taxonID	type	description	language	source
039987C66546FF9F7431C28AFB3AF992.taxon	materials_examined	Type species: Dilophotes exilis Waterhouse 1878: 116 (by original designation). Stenolycus Ohbayashi, 1956: 58, syn. n. Type species: Stenolycus ohirai Ohbayashi, 1956: 58 (by original designation).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66546FF9F7431C28AFB3AF992.taxon	diagnosis	Diagnosis. Dilophotes, together with Macrolycus Waterhouse, 1878, forms the tribe Macrolycini (Lycinae). Macrolycini are easily recognised by cleft claws and the structure of pronotal carinae (Figs 6 ­ 8). These genera differ externally in the number of longitudinal elytral costae. Dilophotes has costa 1 considerably shorter than costa 2 and 4, costa 3 is vestigial and distinct only at the very base of elytra. Costae 2 and 4 reach the apical part of elytra (Fig. 3). Macrolycus has four longitudinal costae on each elytron, although some of them may be very weak, especially apically. Dilophotes often have a considerably sclerotized and exposed internal sacs, which sometimes form a very complex structure (Figs 12, 15, 18, 20). In addition, the phallobase of Dilophotes is long and asymmetrical (Figs 14, 16, 22). Macrolycus has a simple, slender, membranous internal sac and at most its apical part is exposed; its phallobase is small and rounded Bocak and Bocakova (1990). Redescription. Body small to medium sized, length 5.1 ­ 13.2 mm, mostly 6.0 ­ 8.0 mm, parallel­sided to slightly widened posteriorly, moderately dorsoventrally depressed, weakly sclerotized. Many species brightly coloured, with dark red to orange yellow pronotum and elytra. Only few species concolorous black. Head hypognathous, small, slightly transverse, with short, conical rostrum. Eyes small, almost regularly smaller than their frontal interocular distance, hemispherically prominent, seldom with maximum eye diameter considerably larger than interocular distance. Frons concave, with deep depression medially, antennal tubercles flat, but distinct. Clypeus widely concave, labrum transverse, emarginate apically. Antennae slender, compressed, reaching to two thirds the length of elytra, either uniform in sexes or male antennae flabellate and female antennae weakly serrate. Flabellae of antennomeres 3 ­ 10 up to four times longer than trunks of antennomeres. Mandibles elongate, slender, curved, their inner margins simple. Maxillae with long setose galea. Maxillary palpi 4 ­ segmented, basal palpomere very small, triangular, palpomere 2 almost three times longer than basal one, palpomere 3 shorter, apical palpomere robust, as long as wide, securiform or parallel­sided. Labial palpi 3 ­ segmented, considerably shorter, basal palpomeres minute, their combined length slightly longer than those of apical palpomeres, which resembles in shape those of maxillary palpi. Pronotum flat, with only short longitudinal pronotal keel present anteriorly, sometimes with a transverse mound present at middle of pronotum. Frontal margin usually widely rounded, frontal angles inconspicuous, lateral margins straight or slightly concave, posterior angles projected, sometimes very acutely. Disc rugose at frontal and lateral margins, shiny in middle and at base. Scutellum small, simply rounded at posterior margin. Elytra flat, slightly sclerotized, with 3 distinct longitudinal costae in anterior half of elytra. Costa 1 regularly shortened, seldom reaching beyond half of elytra, costae 2 and 4 complete, running entire length of elytra, costa 3 vestigial, scarcely distinct at base of elytra. Transverse costae absent, interstices finely rugose, densely pubescent. Legs slender, compressed, claws cleft apically. Male genitalia consisting of phallobase, phallus, and internal sac (Fig. 14). Parameres absent. Phallobase long, slender, asymmetrical, phallus mostly slender, simple. Internal sac often completely sclerotized and in some cases consisting of a pair of basal rods and lower and upper plate (Figs 12, 15). Female genitalia are characterized by elongate styli and relatively short coxites. Valvifers fused with coxites and bases of coxites connected by bridge (Figs 10, 21, 24).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66546FF9F7431C28AFB3AF992.taxon	description	Sexual dimorphism. There is only very slight sexual dimorphism in body shape and size. Males and females differ only in several species in the shape of antennae. Dilophotes ohirai (Ohbayashi, 1956), D. vandykei (Nakane, 1970), D. luteus sp. n., D. anthracinus sp. n., and D. lizipingensis sp. n. have flabellate antennae in males (Fig. 4) and very slightly serrate antennae in females (Fig. 5). Most species have relatively small eyes without any substantial sexual dimorphism. Only males of D. luteus sp. n. and D. kubani sp. n. have very large eyes. Unlike many other genera, Dilophotes females have large eyes similar to those of the males.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66546FF9F7431C28AFB3AF992.taxon	discussion	Remarks. Waterhouse (1878) presented a description of the genus without proposing its formal name. The paragraph was headed " Genus 38 " and the single included species was Lycus exilis. Waterhouse (1879) then named this genus Dilophotes and included two species in it: Dilophotes exilis (Waterhouse, 1978) and D. pygmaeus Waterhouse, 1879. In this valid description he designated D. exilis as the type species of the genus.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66546FF9F7431C28AFB3AF992.taxon	distribution	Distribution. Dilophotes occurs in the whole Oriental Region and in he eastern part of the Palaearctic Region. Unlike its relative Macrolycus, its range does not reach into Northern China or the Russian Far East. This revision shows that a very high diversity of Dilophotes exists in the tropical forests of northern Laos and it indicates the restricted ranges of most known species. The distribution of species is shown in Figs 1 ­ 2. Ecology. The Chinese Dilophotes species were collected from lower mountain forests up to the high mountain forests in Northern Sichuan and Southern Shaanxi. Similarly, the species from Bhutan were collected in high mountains. Adults were slow moving or sitting on leaves of shrubs and trees in clearings in montane forests. Therefore most specimens were collected by sweeping or as individuals. The Chinese species D. atrorufus was collected in numbers on flowering Castanopsis. Larvae of Dilophotes are unknown, but development in rotting wood is expected as in the related genus Macrolycus. The Indochinese species were collected in habitats from lowland