taxonID	type	description	language	source
039B8798FFA7FFEDFED759DBFEF016D3.taxon	materials_examined	Material examined. Holotype Ψ, 22 May 2004, Turkey, Trabzon province, Yomra district, Çardaklı village (40 ° 88 ˏ N, 39 ° 91 ˏ E), litter and grass from muddy soil, 758 m a. s. l. Paratypes, same date and locality as holotype, 1 Ψ, 5 ɗ, 4 DN; 14 June 2004, Yomra district, Fındıklı village, (40 ° 86 ˏ N, 39 ° 95 ˏ) extremly wet weeds and litter, 4 Ψ, 9 ɗ, 4 DN, 750 m a. s. l.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA7FFEDFED759DBFEF016D3.taxon	description	Female (Holotype). Dorsum (Fig. 1). Idiosoma subcircular, dorso-ventrally flattened, 615 µm long and 490 µm wide, well sclerotised and yellow-brown in colour. Dorsal plate entire, surface covered with small circular markings. Plate surrounded by an entire marginal plate. Marginal plate smooth, not ornamented, and bearing 12 pairs of smooth, lanceolate setae. Dorsal plate with 33 – 35 pairs of relatively long lanceolate setae with fine spicles in proximal half, and whip-like in distal half. All setae long enough to pass insertion of following seta. Posterior end of dorsum lacking circular ornamentation. Ven te r (Fig. 2). Surface of sternal plate ornamented with small circles. Sternal setae st 1 - st 5 straight, short and simple. Sternal setae st 1 inserted level with coxae II, close to anterior end of epigynium; st 2 - st 5 lateral to genital aperture, each accompanied by a circular pore. Genital plate 67 µm long and 45 µm wide. Surface of epigynial plate smooth, with fine spicules posteriorly, shape of plate helmet-like, between coxae II – IV, and bearing a chitinised anchor-like process in its anterior half. A pair of lyrifissures is located near the anterior border of the sternal plate. Ventral region with 7 pairs of ventral opisthonotal setae and numerous lateral setae. All ventral opisthonotal setae serrated proximally, resembling dorsal plate setae. Peritremes located at level of coxae II – III, anterior prolongation hook-like, its tip directed outward. Stigmatal openings not bearing a posterior prolongation. Pedofossal and metapodal area without surface ornamentation, endopodal plates and sternal region ornamented with a few small circular markings. Pedofossal grooves distinct and well developed, and without ornamentation. Post-anal seta (pas) absent. Gnathosoma (Fig. 3). Chelicerae dentate, without nodus, movable digit 60 µm long, fixed digit and its basal elongation (Cmp) total length 157.5 µm long (Fig. 3 A). Hypostomal setae h 1 simple, smooth, needle-like, almost reaching end of laciniae; setae h 2, h 3, h 4 serrated. Setae h 2 with 4 – 6 denticles on both sides; h 4 with 5 – 6 fine denticulations laterally, h 3 longer than h 2 – h 3, with 2 – 3 lateral denticles. Corniculus hornlike. Hypostomal constrictions present between h 3 – h 4; surface of pro- and deutosternum finely denticulate (Fig. 3 B). Tritosternum vase-shaped basally, with three branches, and each branch bifurcate apically, its medial lacinia longer than others; tips of laciniae denticulate (Fig. 3 C). Ligula triangular, its surface with spiculate, basal margins with 3 denticles (Fig. 3 D). Epistome with triangular base and denticulate lateral margins, denticles gradually lengthening from tip to base (Fig. 3 F). Palps short and robust, palp tarsus with a set of tactile setae and thick prong-like setae, ventral setae of trochanter and some setae of femur and genu finely denticulate (Fig. 3 E). Legs (Fig. 4). Coxae I large, tarsi I without claw, legs II – IV with a pair of claws at tip of the ambulacral prolongation; leg setae both simple and denticulate. Whip-like seta of tarsus I longer than tarsus. Femurs and trochanters I – IV bearing a membranous flange. Male. Ven t er (Fig. 5). Genital plate oval, elongate, located between coxae III – IV, 67 µm long and 45 µm wide. Sternal region with scattered circular markings, 5 pairs of short and thorn-like (st 1 – st 5) setae. Setae st 1 arise from level of coxae II, st 2 between coxae II – III, st 3 close to anterior end of coxa III, st 4 between coxae III – IV, outside genital plate, and st 5 behind genital plate. A circular pore accompanies each setae near its base. Other morphological features, setal arrangement, body shape and patterns as for female. Deutonymph. Idiosoma 560 – 600 µm long, 460 – 500 µm wide. Dorsum (Fig. 6). Dorsal plate with 33 – 35 pairs of relatively long setae, setae with fine spicules in proximal half, and whip-like in distal half as for adults. All setae long enough to pass insertion of following seta. Lateral setae dagger-like, curled upward at tip. Two pairs of posteriorly situated setae similar to spiculate dorsal setae. Ven te r (Fig. 7). Sternal plate ornamented with fine punctation. Sternal plate anvilshaped, bearing five pairs of simple, smooth, needle-like setae st 1 – st 5; 310 µm long and 130 µm wide. Anal plate setae similar in shape to dorsal shield setae. Anterior end of peritreme long and slightly twisted exteriorly. Stigmata located between coxae II – III, without a posterior prolongation. Pedofossae distinct, well developed; posterior margin of pedofossae IV pointed. Ventri-anal plate convex anteriorly, without ornemantation, 115 µm long and 245 µm wide. Anal plate with 2 pairs of smooth pointed setae. Distance between coxae II, III and IV: 97, 150, 195 µm, respectively. Gnathosoma. Hypostomal setae h 1 simple, needle-like, smooth; setae h 2 – h 4 denticulate. Epistome, tritosternum, chelicera and other gnathosomal features as in adults, but slightly less robust. Protonymph and larva. Unknown.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA7FFEDFED759DBFEF016D3.taxon	discussion	Remarks. Uropoda turcica sp. nov. is closely related to species of Cilliba, by having a well marked pedofossae, a dorso-ventrally flattened body, many marginal setae, five pairs of sternal setae, five pairs of simple ventral setae and two pairs of simple adanal setae, a V-shaped peritreme without posterior prolongation, 6 - branched tritosternum with broad base, a chelicerae without nodus, and leg I tarsi without claws. The new species differs from all congeners by virtue of its dorsal, ventri-anal and adanal setae which are spiculate proximally part and elongate and whip-like distally. The submarginal setae are longer and more robust than those of other closely related species. Additionally, in comparison to other congeneric species, the female of the new species has some distinctive characters, such as a large helmet-shaped epigynium with a chitinised anchor-like process in its anterior half, without surface ornamentation, located between coxae II – VI, and a porous sternum marked with well-developed endopodal lines. Deutonymphs of the new species are differentiated from those of related species by having dorsal, ventri-anal and adanal setae basally serrated, but slimmer than those of adults.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA2FFE7FED75ED0FD2A1593.taxon	materials_examined	Material examined. Holotype Ψ, Giresun province, Tirebolu district (41 ° 06 ’ N, 39 ° 14 ’ 25 ” E), in soil around chestnut tree-stump (Castanea sativa), 10 June 2006, 110 m a. s. l. Paratypes, 8 Ψ, 17 ɗ, same data as holotype; 1 Ψ, 1 ɗ, Dere village, (40 ° 72 ’ N, 38 ° 46 ’ E), soil and litter, 21 April 2001. Soil samples were extremely wet.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA2FFE7FED75ED0FD2A1593.taxon	description	Female (holotype). Idiosoma 475 µm long, 375 µm wide. Body oval, well sclerotised. Dorsum (Figs 8, 9). Dorsal plate not heavily sculptured, brown in colour, without ornamentation. Four pair of post-dorsal setae finely pilose lateraly. Posterior bulge triangular, with fine spines, pilose anteriorly. Dorsal plate with about 15 pairs of dorsal setae, smooth and thorn-like. Marginal plate with fine spines. All marginal plate setae dagger-like with sawtooth denticles on one side, each arising on a chitinous tubercle. Submarginal setae finely denticulate laterally. Ven te r (Fig. 10). Opisthogastric setae very long, needle-like, bearing a medial keel throughout their length. A pair of short, thorn-like ventri-anal setae inserted on smooth cuticle in front of anus. Genital operculum tongue- or helmet-shaped; length 130 µm and width 100 µm. Surface of operculum without ornamentation. Sternal seta st 1 at level of coxae II, st 2 close to anterior margin of epigynial plate, st 3 - st 5 located lateral to plate. Pre- and para-anal setae short and simple. Post-anal seta resembles pre- and post-anal setae. A pair of additional sternal setae occurs just behind epigynium. Scabellum clear and well developed, with four pairs of pilose capitular setae. Peritremes simple, short, located