taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372422/files/figure.png	https://doi.org/10.5281/zenodo.15372422	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372426/files/figure.png	https://doi.org/10.5281/zenodo.15372426	FIGURE 3. Morphological details of Aphanocephalus: A) antenna; B) maxillary palp; C) labrum; D) prementum and labial palps; E) right mandible, ventral; F) left mandible, inner; G) left mandible, ventral; H) tentorium.	FIGURE 3. Morphological details of Aphanocephalus: A) antenna; B) maxillary palp; C) labrum; D) prementum and labial palps; E) right mandible, ventral; F) left mandible, inner; G) left mandible, ventral; H) tentorium.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372428/files/figure.png	https://doi.org/10.5281/zenodo.15372428	FIGURE 4. Morphological details of Aphanocephalus: A) A. pallidipennis, abdomen, male, apical part of male genitalia visible; B) A. nitidus, abdomen, female, female genitalia and coxites visible; C) A. sp., sternite VIII, male; D) A. sp., female genitalia; E) A. sp., metendosternite; F) A. sp., female, coxites and proctiger, horizontal.	FIGURE 4. Morphological details of Aphanocephalus: A) A. pallidipennis, abdomen, male, apical part of male genitalia visible; B) A. nitidus, abdomen, female, female genitalia and coxites visible; C) A. sp., sternite VIII, male; D) A. sp., female genitalia; E) A. sp., metendosternite; F) A. sp., female, coxites and proctiger, horizontal.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372444/files/figure.png	https://doi.org/10.5281/zenodo.15372444	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372446/files/figure.png	https://doi.org/10.5281/zenodo.15372446	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372448/files/figure.png	https://doi.org/10.5281/zenodo.15372448	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A31DF4EFF29D6085C10FDAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372434/files/figure.png	https://doi.org/10.5281/zenodo.15372434	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3DDF41FF29D0025AECFACB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372444/files/figure.png	https://doi.org/10.5281/zenodo.15372444	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372434/files/figure.png	https://doi.org/10.5281/zenodo.15372434	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3CDF46FF29D2C55BFBFCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372422/files/figure.png	https://doi.org/10.5281/zenodo.15372422	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3BDF47FF29D4305C74FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3BDF47FF29D4305C74FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372444/files/figure.png	https://doi.org/10.5281/zenodo.15372444	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3BDF47FF29D4305C74FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372434/files/figure.png	https://doi.org/10.5281/zenodo.15372434	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3BDF47FF29D4305C74FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3BDF47FF29D4305C74FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3ADF59FF29D5E45D7BFED1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3ADF59FF29D5E45D7BFED1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372444/files/figure.png	https://doi.org/10.5281/zenodo.15372444	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3ADF59FF29D5E45D7BFED1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A3ADF59FF29D5E45D7BFED1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A24DF59FF29D6095C82F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A24DF59FF29D6095C82F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372446/files/figure.png	https://doi.org/10.5281/zenodo.15372446	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A24DF59FF29D6095C82F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372434/files/figure.png	https://doi.org/10.5281/zenodo.15372434	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A24DF59FF29D6095C82F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A27DF5CFF29D7F05E59FB25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A27DF5CFF29D7F05E59FB25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372446/files/figure.png	https://doi.org/10.5281/zenodo.15372446	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A27DF5CFF29D7F05E59FB25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A27DF5CFF29D7F05E59FB25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A21DF5DFF29D2485D7BFC21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A21DF5DFF29D2485D7BFC21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372446/files/figure.png	https://doi.org/10.5281/zenodo.15372446	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A21DF5DFF29D2485D7BFC21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A21DF5DFF29D2485D7BFC21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A21DF5DFF29D2485D7BFC21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372422/files/figure.png	https://doi.org/10.5281/zenodo.15372422	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A20DF5EFF29D3595DECFBDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A20DF5EFF29D3595DECFBDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372448/files/figure.png	https://doi.org/10.5281/zenodo.15372448	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A20DF5EFF29D3595DECFBDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A20DF5EFF29D3595DECFBDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A20DF5EFF29D3595DECFBDD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A23DF5FFF29D3055B60FCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A23DF5FFF29D3055B60FCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372448/files/figure.png	https://doi.org/10.5281/zenodo.15372448	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A23DF5FFF29D3055B60FCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A23DF5FFF29D3055B60FCDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A22DF51FF29D4305E87FDD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A22DF51FF29D4305E87FDD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372448/files/figure.png	https://doi.org/10.5281/zenodo.15372448	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 12. Male genitalia of Aphanocephalus species: A–C) A. leai nom. nov.; D–F) A. nitidus Lea; G–I) A. occidentalis sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A22DF51FF29D4305E87FDD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A22DF51FF29D4305E87FDD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372440/files/figure.png	https://doi.org/10.5281/zenodo.15372440	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	FIGURE 8. Habitus images of Australian Aphanocephalus: A, B) A. leai nom. nov.; C, D) A. nitidus Lea; E, F) A. occidentalis sp. nov.; G–L) A. pallidipennis Lea, various color forms.