to lower mountains and they were often collected in the lower strata of dense forest canopy (L. Bocak, personal communication). Lycids are protected from predators by the presence of several odiforous and bitter compounds (Moore & Brown, 1981), and therefore have a very strong tendency to form mimicry complexes. I have found that the Palaearctic species are dark red to reddish brown, similar to the other lycids that occur in the same habitat (Macrolycus, Calochromus Guérin­Méneville, 1833 and Plateros Bourgeois, 1879). The Bhutanese species Dilophotes bhutanensis sp. n. is similar in the body size, shape and colouration the sympatrically occurring Macrolycus bowringi Waterhouse, 1878, unlike the second sympatrical species, D. holzschuhi sp. n. which is similar to the much smaller, dark red coloured Plateros species. Similarly, I found that in Laos's fauna the lightly coloured species are similar to some Cautires Waterhouse, 1879 and Xylobanus Waterhouse, 1879 as well as the dark red species of some Plateros and Conderis Waterhouse, 1879. Species level relationship. Dilophotes species are, in many cases, hardly distinguishable when comparing external characters. The body colouration is usually the same in several sympatrically occurring species. I found the principal variation in the shape of male genitalia, the shape of antennae (Figs 4, 5), and the relative size of the eyes. Unfortunately, only few specimens were collected in copula or under conditions where I could associate males and females of the respective species. Therefore, the characters based on female genitalia were not been used for the study of species level relationships. Based on the structure of the male genitalia I have distinguished several species groups. The shape of phallus and the degree of sclerotization and the shape of the internal sac are very diversified in Dilophotes. Unfortunately, the only related genus, Macrolycus, has a very different type of phallus and therefore I was not able to homologize the characters found in the male genitalia. The eyes are almost regularly considerably smaller than their interocular distances. Only two species, D. luteus sp. n. and D. kubani sp. n. have very large eyes. These species differ in both the shape of their antennae and the type of their male genitalia. Therefore, the size of eyes is not indicative of their relationships. Two types of male antennae were found in Dilophotes. Dilophotes ohirai (Ohbayashi, 1956), D. vandykei (Nakane, 1970), D. luteus sp. n., D. anthracinus sp. n. and D. lizipingensis sp. n. have flabellate antennae in males, the remaining species have very similar slender filiform to serrate antennae in both sexes. Until now, all the species with flabellate antennae were placed in Flabellodilophotes Pic, 1912 (Dilophotellus Kleine, 1925) or Stenolycus Ohbayashi, 1956. I have found that the presence and / or lack of lamellae on antennae did not correlate with the shape of the male genitalia. Flabellodilophotes is a polyphyletic, typologically based genus and can not be accepted in the classification. I examined some Oriental representatives of Flabellodilophotes and both serrate and flabellate male antennae were found for example in the D. moxiensis group, which is defined in this article (e. g. D. costatus Kleine, 1930). As I did not study the type species of Flabellodilophotes, I do not formally synonymize it with Dilophotes. On the other hand, species with flabellate antennae from the studied area are here classified in Dilophotes. Nakane (1969) redescribed the Japanese species Stenolycus ohirai in detail, which is a type species of Stenolycus. I compared the male genitalia with those of species occurring in continental Asia and I have not found any principal differences between them. Stenolycus ohirai belongs to the Dilophotes anthracinus group and I propose Stenolycus to be a junior synonym of Dilophotes. Kasantsev (2000) described a new subgenus Biphylodes as a Dilophotes without elytral costa 1. The male genitalia of the type species are similar to those of D. moxiensis sp. n. and D. pacholatkoi sp. n. These species have well developed costa 1 as all other Dilophotes. Therefore, Biphilodes as a subgenus is ill based and refers only to one species within D. moxiensis group as defined further. Additionally, some species now classified by Kleine and Pic in Flabellodilophotes Pic, 1912 have the same type of aedeagus and when the male of the Flabellodilophotes species is found, Biphilodes Kasantsev, 2000 can be a junior synonym of Flabellodilophotes Pic, 1912 or Dilophotellus Kleine, 1925.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66541FF9F7431C0D7FB3CF884.taxon	diagnosis	Diagnosis. The Dilophotes atricollis group is characterized by serrate male antennae and a slender phallus with a characteristicaly sclerotized internal sac (Figs 9, 11 ­ 15).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF9E7431C75FFC43FB64.taxon	diagnosis	Diagnosis. Dilophotes atricollis has a characteristic colour pattern. It is the only species of Dilophotes with a dark brown to black pronotum and brightly orange red elytra. The male genitalia show a close relationship with the other species from the D. atricollis group and especially with D. berezowskii, from which it differs in colouration and by its relatively larger eyes. Redescription. Body small to medium sized, slender, black. Pronotum black, covered with dense light brown pubescence, elytra orange red, with dense red pubescence. Head with short rostrum. Eyes small, interocular distance 1.33 times maximum eye diameter. Antennae serrate. Pronotum 1.5 times wider than long, frontal margin rounded, frontal angles inconspicuous. Lateral margins straight, posterior angles slightly projected. Disc with sharp longitudinal costa reaching from middle of frontal margin to middle of disc, keel inconspicuous basally. Scutellum slightly emarginate apically. Elytra slender, each elytron with three longitudinal costae. Costa 1 inconspicuous in apical part. Male genitalia with complex, well sclerotized internal sac, very similar to those of D. berezowskii.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF9E7431C75FFC43FB64.taxon	description	Measurements. BL 7.0 mm, WH 1.43 mm, PL 0.88 mm, PW 1.32 mm, Ediam 0.36 mm, Edist 0.48 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF9E7431C75FFC43FB64.taxon	materials_examined	Material examined. 1 male, " Taiwan, Fenchihu, 1400 m, J. A S. Klapperich leg. "; 1 female, " Taiwan, Tapang (1800), Taichung­hsien, 27.7. 1983, T. Hattori leg. "; l male " Taiwan, Alishan, 2 400 m, 10. 6. 