or 12 vs. 6–9, rarely 5 or 10). It is further diagnosed from  A. jacksonii by its deeper caudal peduncle (caudal peduncle depth 9.7–11.4% SL vs. 4.8–7.9% SL), shorter caudal peduncle (caudal peduncle 14.4–16.4% SL vs. 16.7–20.6% SL), and less slender body (body depth at anus 13.9–14.9% SL vs. 9.6–13.2% SL). It is further diagnosed from  A. ruziziensis by having more branchiostegal rays (8–9 vs. 6–7), a deeper caudal peduncle (caudal peduncle depth 9.7–11.4% SL vs. 8.6–9.5% SL), shorter caudal peduncle (caudal peduncle 14.4–16.4% SL vs. 18.7–20.3% SL), and less slender body (body depth at anus 13.9–14.9% SL vs. 11.1–12.9% SL).  Amphilius frieli is further diagnosed from  A. pedunculus ,  A. crassus , and  A. lujani by its narrower interorbital width (25.1–27.7% HL vs. 28.1–35.8% HL), and from  A. crassus by having more branchiostegal rays (8–9 vs. 6–7, rarely 8), more branched pectoral-fin rays (9–10 vs. 7–8, rarely 9), and a shorter dorsal-fin insertion to adipose-fin insertion length (37.5–41.2% SL vs. 42.2–44.6% SL). It is further diagnosed from  A. lujani by its deeper caudal peduncle (caudal peduncle depth 9.7– 11.4% SL vs. 8.1–9.5% SL), shorter caudal peduncle (caudal peduncle 14.4–16.4% SL vs. 16.8–20.9% SL). </p>
            <p>Description. Morphometric data are in Table 5. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising gently from tip of snout to dorsal-fin origin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital openings located at midpoint of adpressed pelvic fin, closer to pelvic-fin insertionthan to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately joined at isthmus forming a V -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, slightly papillate. Rictal lobe large and slightly papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming crescent shaped band. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth, to pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to origin of pectoral-fin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to edge of branchiostegal membrane. Branchiostegal membrane with 8 (25), or 9 (12) rays. Gill rakers on first epibranchial 2 (1) or 3 (37); rakers on first ceratobranchial 7 (22), or 8 (15); total gill rakers on first arch 10 (22) or 11 (15).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit; covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over tip of pectoral fin. Dorsal fin with i,6 (37) rays, and fin margin straight. Pectoral fin with i,9 (37) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior of dorsal-fin insertion. Pelvic fin with i,5 (37) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base, fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, deeply incised posteriorly. Caudal fin deeply forked with tips of lobes rounded; fin with i,5,6,i (36) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,6 (2), ii,7 (2), iii,6 (24), or iii,7 (10) rays. Anal-fin margin almost straight.</p>
            <p>Coloration. Body variably mottled with dark saddles. First saddle posterior of head, second saddle at dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddle connected laterally by broad stripe. Venter light brown with fourth and fifth saddles meeting those of opposite side. Dorsal, and anal fins light brown with dark medial band. Adipose fin dark brown to black, with anterior and posterior distal edges cream-colored. Pectoral and pelvic fins positioned horizontally with upper surfaces cream-colored with dark fin base and dark medial band. Lower surfaces light yellow. Caudal fin cream-colored with dark markings on upper and lower lobes. Caudal coloration asymmetrical, with lower lobe almost completely dark (except for small cream colored mark at base of caudal and cream-colored tip). Upper lobe with less dark pigment, with dark blotch covering dorsal edge, but caudal base with large cream-colored patch and tip cream-colored.</p>