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2CDF54FF29D50D5CDCFF35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372422/files/figure.png	https://doi.org/10.5281/zenodo.15372422	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	FIGURE 2. Morphology of Aphanocephalus: A) A. bimaculatus, head and mouthparts, ventral; B) A. pallidipennis, antenna; C) A. bimaculatus, head and prosternum, ventral; D) A. lawrencei, head and prosternum, ventral; E) A. pallidipennis, head and prosternum, ventral; F) A. sp., tarsal claw; G) A. pallidipennis, terminal antennomere; H) A. sp., tarsus; I) A. bimaculatus, meso-, metaventrite and first abdominal ventrite; J) A. nitidus, meso-, metaventrite and first abdominal ventrite.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372434/files/figure.png	https://doi.org/10.5281/zenodo.15372434	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	FIGURE 6. Type material and labels of Aphanocephalus species: A) A. potamophilus Lea, lectotype (SAMA); B) A. quadrimaculatus Lea, lectotype (SAMA); C) A. quadrinotatus Lea, lectotype (SAMA); D) A. austerus John, lectotype (SMNS); E) A. hackeri John, lectotype (SMNS). (Figs A–C licensed under CC-BY-NC-SA 4.0, Museum of South Australia; photos D and E kindly provided by Kevin Weissing, SDEI)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372444/files/figure.png	https://doi.org/10.5281/zenodo.15372444	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	FIGURE 10. Male genitalia of Aphanocephalus species:A–C) A. bimaculatus Grouvelle; D–F) A. blackburni nom. nov.; G–I) A. calderi sp. nov. A, D, G—penis, lateral; B, E, H—tegment, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A29DF56FF29D62D5CF4FA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372430/files/figure.png	https://doi.org/10.5281/zenodo.15372430	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	FIGURE 5. Type material and labels of Aphanocephalus species:A) Eleothreptus punctulatus Blackburn, lectotype (NHM); B) A. bimaculatus Grouvelle, lectotype (NHM); C) A. bimaculatus Lea, lectotype (SAMA); D) A. nitidus Lea, lectotype (SAMA); E) A. pallidipennis Lea, lectotype (SAMA); F) A. poropterus, lectotype (SAMA). (Figs C–F licensed under CC-BY-NC-SA 4.0, Museum of South Australia)	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2BDF57FF29D1215B79F941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372436/files/figure.png	https://doi.org/10.5281/zenodo.15372436	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	FIGURE 7. Habitus images of Australian Aphanocephalus: A, B) A. bimaculatus Grouvelle; C, D) A. blackburni nom. nov.; E, F) A. calderi sp. nov.; G, H) A. hackeri John, I, K) A. kukuyalanji sp. nov.; J, L) A. lawrencei sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372418/files/figure.png	https://doi.org/10.5281/zenodo.15372418	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	FIGURE 1. Morphology of Aphanocephalus: A) A. bimaculatus, habitus; B) A. potamophilus, habitus; C) A. pallidipennis, habitus; D) A. bimaculatus, pronotum; E) A. punctulatus, pronotum; F) A. pallidipennis, pronotum; G) A. lawrencei, lateral; H) A. queenslandicus, lateral, pseudopores visible; I) A. bimaculatus, head, frontal, with antennal pores visible; J) A. pallidipennis, posterior angle of pronotum fitting into excavation of elytra; K) A. lawrencei, elytral pores; L) A. pallidipennis, elytral pores.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A2ADF68FF29D1B35FBAF97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372446/files/figure.png	https://doi.org/10.5281/zenodo.15372446	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 11. Male genitalia of Aphanocephalus species:A–C) A. hackeri John; D–F) A. kukuyalanji sp. nov.; G–I) A. lawrencei sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A15DF69FF29D0655A1FF838.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A15DF69FF29D0655A1FF838.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A15DF69FF29D0655A1FF838.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372462/files/figure.png	https://doi.org/10.5281/zenodo.15372462	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	FIGURE 15. Female spermatheca of Aphanocephalus species: A) A. bimaculatus Grouvelle; B) A. kukuyalanji sp. nov.; C) A. lawrencei sp. nov.; D) A. pallidipennis Lea; E) A. potamophilus Lea; A. punctulatus (Blackburn); G) A. queenslandicus sp. nov.; H) A. slipinskii sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A15DF69FF29D0655A1FF838.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A15DF69FF29D0655A1FF838.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372452/files/figure.png	https://doi.org/10.5281/zenodo.15372452	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 13. Male genitalia of Aphanocephalus species: A–C) A. pallidipennis John; D–F) A. potamophilus Lea; G–I) A. punctulatus (Blackburn). A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A16DF6BFF29D7F05D57F9E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372442/files/figure.png	https://doi.org/10.5281/zenodo.15372442	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	FIGURE 9. Habitus images of Australian Aphanocephalus: A, B) A. potamophilus Lea; C, D) A. punctulatus (Blackburn); E–G) A. queenslandicus sp. nov.; H, I) A. slipinskii sp. nov.; J, K) A. weiri sp. nov.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A16DF6BFF29D7F05D57F9E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372458/files/figure.png	https://doi.org/10.5281/zenodo.15372458	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	FIGURE 14. Male genitalia of Aphanocephalus species: A–C) A. queenslandicus sp. nov.; D–F) A. slipinskii sp. nov.; G–I) A. weiri sp. nov. A, D, G—penis, lateral; B, E, H—tegmen, lateral; C, F, I—tegmen, inner.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
039A2D1C0A16DF6BFF29D7F05D57F9E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15372464/files/figure.png	https://doi.org/10.5281/zenodo.15372464	FIGURE 16. Distribution map of Australian Apahnocephalus species.	FIGURE 16. Distribution map of Australian Apahnocephalus species.	2025-05-08	Szawaryn, Karol		Zenodo	biologists	Szawaryn, Karol			