1977, J. & S. Klapperich leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF9E7431C75FFC43FB64.taxon	distribution	Distribution. Dilophotes atricollis is endemic to Taiwan.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF917431C35DFC2CFD4A.taxon	diagnosis	Diagnosis. Dilophotes atrorufus is the only Japanese species with serrate antennae in the male and has a characteristic slightly curved apical part of phallus and a smaller but well sclerotized internals sac (Fig. 11). Although the upper and lower plates of internal sac are not distinctly separated, the species is placed in the D. atricollis group.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF917431C35DFC2CFD4A.taxon	description	Description. Body medium sized, slender, black. Pronotum black with brown pubescence, elytra slender with reddish brown pubescence. Head with short rostrum. Eyes small, interocular distance 1.63 times maximum eye diameter. Antennae serrate, considerably flattened. Maxillary palpi slender. Pronotum 1.4 times wider than long, frontal margin slightly emarginate anteriorly, anterior angles distinct, lateral margins slightly concave, posterior angles projected. Pronotal disc with sharp costa reaching slightly beyond middle of pronotal disc. Scutellum simply rounded apically. Each elytron with three longitudinal costae, costa 1 inconspicuous in apical half. Male genitalia slender, long, phallus distinctly curved apically, internal sac complex, sclerotized, upper and lower plates fused (Fig. 11). Measurements. BL 9.0 mm, WH 1.98 mm, PL 1.21 mm, PW 1.65 mm, Ediam 0.40 mm, Edist 0.65 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF917431C35DFC2CFD4A.taxon	materials_examined	Material examined. 1 male, 1 female, " Japon, Yumoto, pres Nikko, 6 Aout, 1911, Edme Gallois leg. "; 1 specimen, " Japonia, 24. 6. 1958, Mt. Iwawaki, Sh. Mizobe lgt. "; 1 male, 1 female, " Tokyo near Kamikochi, Japan, 11. 8. 1950, T. Nakane lgt. "; 1 specimen, " Japon, Chuzenji, 2. 8. 1911, Edme Gallois lgt. "; 1 specimen, " Japon, Yumoto pres Nikko, 11 Aout, 1911, Edme Gallois " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66540FF917431C35DFC2CFD4A.taxon	distribution	Distribution. Dilophotes atrorufus is endemic to Japan.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654FFF907431C505FD15FCC2.taxon	diagnosis	Diagnosis. Dilophotes berezowskii has male genitalia very similar to that of D. atricollis. These species differ in the pronotal colouration. D. berezowskii is a very widespread species and specimens from different elevations differ in the shade of red body parts, but the colouration of pronotum and elytra is always very similar. Redescription. Body medium­sized, slender, black, pronotum and elytra red to dark brown, with dense red pubescence. Head with short rostrum, frons densely pubescent. Eyes small, interocular distance 1.64 times maximum eye diameter. Antennae slightly serrate, terminal antennomeres almost parallel­sided. Maxillary palpi slender. Pronotum 1.2 times wider than long, frontal margin slightly emarginate in middle, frontal angles inconspicuous, lateral margins very slightly concave, posterior angles moderately projected (Fig. 7). Pronotal disc with sharp longitudinal costa attached to frontal margin and reaching middle of pronotal disc, keel inconspicuous posteriorly. Pronotal disc surface densely pubescent, shiny. Posterior margin of scutellum straight. Elytra slender, flat, slightly widened posteriorly, each elytron with three longitudinal costae, costa 1 very weak, reaching three quarters of elytral length, costa 3 very strong basally. Male genitalia with phallus slender, internal sac complex, sclerotized (Fig. 9).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654FFF907431C505FD15FCC2.taxon	description	Measurements. BL 8.0 mm, WH 1.76 mm, PL 1.15 mm, PW 1.43 mm, Ediam 0.34 mm, Edist 0.56 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654FFF907431C505FD15FCC2.taxon	materials_examined	Material examined. 4 males, 2 females, " China, Abazhou, Nanping, Jiuzhaigou, 2000 m, 33.15 ° N, 104.15 ° E, 8. ­ 13. 6. 1990 ", 3 males, " China, Shaanxi, Quinling Shan, 35 km S of Baoji, 21. ­ 23. 6. 1998 "; 2 males, " Yunnan, Habashan mts., SE slope, 27.20 ° N, 100.11 ° E, 2000 ­ 3800 m, 10. ­ 13. 7. 1992 "; 2 males, " China, W Hubei prov., Dashennongjia Nat. Res., Muyu, E slope, 2000 m, 12. ­ 16. 6. 1997 "; 1 male, " China, Shaanxi prov., Qing Ling Shan mts., road Baoji, Taibai vill., 21. ­ 26. 6. 1998 "; 1 male, " China, Yunnan prov., Jinsha riv., Daju, 27.18 ° N, 100.13 ° E, 15. ­ 17. 7. 1990 "; 1 male, " China, N Yunnan, Yulongshan mts., E slope, 3200 m, 27.13 ° N, 100.16 ° E, 14. 7. 1990 "; 1 male, " China, Yunnan, Hengduan mts., 28.06 ° N, 98.54 ° E, 5. ­ 8. 7. 1996 "; 1 male, " Yunnan, Yulongshan, 3300 m, 27.07 ° N, 100.14 ° E, 9. 7. 1996 "; 1 male, " China, NW Sichuan, 30 km NW Muli, Bowa, mixed forest, 3500 m, 28.07 ° N, 101.05 ° E, 24. 7. 1995 " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654FFF907431C505FD15FCC2.taxon	distribution	Distribution. berezowskii is a widely distributed species in China (Sichuan, Shaanxi, Hubei, Yunnan).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654FFF907431C505FD15FCC2.taxon	discussion	Remark. There exists the considerable variability in the colouration of pronotum and elytra. Specimens from lower elevations in Sichuan (about 2000 m a. s. l.) are reddish. Specimens from higher elevations in Yunnan, humid mountains in Shaanxi Province, and the Shennongjia mountains are dark red. Kasantsev (2000) described this species according to two specimens from one locality, therefore he did not mentioned any variability and was able to distingiush this species from D. diversipennis Pic, 1912.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654EFF937431C5FFFB1AFE22.taxon	materials_examined	Type material. Holotype, male, " W­Bhutan, Thimphu Distr., E of Dochu­la, Menshunang, 2400 m, 7. 7. 1988, C. Holzschuh leg. " (LMBC). Paratypes: 2 females, same locality data (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654EFF937431C5FFFB1AFE22.taxon	diagnosis	Diagnosis. Dilophotes bhutanensis sp. n. is the only Himalayan Dilophotes species with a light orange pronotum and elytra. The structure of internal sac indicates a close relationship to D. atricollis and it differs from species with similar male genitalia by its straight, dentate part of lower plate (Fig. 12).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654EFF937431C5FFFB1AFE22.