            <p>Distribution. Upper Congo River basin upstream of Kabalp, Democratic Republic of the Congo including tributaries of the Upper Congo mainstem, the Lufira River drainage, and the Kafubu, Chambeshi, Lwela, and Luongo River systems of the Luapula River drainage (Fig. 6).</p>
            <p>Etymology. Named for John P. Friel in recognition of his excellent contributions to the study of African fishes.</p>
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	https://treatment.plazi.org/id/03A71476256DFF9B2A9DE8CBFA5BF979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A714762572FF9E2A9DEAD1FA8DFA83.text	03A714762572FF9E2A9DEAD1FA8DFA83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius crassus Thomson & Hilber	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius crassus Thomson &amp; Hilber ,  new species</p>
            <p>(Fig. 8, Table 6)</p>
            <p> Amphilius platychir (non Günther).— Bailey, 1969: 192 (in part), Kilombero (BMNH 1969.2.11.192–194) [Rufiji River basin]. </p>
            <p>Holotype. UF 184237, ex. UF 170743, Tanzania, Sonjo River at bridge in Man'gula on road from Mikumi to Ifakara, altitude 302 m, Rufiji River basin, 07° 48' 29.6”S, 36° 53' 47.6"E (86.3 mm SL).</p>
            <p>Paratypes. Rufiji River basin: AMNH 258334, same data as holotype, ex. CU 93734 (10: 43.4–61.5); AUM 57571, same data as holotype, ex. CU 93734 (10: 40.7–66.8); BMNH 1969.2.11.192–194, Tanzania, Near Kiberege, trib of Kilombero River, ca. 07°57'S, 36°52'E (3: 30.8–31.7); CU 93731, Idete River at bridge in Idete on road from Ifakara to Taveta, altitude 310 m, 08°06'14”S, 36°29'17"E (7: 27.3–67.3); CU 93732, Tanzania, 12 Bridges River at overhead bridge for train on road from Mikumi to Ifakara, altitude 419 m, 07°27'53”S, 37°00'52"E (4: 49.7–72.4); CU 93734, same data as holotype, (40: 36.7–69.9); MRAC B3–06–P–7–16, same data as holotype, ex. UF 170743 (10: 43.3–60.5); SAIAB 187279, same data as holotype, ex. UF 170743 (10: 45.1–57.0); UF 170706, same data as CU 93732 (5: 47.6–80.3); UF 170729, same data as CU 93731 (8: 28.9–84.7); UF 170743, same data as holotype (40: 39.8–84.7).</p>
            <p>Non-types. Wami River basin: CU 93730, Tanzania, Divue River above &amp; below falls along road from Dumila to Turiani, altitude 374 m, 06°10'26.8”S, 37°34'59.8"E (10: 27.1–57.5); CU 93733, Tanzania, Wami River at bridge in Rudewa on road from Dumila to Kilosa, altitude 433 m, 06°40'45.2”S, 37°07'27.0"E (8: 27.1–41.2); UF 170707, same data as CU 93733 (10: 25.9–77.0); UF 170708, same data as CU 93730 (10: 40.0–59.4).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  Amphilius n. sp. . Rufiji is diagnosed from A.  jacksonii ,  A. ruziziensis ,  A. pedunculus , and  A. lujani by its shorter caudal peduncle (caudal peduncle length 13.3– 15.5% SL vs. 16.0–20.9% SL), and from  A. jacksonii ,  A. ruziziensis , and  A. lujani by its deeper caudal peduncle (caudal peduncle depth 10.1–12.0% SL vs.4.8–9.5% SL). It is diagnosed from  A. ruziziensis , A. n. sp.. Malagarasi, and  A. frieli by its longer dorsal-fin insertion to adipose-fin insertion length (42.2–44.6% SL vs. 37.5–41.7% SL).  Amphilius crassus is further diagnosed from  A. jacksonii by its deeper body (body depth at anus 14.5–17.4% SL vs. 9.6–13.2% SL), and from  A. ruziziensis by wider interorbital width (28.9–34.1% HL vs. 23.4–25.1% HL) and deeper body (body depth at anus 14.5–17.4% SL vs. 11.1–12.9% SL). It is further diagnosed from  A. lujani by having fewer branched pectoral-fin rays (7–8, rarely 9 vs. 9), and from  A. frieli by its fewer branchiostegal rays (6– 7, rarely 8 vs. 8–9), fewer branched pectoral-fin rays (7–8, rarely 9 vs. 9–10), fewer gill rakers on the first gill arch (6–8, rarely 9 vs. 10–11, rarely 9 or 12), and wider interorbital width (28.9–34.1% HL vs. 25.1–27.7% HL). </p>