at level of coxae II – III; anterior prolongation directed externally; posteriorly with stigmata located in pedofossae III. Pedofossal grooves deep and well marked; pedofossae IV directed internally accomodating distal part of legs IV. Two pairs of short pilose setae behind anal region near posterior margin of idiosoma. Gnathosoma (Fig. 11). Chelicera without nodus; movable digit with one tooth; fixed digit 33 µm long, with anterior sensory angular point and a sensillar setae near distal part, movable digit 22 µm long, middle section 135 µm long (Fig. 11 A). Epistome lancet-like, bifurcate and pilose distally, basally denticulate, witha row of median denticles (Fig. 11 B). Base of tritosternum vase- or glass-shaped, its laciniae branched, internal branches denticulate and paraxial branches shorter and smooth (Fig. 11 C). Laciniae of hypostome finely denticulate (Fig. 11 D). Hypostome with four pairs of setae; hypostomal setae h 1 long and forked distally, setae h 2 - h 4 denticulate. Hypostomal setae h 4 shorter than the others, serrated, not reaching the base of h 3; h 3 long, exceeding base of h 2; h 2 half length of h 1. Hypostome articulated between h 3 - h 4. Corniculi horn-like; deutosternum with fine superficial denticulations up to h 4 (Fig. 11 D). Male. Idiosoma 475 µm long and 375 µm wide. Ven te r (Fig. 12). Ventral surfaces of idiosoma smooth and without ornamentation. Sternal area smooth, without ornamentation, sternal setae simple, short and thorn-like. Genital aperture located between coxae IV, its opening 30 µm in diameter. Sternal setae st 1 at level of coxae II, st 2 behind coxae II – III, st 3 between coxae III – IV, st 4 close to anterior margin of genital plate, st 5 near posterior level of genital plate. Pre- and post-anal setae short and simple. Genital plate circular and 50 µm diameter. Posterior opisthosomal leaflike setae 105 – 130 µm, and column length 25 – 35 µm at base. Legs, chelicerae, epistome, tritosternum, setal arrangement, idiosoma shape and pattern, and other morphological characters resemble those of female. Deutonymph. Idiosoma 460 µm long, 360 µm wide. Dorsum (Fig. 13). Dorsal plate without ornamentation, light brown in colour, weakly sclerotised, anterior and posterior margins slightly indented. All dorsal, marginal and lateral setae simple and hair-like, dorsal setae shorter than marginal setae. Dorsal plate with about 20 pairs of setae, I 5 shortest, marginal series with 30 pairs. Ven te r (Fig. 14). Sternal plate 135 – 150 µm long and 50 – 55 µm wide, anal plate 105 – 125 µm long and 150 – 155 µm wide. Sternal and anal plates with web-like ornamentation. Endo- and exopodal plates with punctate ornamentation. Sternal plate pitcher-shaped, extending from coxae I to behind coxae IV, bearing five pairs of short setae st 1 – st 5. Interscutal membrane around ventral plates striated; ventri-anal plate setae short and thorn-like, except a pair of long ventri-anal setae at posterior margin. Other ventral setae in soft skin long and needle-like. Coxae I large, close together; pedofossae distinct, well developed. Anterior extension of peritreme long, medially elbowed, stigmata at level of coxae III (Fig. 14).	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA2FFE7FED75ED0FD2A1593.taxon	discussion	Remarks. Adults of the new species differ from their congeners by the presence of leaf-like setae on posterior idiosoma. These setae are very similar to grass leaves, which bear a median keel and a few basal denticles. Each seta arise from a large chitinous based limited by wide flaps. In the other known species of Crinitodiscus, setae like this are absent (Athias-Binche & Błoszyk, 1985; Kontschán, 2003; Bal, 2005). The new species can also be distinguished from other species by the following combination of characters: dorsal and ventral plates without ornamentation, a series of asymmetrical anterior setae on the dorsal shield; marginal setae leaf-like, with denticles laterally and medially and a median keel; sublateral setae bearing a few minute denticles; four pairs of widened and denticulate posterior dorsal setae in front of the postdorsal hump; this hump is high and conspicuous, its lateral sides with web-like ornamentation and superficial spines; epigynium without ornamentation and not bearing a hyaline lobate process or a prolongation. Also, there is a pair of very long ventral setae behind the epigynium; two pairs of short pilose ventral setae behind the anus; and two pairs of very short ventral setae between the epigynium and anus.