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum and elytra bright orange, covered with dense orange pubescence. Head with short rostrum. Eyes small, interocular distance 1.41 times maximum eye diameter. Antennae slightly serrate. Maxillary palpi slender. Pronotum 1.33 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins almost straight, posterior margin strongly convex in middle and posterior angles acutely projected. Pronotal disc surface light orange, shiny, with dense, brightly orange pubescence. Disc with longitudinal costa reaching middle of disc, then inconspicuous. Scutellum black with dark pubescence, slightly emarginate posteriorly. Elytra slender, flat, each elytron with three longitudinal costae. Costa 1 reaching to middle of elytral length, absent apically. Male genitalia slender, with complex, sclerotized internal sac. Lower plate apically dentate, dentate margin straight (Fig. 12). Measurements. BL 8.5 mm, WH 1.81 mm, PL 1.15 mm, PW 1.54 mm, Ediam 0.40 mm, Edist 0.56 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654EFF937431C5FFFB1AFE22.taxon	distribution	Distribution. Dilophotes bhutanensis sp. n. in known at present only from the type locality in Bhutan.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654EFF937431C5FFFB1AFE22.taxon	etymology	Etymology. The specific epithet refers to the holotype locality in Bhutan.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF937431C79DFD2AFA7F.taxon	materials_examined	Type material. Holotype, male, " W­Bhutan, Thimphu Distr., E of Dochu­la, Menshunang, 2400 m, 7. 7. 1988, C. Holzschuh leg. " (LMBC). Paratype: 1 male, " NE India, Menghalaya, 3 km E of Tura, 500 ­ 1150 m, 23.30 ° N, 90.14 ° E, 1. ­ 8. 5. 1999, Dembický & Pacholátko leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF937431C79DFD2AFA7F.taxon	diagnosis	Diagnosis. The internal sac is similar to that of D. qinlinganus, but the upper plate is smaller and straight dorsally (Fig. 13). Externally, D. holzschuhi sp. n. is reminiscent of the dark coloured species from southern China and Laos, which belong to different species groups.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF937431C79DFD2AFA7F.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum light reddish brown, elytra red, both covered with dense red pubescence. Head with short rostrum. Eyes small, interocular distance 1.45 times maximum eye diameter. Antennae very slightly serrate, subapical antennomeres almost parallel­sided. Maxillary palpi slender. Pronotum 1.27 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins almost straight, posterior margin weakly convex in middle. Disc with sharp longitudinal costa anteriorly. Scutellum black. Elytra slender, flat, each elytron with three longitudinal costae. Costa 1 inconspicuous in apical half. Male genitalia slender, long, both lower and upper plates of internal sac small, dorsal margin of upper plate straight basally (Fig. 13). Measurements. BL 1.54 mm, WH 1.48 mm, PL 0,99 mm, PW 1.26 mm, Ediam 0.36 mm, Edist 0.53 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF937431C79DFD2AFA7F.taxon	distribution	Distribution. Dilophotes holzschuhi sp. n. occurs in mountainous areas in Bhutan and in south eastern India at Bangladesh border.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF937431C79DFD2AFA7F.taxon	etymology	Etymology. The specific epithet is a patronym in honour of Carolus Holzschuh (Vienna), a collector of the holotype.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF957431C07AFAB7FC2A.taxon	materials_examined	Type material. Holotype, male, " Laos, Saisombun zone, Phou Khao Khouay N. P., Tad Leuk, 18.23 ° N, 103.04 ° E, 150 m, 20. 5. 2001, E. Jendek & O. Šauša leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF957431C07AFAB7FC2A.taxon	diagnosis	Diagnosis. Dilophotes laosensis sp. n. is externally very similar to D. holzschuhi sp. n. and other dark red coloured species with small bodies. It differs by the relatively small lower plate of internal sac and the widely curved dorsal margin of upper plate (Fig. 14).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF957431C07AFAB7FC2A.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum dark brown with light brown lateral margins, covered with light brown pubescence, elytra brightly red, with light red pubescence. Head with short rostrum. Eyes small, interocular distance 1.41 times maximum eye diameter. Antennae slightly serrate. Maxillary palpi slender. Pronotum 1.25 times wider than long. Frontal margin considerably projected, very slightly emarginate in middle, lateral margins almost parallel­sided anteriorly, distinctly divergent in posterior half, posterior angles slightly projected, posterior margin weakly convex medially. Pronotal disc with sharp longitudinal costa attached to middle of anterior margin. Scutellum simply rounded posteriorly. Elytra very slender, flat, each elytron with three longitudinal costae. Costa 1 inconspicuous in apical part. Male genitalia slender, internal sac sclerotized, complex, lower plate relatively small (Fig. 14). Measurements. BL 6.0 mm, WH 1.45 mm, PL 0.88 mm, PW 1.1 mm, Ediam 0.34 mm, Edist 0.48 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF957431C07AFAB7FC2A.taxon	distribution	Distribution. Dilophotes laosensis sp. n. is known only from a lower mountain locality in the vicinity of Vientiane.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654DFF957431C07AFAB7FC2A.taxon	etymology	Etymology. The specific epithet refers to the country, where the species was collected.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	materials_examined	Type material. Syntypes, male, female, " Indochina " (MHNP).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	diagnosis	Diagnosis. Dilophotes particularis belongs to the D. atricollis group on the basis of the shape of male genitalia. It differs externally by the red margins and infuscated middle part of pronotum.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	description	Description of the male syntype. Body medium sized, slender, black. Pronotum light reddish brown, with dense light brown pubescence, elytra red, covered with red pubescence. Head with short rostrum. Eyes small, interocular distance 1.39 times maximum eye diameter. Maxillary palpi slender. Antennae serrate, moderately robust. Pronotum 1.4 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins slightly concave, posterior angles slightly projected. Posterior margin weakly convex in middle. Pronotal disc with sharp longitudinal costa attached to frontal margin reaching beyond middle, inconspicuous at posterior margin. Disc surface densely pubescent, shiny. Scutellum with slightly emarginate posterior margin. Elytra slender, flat, each elytron with three longitudinal costae, costa 1 very weak in apical half. Male genitalia with long slender phallus and complex, sclerotized internal sac, very similar to that of D. berezowskii. Measurements. BL 7.0 mm, WH 1.65 mm, PL 0.99 mm, PW 1.43 mm, Ediam 0.36 mm, Edist 0.51 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	materials_examined	Other material examined. 