            <p>Description. Morphometric data are in Table 6. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising steeply from tip of snout to dorsal-fin origin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital openings located at midpoint of adpressed pelvic fin, closer to pelvic-fin insertion than to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately joined at isthmus forming a V -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, slightly papillate. Rictal lobe large and slightly papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming crescent shaped band. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth, to pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to origin of pectoral-fin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to edge of branchiostegal membrane. Branchiostegal membrane with 6 (22), 7 (123), or 8 (2) rays. Gill rakers on first epibranchial 1 (1), 2 (122) or 3 (24); rakers on first ceratobranchial 4 (19), 5 (95), or 6 (33); total gill rakers on first arch 6 (19), 7 (79) 8 (42) or 9 (7).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit; covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over tip of pectoral fin. Dorsal fin with i,6 (147) rays, and fin margin straight. Pectoral fin with i,7 (13), i,8 (130), or i,9 (4) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior of dorsalfin insertion. Pelvic fin with i,5 (147) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base, fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, deeply incised posteriorly. Caudal fin deeply forked with tips of lobes rounded; fin with i,5,5,i (4), i,5,6,i (141) or i,6,6,i (2) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,6 (56), ii,7 (34), iii,5 (3), iii,6 (50), or iii,7 (5) rays. Anal-fin margin almost straight.</p>
            <p>Coloration. Body variably mottled with dark saddles. First saddle posterior of head, second saddle at dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddles connected laterally by broad stripe. Venter light brown with fourth and fifth saddles meeting those of opposite side. Dorsal, and anal fins light brown with dark medial band. Adipose fin dark brown to black, with anterior and posterior distal edges cream-colored. Pectoral and pelvic fins positioned horizontally with upper surfaces cream-colored with dark fin base and dark medial band. Lower surfaces light yellow. Caudal fin cream-colored with dark markings on upper and lower lobes. Caudal coloration variable but usually spotted and asymmetrical, with lower lobe more pigmented.</p>
            <p>Distribution. Rufiji and Wami river basins, eastern Tanzania (Fig. 6).</p>
            <p> Etymology. From the Latin noun  crassus (m), meaning fat or stout in reference to the deep, stout body. </p>
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	https://treatment.plazi.org/id/03A714762572FF9E2A9DEAD1FA8DFA83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
03A714762577FF9D2A9DE9C8FC32FC33.text	03A714762577FF9D2A9DE9C8FC32FC33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphilius lujani Thomson & Page	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amphilius lujani Thomson &amp; Page ,  new species</p>
            <p>(Fig. 9, Tables 7)</p>
            <p> Amphilius jacksonii .— Seegers et al., 2003: 37, Lake Victoria drainage (affluent rivers) [Nile basin] </p>
            <p>Holotype. UF 184238, ex. UF 169233, Uganda, Manafwa River at Bumwangu, Lake Kyogo drainage, Nile basin, 00°56'26.2”N, 34°16'49.2"E (104.1 mm SL).</p>