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA8FFF9FED75E93FEDB10DB.taxon	materials_examined	Material examined. Holotype Ψ, Bingöl province, Palulu Mountain (38 ° 57 ’ N, 40 ° 14 ’ E), 25 May 2004, 1,250 m a. s. l. Paratypes, 3 Ψ, 1 ɗ, same data as holotype. Litter and bark from deciduous tree stump samples were moist.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA8FFF9FED75E93FEDB10DB.taxon	description	Female (Holotype). Dorsum (Fig. 15). Idiosoma oval, 570 µm long and 420 µm wide, well sclerotised, brown. Dorsal plate entire, strongly convex, without ornamentation. Plate surrounded by an entire marginal plate. Marginal plate narrow, bearing 12 pairs of setae. Dorsal plate with 40 – 50 relatively short, simple thorn-like setae, setae not reaching insertion of following setae. Posterior end of dorsum with four knife-like setae, these powerful setae longer and wider than other dorsal setae. Ven te r (Fig. 16). Epigynial plate 150 µm long and 110 µm wide, helmet-like, bearing a crown-like anterior extension; posterior margin behind coxae IV, anterior margin behind coxae I. Surface of plate with punctuations and many shining pores. Ventral region with 4 pairs of sternal (st 1 - st 4) setae, and 7 pairs of ventri-anal setae. Sternal setae st 1 inserted at the level coxae II, close to epigynium; st 2 - st 3 lateral to genital aperture, st 4 behind epigynial plate. Anterior sternal region with a chitinous wrinkled surface process. Ventrianal plate ornamented with micropunctations and small porous shining markings. All ventral setae simple, smooth, needle-like. Peritremes at level of coxae II and pedofossae III, anterior prolongation directed exteriorly. Stigmatal openings with short posterior prolongations. Endopodal line absent. Exopodal plate ornamented with shining pores. Pedofossal grooves distinct and well developed, fossae IV also with shining pores. Anal opening 13 µm in diameter. Post-anal seta present and similar in form to the other ventral setae. Gnathosoma (Fig. 17). Chelicerae unidentate, with large nodus, movable digit 30 µm, fixed digit 52 µm, and middle section 160 µm long (Fig. 17 A). Setae h 1 simple, smooth, thorn-like, almost reaching end of laciniae. Setae h 2 smooth, short and stout, reaching base of h 1. Setae h 3, h 4 branched; h 4 reaching past base of h 3, and h 2 reaching base of h 1. Corniculus horn-like, with blunt end. Hypostomal constrictions present between h 3. Deutosternum with superficial denticulation (Fig. 17 C). Epistome with triangular base and denticulate lateral margins, denticles gradually lengthening from tip to base (Fig. 17 B). Tritosternal base vase-shaped, medio-lateral lacinia laterally denticulate and tip bifid, lateral part of laciniae denticulate and shorter than middle part; lateral branches denticulate on one side (Fig. 17 D). Palps short and robust, palp tarsus with a set of tactile setae, ventral setae of trochanter finely denticulate (Fig. 17 E). Legs (Fig. 18). Legs I – II directed anteriorly, and III – IV posteriorly. Tarsi II – IV with a pair of claws at tip of ambulacral prolongation; tarsi I with a lighter claw on a delicate ambulacrum. Whip-like setae longer than tarsi I. Setae on all legs simple and thorn-like. Femurs and trochanters bearing a membraneous flange. Male. Dorsum. All dorsal features as in females. Length of idiosoma 575 µm, width 430 µm. Ven te r (Fig. 19). Surface of sternal plate ornamented with micropunctations and small rounded hollows. Sternal setae straight and thorn-like. Sternal setae st 1 inserted level with coxae II, st 2 - st 4 anterior of genital aperture on line, st 5 close to posterior end of plate. Genital aperture circular, between coxae IV, and surrounded thick chitinous wall. Genital plate 45 µm long and 40 µm wide. Surface of genital plate with micropunctation. Sternal plate with a chitinous wrinckled surface ornamentations in front of genital plate. Legs. A sexual dimorphism occurs in femur II which bears a powerful apophysis (Fig. 18 E). Other morphological features, setal arrangement, body shape and patterns as for female. Deutonymph, protonymph and larva. Unknown.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