1 male, " Nepal, Kathmandu, Godavari, 21. ­ 27. 6. 1989, C. Holzschuh lgt. "; 1 female, " N Vietnam, Tam Dao, 900 – 1400 m, 3. ­ 11. 6. 1988, J. Jelinek leg. "; 1 female, " N Vietnam (Tonkin), pr. Vinh Phu, Tam Dao, 6. ­ 9. 5. 1990, Vit Kubáò leg. "; 1 specimen " N Vietnam (Tonkin), pr. Vinh Phu, Tam Dao, 6. ­ 9. 5. 1990, L. Dembický leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	distribution	Distribution. Dilophotes particularis is known from several specimens collected in northern Vietnam and one specimen found in the Kathmandu Valley in Nepal. The exact collection locality of the type series is unknown, but I have studied specimens collected recently in Tonkin (Tam Dao), Vietnam and these specimens are very similar to syntypes.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654BFF947431C5AFFB00FC22.taxon	discussion	Remark. D. qinlinganus was thoroughly described by Kasantsev (2000). It differs from close D. holzschuhi sp. n. in the shape of upper plate of internal sac (Fig. 13).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654AFF977431C5A7FE23FEFA.taxon	materials_examined	Type material. Holotype, male, " Laos, 15 km NW Louang Namtha, 21.07 ° N / 101.21 ° E, alt. 750 + 100 m, 13. ­ 24. 5. 1997, E. Jendek & O. Šauša leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654AFF977431C5A7FE23FEFA.taxon	diagnosis	Diagnosis. Dilophotes sausai sp. n. belongs to the group of dark coloured species with small, slender bodies. The dentate margin of lower plate of the internal sac is similar to D. bhutanensis sp. n., but unlike that species the dentate part is widely rounded as in D. sausai sp. n.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654AFF977431C5A7FE23FEFA.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum light brown with brown pubescence, elytra brownish red, covered with red pubescence, each elytron with longitudinal infuscation in basal half. Head with short rostrum. Eyes small, interocular distance 1.43 times maximum eye diameter. Antennae slightly serrate. Maxillary palpi slender. Pronotum 1.33 times wider than long. Frontal margin widely rounded, frontal angles distinct, lateral margins concave, posterior angles slightly projected, posterior margin rounded. Pronotal disc with sharp, slender, longitudinal costa, rugose at frontal margin, shining at base, middle of disc dark brown, with light red pubescence. Elytra slender, flat, each elytron with three longitudinal costae. Costa 1 inconspicuous in apical quarter. Internal sac of male genitalia well sclerotized, lower plate with dentate, rounded margin (Fig. 15). Measurements. BL 6.0 mm, WH 1.32 mm, PL 0.82 mm, PW 1.1 mm, Ediam 0.35 mm, Edist 0.51 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654AFF977431C5A7FE23FEFA.taxon	distribution	Distribution. Dilophotes sausai sp. n. is known only from the type locality in northernmost Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C6654AFF977431C5A7FE23FEFA.taxon	etymology	Etymology. The specific epithet is a patronym in honour of Ondrej Šauša, a collector of the species.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF977431C7FFFA82FD1D.taxon	discussion	Remark. D. tibetanus was recently thoroughly described by Kasantsev (2000). This species is characteristic in the simple lower plates of internal sac. The close species, D. bhutanensis sp. n. and D. sausai sp. n. have lower plates with dentate margins (Figs 12, 15).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF967431C26DFEF4FEFA.taxon	materials_examined	Type material. Holotype, male, " Laos, Attapu prov., Bolaven Plateau, 15 km SE of Ban Houaykong, Nong Lom lake env., 15.02 ° N, 106.35 ° E, alt. 800 m, 18. ­ 30. IV. 1999, E. Jendek & O. Šauša leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF967431C26DFEF4FEFA.taxon	diagnosis	Diagnosis. Dilophotes bolavensis sp. n. is externally very similar to the all dark red coloured species. It has a plate­shaped internal sac and slender phallus. This differs from similar male genitalia of D. jendeki sp. n. in the much higher ratio between the length of internal sac and phallus (Figs 16, 17).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF967431C26DFEF4FEFA.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum and elytra with red pubescence. Head with short rostrum. Interocular distance 1.19 times maximum eye diameter. Antennae slightly serrate. Pronotum 1.30 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins slightly concave, posterior angles moderately projected. Pronotal disc with longitudinal costa reaching middle, then inconspicuous. Disc surface black, shiny, with light pubescence. Elytra slender, flat, each elytron with three longitudinal costae. Costae 1 and 3 weak in apical part. Male genitalia with long slender phallus, phallus five times length internal sac, internal sac small, plateshaped (Fig. 16). Measurements. BL 5.25 mm, WH 1.21 mm, PL 0.71 mm, PW 0.93 mm, Ediam 0.40 mm, Edist 0.47 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF967431C26DFEF4FEFA.taxon	distribution	Distribution. D. bolavensis sp. n. is known only from the type locality.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66549FF967431C26DFEF4FEFA.taxon	etymology	Etymology. The specific epithet is derived from the local name of the Bolaven Plateau in Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C7FFFEC7FD45.taxon	discussion	Remark. This species belongs according to the type of male genitalia to the D. moxiensis group and differs from other species of the group in the absent elytral costa 1. The status of the subgenus Biphilodes was discussed above in frame of the relationship within the whole genus.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C57DFE1EF89F.taxon	materials_examined	Type material. Holotype, male, " Laos, Attapu prov., Bolaven Plateau, 15 km SE of Ban Houaykong, Nong Lom lake env., 15.02 ° N, 106.35 ° E, alt. 800 m, 18. ­ 30. IV. 1999, E. Jendek & O. Šauša leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C57DFE1EF89F.taxon	diagnosis	Diagnosis. Dilophotes jendeki sp. n. resembles D. bolavensis sp. n. in both general appearance and the shape of male genitalia. These species differ in the relative size of internal sac (Figs 16, 17).