            <p>Paratypes. Lake Kyogo drainage: AUM 47147, same data as holotype (16: 37.7–112.3); AUM 47149, Uganda, Malaba River at Lwakhakha, border between Uganda and Kenya, 00°47'04”N, 34°22'44"E (7: 46.3–98.1); BMNH 1961.6.13.19, Uganda, Malawa [=Malaba] River, ca. 00°35'N, 34°03'E (1: 99.4); BMNH 1962.2.6.62–63, Uganda, Malawa [=Malaba] River, ca. 00°35'N, 34°03'E (2: 80.0–107.9); BMNH 1965.10.15.18–21, Kenya, Malikisi River, ca. 01°37'N, 34°13'E (4: 35.7–48.0); CU 97333, same data as holotype, ex. AUM 47147 (3: 50.3– 90.5); MRAC B3–06–P–17–19, same data as holotype, ex. UF 169233 (3: 56.0–88.1); SAIAB 187280, same data as holotype, ex. UF 169233 (3: 47.9–98.0); UF 169233, same data as holotype (15: 43.0–107.6); UF 169235, same data as AUM 47149 (7: 45.4–90.1).</p>
            <p>Non-types. Lake Kyogo drainage: BMNH 1965.10.15.11–13, Uganda, Sironko River, ca. 01°29'N, 34°14'E (2: 103.7–119.5); BMNH 1965.10.15.14–15, Uganda, Sironko River, ca. 01°29'N, 34°14'E (1: 101.1); BMNH 1965.10.15.16–17, Uganda, Zuzu River, tributary of the Manafwa River, ca. 01°13'N, 34°21'E (2: 40.9–42.1). Lake Manyara basin: BMNH 1969.2.20.1, Tanzania, Stream entering Lake Manyara, ca. 03°38'S, 35°41'E (1: 45.0). Nzoia River system (NE Lake Victoria tributary): SAIAB 65048, Kenya, Moi Brgidge, Little Nzoia River, ca. 00°55'N, 35°07'E (1: 74.5).</p>
            <p> Diagnosis. Diagnostic characters are summarized in Table 2.  Amphilius lujani is diagnosed from  A. pedunculus ,  A. frieli , and  A. crassus by having a more slender caudal peduncle (depth 8.1–9.5% SL vs. 9.9–12.3% SL). It is diagnosed from  A. jacksonii by having a deeper caudal peduncle (depth 8.1–9.5% SL vs. 4.8–7.9% SL) and a deeper body (body depth at anus 13.5–15.8% SL vs. 9.6–13.2% SL).  Amphilius lujani is diagnosed from  A. ruziziensis by having a wider interorbital width (28.3–31.7% HL vs. 23.4–25.1% HL) and a deeper body (body depth at anus 13.5–15.8% SL vs. 11.1–12.9% SL). It is further diagnosed from  A. frieli by having fewer gill rakers on the first gill arch (6–8, rarely 5 or 9 vs. 10–11, rarely 9 or 12) and a longer caudal peduncle (length 16.8–20.9% SL vs. 14.4–16.4% SL).  Amphilius lujani is further diagnosed from  A. crassus by having more branched pectoralfin rays (9 vs. 7–8, rarely 9) and a longer caudal peduncle (length 16.8–20.9% SL vs. 13.3–15.5% SL). </p>
            <p>Description. Morphometric data are in Table 7. Body elongate, ventral profile flattened ventrally to anal-fin base, then tapered dorsally to end of caudal peduncle. Dorsal profile rising gently from tip of snout to dorsal-fin origin, then nearly horizontal to end of caudal peduncle. Greatest body depth at dorsal-fin origin. Caudal peduncle laterally compressed, with crenellated epidermal fold. Anus and urogenital openings located at midpoint of adpressed pelvic fin, closer to pelvic-fin insertionthan to origin of anal fin. Skin smooth. Lateral line complete, extending from dorsal edge of opercular cavity to caudal-fin base.</p>
            <p>Head and anterior part of body depressed and broad. Head wedge-shaped in lateral view. Snout broad, blunt when viewed from above. Head becoming wider from tip of snout to pectoral-fin base. Branchiostegal membranes moderately joined at isthmus forming a V -shaped connection.</p>
            <p>Mouth broad, gently curved, subterminal. Lips moderately fleshy, strongly papillate. Rictal lobe large and papillate. Anterior portion of premaxillary tooth band exposed with mouth closed. Premaxillary tooth patches joined, forming crescent shaped band. Premaxillary and dentary teeth short, conical. Dentary tooth patches forming U -shaped band, separated medially.</p>