039B8798FFA8FFF9FED75E93FEDB10DB.taxon	discussion	Remarks. Hirschmann (1989) reviewed the cosmopolitan genus Uroobovella and created 23 species-groups for easy identification. All distinctive characters for the speciesgroups of the genus Uroobovella are summarised below, emphasising the distinctive characters of the new species. U. anatolica sp. nov. is differentiated from the vinicolora, pergibba, costai, schulzi, tricuspis, hilli, marginata, ruhmi, coprophila, flagelliger, fimicola, appendiculata, minima, ipidis species-groups and all related species by the presence of five pairs of sternal setae in males (4 in females). Hovewer, the foveolata, venusta, elegans, formosana, rackei, micherdzinski, fracta, carinata, pulchella and vitzhumi species-groups and related species share the same number of sternal setae (5 pairs in male 5, 4 pairs in females). The new species is differentiated from the foveolata, venusta and elegans species-groups, by the presence of a pair of claws on pretarsus I. Members of formosana species-group possess a pair of claws on legs I, but, lack marginal plates and endopodal lines, and their males do not have an apophysis on legs II. On the other hand, in U. rackei, there is a well-developed marginal plate, but the dorsal and ventral plates are separated by a soft chitinised area containing platelets. The new species is differentiated from U. rackei by the relatively hard and heavily sclerotised processes between the dorsal and ventral plates. The species groups micherdzinski, fracta, carinata, pulchella have a marginal plate as a distinctive character in common with the new species. Hovewer, the new species is differentiated from the micherdzinski and fracta species-groups by the presence of structural ornamentation on these plates. The carinata species-group has a dorsal hunch-backed carina, which does not exist in the new species. The new species differs from vitzhumi species-group by the absence of lateral shoulder and vertex, deep circular and polygonal hollows on the plates, marginal plates with densely undulating marginal plate internally, and endopodal line and tasseled setae on marginal plate. Finally, the pectinata species-group does not have leg pedofossae, in contrast with the new species. The new species shares some morphological characters with the species of the pulchella group. The members of this species-group have 5 pairs of sternal setae in males and 4 in females; the idiosoma is egg shaped or widely oval; endopodal lines are present, legs I lack claws; with a well-developed marginal plate and with structural plates are present; and the marginal plate is smooth sided medially. The new species is closely related to U. denticulata Hirschmann & Zirngiebl-Nicol 1972 in the pulchella speciesgroup, which has some very similar morphological characters with the new species: with web-like patterns on plates, presence of well developed endopodal lines, and the existence of short simple setae on the dorsal, marginal, lateral, sternal and ventral plates. Hovewer, the epigynium is the most distinctive character for U. denticulata, which has a wide and notched extension apically, as wide as the epigynium. In summary, U. anatolica sp. nov. differs from all of its congeners by the following combinaton of character: males with five (and females four) pairs of sternal setae; tarsi I without claws; dorsal plate smooth and without ornamentation; tongue-like epigynium with a dentate anterior prolongation; leg pedofossae well-developed; marginal plate present and not fragmented posteriorly; all dorsal, marginal, lateral and ventral setae simple, smooth and thorn-like; sternal plate with a chitinous wrinkled surface ornamentations near anterior end of epigynium, ventri-anal and epigynial plates ornamented with micropunctations and small circular pores; soft cuticle supporting dorsal and ventral plates absent, and cuticle of this region hardened and sclerotised; medial borders of marginal plate smooth and without chitinous undulations; hyaline prolongation of fixed digit of chelicera knife-like; peritremes hook-like anteriorly and the anterior end inclined inward, and posterior prolongation smooth; and males with apophysis in leg femora II.	en	Bal, Durmuş Ali (2006): New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae). Zootaxa 1368: 19-40, DOI: 10.5281/zenodo.174843