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C57DFE1EF89F.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum dark brown to black, with light brown pubescence, elytra dark brown, very slightly infuscate at humeri, covered with dark red pubescence. Head with short rostrum. Eyes small, interocular distance 1.36 times maximum eye diameter. Maxillary palpi slender. Antennae slightly serrate, slightly surpassing elytral midlength, more slender apically. Pronotum 1.4 times wider than long, frontal margin round, lateral margin slightly concave, almost parallelsided in anterior three quarters, posterior angles acutely projected. Posterior margin distinctly convex medially. Pronotal disc with sharp longitudinal costa attached to frontal margin and reaching middle, then inconspicuous. Disc surface pubescent, shiny. Scutellum emarginate apically. Elytra slender, each elytron with three longitudinal costae, costa 1 inconspicuous in apical part. Male genitalia with slender phallus, three times longer than internal sac, internal sac large, complex, sclerotized (Fig. 17). Measurements. BL 5.5 mm, WH 1.4 mm, PL 0.82 mm, PW 1.15 mm, Ediam 0.36 mm, Edist 0.50 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C57DFE1EF89F.taxon	distribution	Distribution. Dilophotes jendeki sp. n. is known only from the type locality in southernmost Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66548FF967431C57DFE1EF89F.taxon	etymology	Etymology. The specific epithet is a patronym in honour of Eduard Jendek, a collector of the type.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF897431C75FFD82FABA.taxon	materials_examined	Type material. Holotype, male, " China, Moxi, east of Gongashan, 2700 m, 20. ­ 24. vii. 1992 " (LMBC). Paratypes: 5 males, 2 females, same locality data (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF897431C75FFD82FABA.taxon	diagnosis	Diagnosis. Dilophotes moxiensis sp. n. resembles D. pacholatkoi sp. n. in the shape of male genitalia (Figs 18 ­ 20). These species differ in the shape of the apical ring, which is elliptical in D. moxiensis sp. n. (Fig. 18) and rounded in D. pacholatkoi sp. n. (Fig. 20). These species differ also in colouration of pronotum and elytra.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF897431C75FFD82FABA.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum and elytra black, only their margins red, both covered with dense, dark red to purple pubescence. Head with short rostrum. Eyes very small, interocular distance 1.64 times maximum eye diameter. Antennae very slender, reaching beyond two thirds of elytra, basal antennomeres slightly serrate, terminal ones parallel­sided. Maxillary palpi slender, apical palpomere 1.7 times longer than wide. Pronotum 1.5 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins slightly concave, posterior angles moderately projected (Fig. 6). Pronotal disc with sharp longitudinal costa attached to frontal margin and reaching middle of pronotum. Disc surface densely pubescent, shiny. Scutellum simply rounded. Elytra very slender, flat, each elytron with three longitudinal costae in humeral part, costa 1 very weak, shortened, distinct in humeral quarter. Male genitalia with short, robust phallus, internal sac sclerotized, apical ring elliptical (Fig. 18). Measurements. BL 9.0 mm, WH 1.94 mm, PL 1.17 mm, PW 1.72 mm, Ediam 0.37 mm, Edist 0.62 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF897431C75FFD82FABA.taxon	distribution	Distribution. China, Sichuan Province. Until now, the species is known only from the type locality.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF897431C75FFD82FABA.taxon	etymology	Etymology. The specific epithet is derived from the local name Moxi, a village at the foothill of Gongashan.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF887431C322FD63FC12.taxon	materials_examined	Type material. Holotype, male, " N. Thailand, Changmai, 25. 4. 1993, P. Pacholátko leg. " (LMBC) Paratypes: 1 male, 1 female, " N Thailand, Changmai, 25. 4. 1993, Pacholátko leg. "; 2 males, " Guizhou, Dakua, 35 km NE Leishan, 20. ­ 24. 6. 1994, Bolm leg. "; 2 males, " Laos, Louangnamtha prov., 21.09 ° N, 101.19 ° E, 900 ­ 1200 m, 5. ­ 31. 5. 1997, Vít Kubáò leg. "; 1 male, " N Vietnam, Tam Dao, 1989, P. Pacholátko leg. "; 1 male, " Yunnan, Gaoligong mts., 25.22 ° N, 98.49 ° E, 17. ­ 24. 5. 1995, Vít Kubáò leg. "; 1 female, " NW Thailand, Mae Hong Son distr., Soppong, 19.27 ° N, 96.20 ° E, 1500 m, 7. ­ 12. 6. 1996, J. Horák leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF887431C322FD63FC12.taxon	diagnosis	Diagnosis. The internal sac of D. pacholatkoi sp. n. is similar to that of D. moxiensis sp. n. and both species have a short and stout phallus. They differ in the shape of apical ring, which is approximately as long as wide in D. pacholatkoi sp. n., larger eyes, and in the colouration of pronotum and elytra.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF887431C322FD63FC12.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum black, elytra red, both body parts covered with red pubescence. Head with short rostrum. Eyes small, interocular distance 1.36 times maximum eye diameter. Antennae slender, weakly compressed, very slightly serrate. Maxillary palpi slender. Pronotum 1.12 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, posterior angles distinct, lateral margins almost straight, posterior margin convex in the middle. Pronotal disc with frontal longitudinal costa reaching two fifths of pronotum length, transverse mound present in middle of disc. Scutellum black, posterior margin rounded. Elytra slender, flat, each elytron with three longitudinal costae. Costa 1 inconspicuous in apical part. Male genitalia as in fig. 20. Measurements. BL 6.0 mm, WH 1.26 mm, PL 0.88 mm, PW 0.99 mm, Ediam 0.36 mm, Edist 0.50 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF887431C322FD63FC12.taxon	distribution	Distribution. Dilophotes pacholatkoi sp. n. is a widely distributed species. It occurs in southern China (Yunnan, Guizhou), Vietnam (Tonkin), northern Laos, and northeaster Thailand.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66557FF887431C322FD63FC12.taxon	etymology	Etymology. The specific epithet is a patronym in honour of Mr. P. Pacholátko, a Czech entomologist and collector of the species.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF887431C257FBC0FB05.taxon	diagnosis	Diagnosis. Dilophotes anthracinus group is characterized by the flabellate male antennae, partially sclerotized internals sac and simple short phallus (Figs 22, 23).