            <p>Three pairs of simple, tapered circumoral barbels. Maxillary barbel large, fleshy and flattened with pointed tip; barbel extending posterolaterally from corner of mouth, to pectoral-fin base. Outer mandibular barbel thin with pointed tip, origin at posterior corner of lower jaw, extending to origin of pectoral-fin. Inner mandibular barbel originates anterolaterally of inner mandibular barbel, extending to edge of branchiostegal membrane. Branchiostegal membrane with 7 (21), or 8 (25) rays. Gill rakers on first epibranchial 2 (36) or 3 (10); rakers on first ceratobranchial 4 (2), 5 (22), 6 (1), or 7 (1); total gill rakers on first arch 6 (2), 7 (16) 8 (23) or 9 (5).</p>
            <p>Eye small, positioned dorsolaterally approximately midway between tip of snout and posterior margin of operculum. Horizontal diameter of eye slightly wider than vertical diameter. Eye without free orbit; covered with skin confluent with dorsal surface of head. Anterior and posterior nares with prominent tubular rims; nares separate but relatively close to each other. Posterior nare located about midway between eye and tip of snout.</p>
            <p>Dorsal-fin origin at point over tip of pectoral fin. Dorsal fin with i,6 (45) or i,7 (1) rays, and fin margin straight. Pectoral fin with i,9 (46) rays; unbranched ray greatly thickened. Pectoral fin with four or five innermost rays progressively shorter making posterior fin margin rounded. Origin of pelvic fin posterior of dorsal-fin insertion. Pelvic fin with i,5 (46) rays with first ray unbranched and greatly thickened. Pelvic fin with straight posterior margin.</p>
            <p>Adipose-fin base longer than anal-fin base, origin anterior to origin of anal-fin base, fin extending past anal-fin insertion. Margin strongly convex with sharply rounded edge, deeply incised posteriorly. Caudal fin deeply forked with tips of lobes rounded; fin with i,5,6,i (46) principal rays. Anal fin with short base, origin posterior to origin of adipose-fin base, with ii,6 (10), ii,7 (12), iii,6 (19), or iii,7 (6) rays. Anal-fin margin almost straight.</p>
            <p>Coloration. Body variably mottled with dark saddles. First saddle posterior of head, second saddle at dorsal fin, third saddle between dorsal and adipose fins, fourth saddle under anterior part of adipose fin, and fifth saddle on caudal peduncle. All saddles connected laterally by broad stripe. Venter light brown with fourth and fifth saddles meeting those of opposite side. Dorsal, and anal fins light brown with faint medial bands. Adipose fin dark brown to black, with posterior distal edge cream-colored. Pectoral and pelvic fins positioned horizontally with upper surfaces cream-colored with darkened fin base and faint medial bands. Lower surfaces light yellow. Caudal fin cream-colored with dark markings on upper and lower lobes. Caudal coloration asymmetrical, with lower lobe almost completely dark (except for small cream-colored mark at base of caudal and cream-colored tip). Upper lobe with less dark pigment, with dark blotch covering dorsal edge, but caudal base with large cream-colored patch and tip cream-colored.</p>
            <p>Distribution. Lake Kyogo drainage, western Uganda; Nzoia River, a northeastern tributary of Lake Victoria, and Lake Manyara basin, Tanzania (Fig. 6).</p>
            <p>Etymology. Named for Nathan K. Lujan who collected the holotype and most paratypes of this species, and who has made excellent contributions to our knowledge of freshwater fishes.</p>
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	https://treatment.plazi.org/id/03A714762577FF9D2A9DE9C8FC32FC33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Thomson, Alfred W.;Page, Lawrence M.;Hilber, Samantha A.	Thomson, Alfred W., Page, Lawrence M., Hilber, Samantha A. (2015): Revision of the Amphilius jacksonii complex (Siluriformes: Amphiliidae), with the descriptions of five new species. Zootaxa 3986 (1): 61-87, DOI: 10.11646/zootaxa.3986.1.3