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF8B7431C2BDFEC1FD3A.taxon	materials_examined	Type material. Holotype, male, " Laos, 15 km N Louang Namtha, 750 m, 21.07 ° N / 101.21 ° E, 13. ­ 24. 5. 1997, E. Jendek & O. Šauša leg. " (LMBC). Paratype: 1 male, same locality data, (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF8B7431C2BDFEC1FD3A.taxon	diagnosis	Diagnosis. Dilophotes anthracinus sp. n. has flabellate male antennae and is the only Dilophotes species in the studied area with a concolourous black pronotum and elytra. In addition, the species has very short phallus and mostly membranous simple internal sac (Fig. 22). The closest species is D. lizipingensis sp. n. with much more slender phallus (Fig. 23), and red parts on pronotum and elytra.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF8B7431C2BDFEC1FD3A.taxon	description	Description of holotype. Body medium­sized, slender, black, pronotum, scutellum and elytra almost black with light pubescence. Eyes small, interocular distance 1.33 times maximum eye diameter. Antennae flabellate, flabellae of middle antennomeres three times longer than each antenomere. Pronotum 1.57 times wider than long. Frontal margins widely rounded, frontal angles distinct, lateral margins slightly concave, posterior angles moderately projected. Pronotal disc with frontal longitudinal costa reaching middle of disc, then inconspicuous. Disc surface black, shiny, with light pubescence. Elytra long, slender, flat, each elytron with three longitudinal costae. Costa 1 weak in apical part, costa 2 strong. Male genitalia with short phallus and partly membranous, simple internal sac (Fig. 22). Measurements. BL 5.5 mm, WH 1.32 mm, PL 0.77 mm, PW 1.21 mm, Ediam 0.36 mm, Edist 0.48 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF8B7431C2BDFEC1FD3A.taxon	distribution	Distribution. Dilophotes anthracinus sp. n. is known only from the type locality in northernmost Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66556FF8B7431C2BDFEC1FD3A.taxon	etymology	Etymology. The specific epithet refers to the black colouration of the pronotum and elytra.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66555FF8B7431C4BFFEC5F884.taxon	materials_examined	Type material. Holotype, male, " China, Sichuan, Liziping, 28. 6. ­ 3. 7. 1991 " (LMBC). Paratypes: 1 male, 1 female, " China, W. Guizhou prov., Leigongshan, Xijiang, 29. 5. ­ 2. 6. 1997, 1200 ­ 1900 m, Bolm leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66555FF8B7431C4BFFEC5F884.taxon	diagnosis	Diagnosis. Dilophotes lizipingensis sp. n. has flabellate male antennae but it is similar in body colouration to the dark red species belong to the other species groups based on their structure of male genitalia. Within groups having flabellate antennae it is well defined by the shape of its phallus and internal sac (Fig. 23).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66555FF8B7431C4BFFEC5F884.taxon	description	Description of holotype. Body medium sized, slender, black, pronotum black with brown margins, elytra dark brown to black, only humeri brownish red, both body part covered with dark red dense pubescence. Eyes small, interocular distance 1.38 times maximum eye diameter. Head with short rostrum. Antennae flabellate, lamella of antennomere 7 3.5 times trunk of antennomere (Fig. 4). Pronotum transverse, 1.5 times wider than long, frontal margin widely rounded, frontal angles inconspicuous, lateral margins very slightly concave, posterior angles slightly projected (Fig. 7). Disc with sharp longitudinal costa from frontal margin almost reaching posterior margin. Disc surface densely pubescent. Scutellum simply rounded. Elytra very slender, flat, each elytron with three longitudinal costae. Costa 1 and 2 weak, costa 3 strong especially in humeral quarter. Male genitalia with short phallus and partly sclerotized internal sac (Fig. 23). Measurements. BL 6.5 mm, WH 1.65 mm, PL 0.93 mm, PW 1.43 mm, Ediam 0.40 mm, Edist 0.55 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66555FF8B7431C4BFFEC5F884.taxon	distribution	Distribution. Dilophotes lizipingensis sp. n. should be widely distributed species in China, but at present it is known only from two distant localities in western Sichuan and Guizhou.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66555FF8B7431C4BFFEC5F884.taxon	etymology	Etymology. The specific epithet is derived from the local name Liziping in Sichuan, China.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8A7431C75FFB54FAB4.taxon	diagnosis	Diagnosis. Dilophotes ohirai is the only Japanese species with flabellate male antennae and red pubescence of pronotum. It reminds D. atrorufus in general appearance but the male genitalia differs substantially by very short phallus. D. vandykei does not have red pubescence on pronotum and its phallus is slender. Redescription of male. Body small, slender, black to dark brown, covered with dark fine pubescence, only pronotum red and sometimes humeri with reddish pubescence. Head small, transverse, moderately shining. Interocular distance 1.72 times maximum eye diameter. Antennae flabellate from antennomere 3, lamella of antennomere 3 as long as trunk of antennomere, lamellae of subsequent antennomeres twice or more longer than trunks of corresponding antennomeres. Maxillary palpi slender, apical palpomere almost parallelsided, bearing membranous processes apically. Pronotum wider than long, infuscate medially, hind angles acutely projected, frontal margin weakly rounded. Frontal keel reaching slightly beyond middle. Elytra almost parallel­sided, slightly widen posteriorly, surface thickly, rugosely punctured. Three longitudinal costae on each elytron, costa one the shortest and weakest. Legs long, slender. Male genitalia short, phallus resembling rolled leaf, widened in middle, internal sac membranous.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8A7431C75FFB54FAB4.taxon	description	Measurements. BL 7.3 mm, WH 1.74 mm, PL 0.88 mm, PW 1.50 mm, Ediam 0.32 mm, Edist 0.56 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8A7431C75FFB54FAB4.taxon	materials_examined	Material examined. 1 male, " Higashimatasawa, Kawakamimura, Nagano, 21. vii. 1980, K. Emoto (KMCK).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8A7431C75FFB54FAB4.taxon	distribution	Distribution. Dilophotes ohirai is endemic to Japan (Honshu).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8A7431C75FFB54FAB4.taxon	discussion	Remark. The reason for new generic placement was given in the introduction.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8D7431C32DFBDEFDD2.taxon	diagnosis	Diagnosis. Dilophotes vandykei resembles other species of D. anthracinus group. This species is characteristic in dark black pronotum, relatively slender phallus, and very long internal sac. Redescription of male. Body small, slender, dark reddish brown, covered with dark brown to black pubescence. Eyes small, interocular distance twice eye diameter. Antennae reaching middle of elytra, antennomeres 3 ­ 10 flabellate, lamellae of middle antennomeres more than twice longer than respective antennomeral trunks. Pronotum wider than long. Frontal margin produced, frontal angles inconspicuous, posterior angles moderately projected. Disc densely punctuated, with moderately sharp longitudinal costa attached to frontal margin and reaching middle of disc and with median longitudinal sulcus basally. Elytra very slender, flat, subparallel­sided, each elytron with three longitudinal costae in humeral part, costa 1 very weak, shortened, distinct in humeral third. Male genitalia with small phallus, internal sac long.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8D7431C32DFBDEFDD2.taxon	description	Measurements. BL 5.5 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8D7431C32DFBDEFDD2.taxon	distribution	Distribution. Dilophotes vandykei is known only from northern Japan (Hokkaido).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66554FF8D7431C32DFBDEFDD2.taxon	discussion	Remarks. I have not studied any specimen of this rare species. The above given description is a shortened form of those published by Nakane (1970). The Nakane's description enables sure identification within the D. anthracinus group.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8D7431C497FD13FC2D.taxon	diagnosis	Diagnosis. The male genitalia are very characteristic (Fig. 25). Their shape does not show close relationship of D. kubani sp. n. to any known Dilophotes, and therefore it is classified as a separate species group.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8C7431C03CFD31FB8A.taxon	materials_examined	Type material. Holotype, male, " Laos, Boli Kham Xai prov., Ban Nape (8 km NE), 18.21 ° N, 105.08 ° E, 600 m, 1. ­ 18. 5. 2001, Vít Kubáò leg. " (LMBC). Paratype: 1 female, " Laos, Khammouan Pr., Nakai env., 600 m, 17.43 ° N, 105.09 ° E, 22. 5. ­ 8. 6. 2001, E. Jendek & O. Šauša leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8C7431C03CFD31FB8A.taxon	diagnosis	Diagnosis. Dilophotes kubani sp. n. resembles D. luteus sp. n. in pronotum and elytra colouration. These species differ substantially in the shape of male genitalia and the shape of male antennae.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8C7431C03CFD31FB8A.taxon	description	Description of holotype. Body medium­sized, slender, black, pronotum and elytra light orange red, with light red pubescence, scutellum brown basally, light brown to red apically. Eyes large, maximum eye diameter 1.21 times interocular distance. Antennae acutely serrate, strongly compressed, very long, reaching apex of elytra. Pronotum 1.75 times wider than long. Frontal margin widely rounded, frontal angles inconspicuous, lateral margins slightly concave, posterior angles moderately projected. Disc with sharp longitudinal costa attached to frontal margin and reaching almost posterior margin. Disc surface shiny, light brown with dense pubescence. Scutellum slightly emarginate in posterior margin. Elytra very slender, flat, each elytron with three longitudinal costae in humeral part, costa 1 very weak, shortened, distinct in humeral quarter. Male genitalia with small phallus, internal sac large (Fig. 25). Measurements. BL 5.25 mm, WH 1.37 mm, PL 0.66 mm, PW 1.15 mm, Ediam 0.44 mm, Edist 0.36 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8C7431C03CFD31FB8A.taxon	distribution	Distribution. Dilophotes kubani sp. n. is known only from the type locality in central Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66553FF8C7431C03CFD31FB8A.taxon	etymology	Etymology. The specific name is a patronym in honour of V. Kubáò, a Czech entomologist and collector of the holotype.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8C7431C2CFFD5FFACC.taxon	diagnosis	Diagnosis. Dilophotes luteus group is characterized by the flabellate male antennae, very unusual simple, curved phallus, and membranous internal sac. The shape of male genitalia does not show any close relationships to other species groups. D. luteus sp. n. is the only species classified in the species group.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8F7431C385FBBAFD62.taxon	materials_examined	Type material. Holotype, male, " Laos, Boli Kham Xai prov., Ban Nape (8 km NE), 18.21 ° N / 105.08 ° E, 600 m, 1. ­ 18. 5. 2001, Vít Kubáò leg. " (LMBC).	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8F7431C385FBBAFD62.taxon	diagnosis	Diagnosis. Dilophotes luteus sp. n. reminds in body size and colouration D. kubani sp. n. These species differ substantially in the shape of male genitalia (Figs 25, 26) and antennae. Additionally, D. luteus sp. n. is the only species with yellow scutellum.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8F7431C385FBBAFD62.taxon	description	Description of holotype. Body medium­sized, slender, black, pronotum, scutellum, and elytra light yellow, covered with light ochre pubescence. Eyes very large, maximum eye diameter 1.36 times interocular distance. Antennae flabellate, lamellae of middle antennomeres three times length of trunks. Pronotum 1.57 times wider than long. Frontal margins widely rounded, frontal angles inconspicuous, lateral margins slightly concave, posterior angles moderately projected. Disc with short frontal longitudinal keel and distinct transverse mound in middle. Disc surface shiny, light brown with dense pubescence. Scutellum slightly emarginate posteriorly. Elytra long, very slender, flat, each elytron with three longitudinal costae in humeral part, costa 1 very weak, shortened, distinct in humeral quarter. Male genitalia with small, strongly curved phallus (Fig. 26). Measurements. BL 6.0 mm, WH 1.32 mm, PL 0.77 mm, PW 1.21 mm, Ediam 0.50 mm, Edist 0.36 mm.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8F7431C385FBBAFD62.taxon	distribution	Distribution. Dilophotes luteus sp. n. is known only from the type locality in central Laos.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
039987C66552FF8F7431C385FBBAFD62.taxon	etymology	Etymology. The specific name refers to the colouration of elytra.	en	Bic, Vlastislav (2002): Review of the genus Dilophotes Waterhouse (Coleoptera: Lycidae) of the Palaearctic Region and Indochina. Zootaxa 59: 1-26, DOI: 10.5281/zenodo.155977
