taxonID	type	description	language	source
039AB4509330FFC914C5548AFDA2B2A7.taxon	materials_examined	ALGERIA: 2 ♂♂, 2 ♀♀, Bou Saada, 29 Apr. 1987, G. Magnani leg. (GS, MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C554D4FDBBB37C.taxon	materials_examined	UGANDA: 3 ♂♂, Kabulamuliro, 1910, Dr. E. Bayon [leg.], det. E. Benderitter 1927 (MSNG).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C5553FFDC7B30A.taxon	materials_examined	IRAN: 1 ♂, paratype, Kerman, 2 Apr. [19] 51, Salavatian leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C55589FDCCB049.taxon	materials_examined	ARMENIA: 2 ♂♂, Armenian SSR, Vedi, 19 Jun. 1982, Kalashjan leg. (GS); 1 ♂, Vedi distr., Gorovan Sands, 22 Jun. 1992, I. Osipov vendit (GS); 2 ♂♂, Vedi, Garavanskie peski [= Gorovan Sands], 6 – 8 Jun. 1994, P. Kazarjan leg. (MU); 2 ♂♂, Ararat prov., Goravan village subr., semi-desert, 3 – 8 Jun. 2013, T. Ghrejyan leg. (MU); 1 ♂, Vedi, Gorovan 17 – 20 Jun. 2013, Skoupy leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C5564AFE9AB0C1.taxon	materials_examined	. LIBYA: 2 ♂♂, Trip. [Tripolitania], Sirte, 21 May 1936, R. e C. Koch leg. (MU). – EGYPT: 1 ♂, Kirdassah [= Kirdasah], 6 May [19] 33, W. Wittmer leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C556B1FD92B09F.taxon	materials_examined	ALGERIA: 1 ♂, paratype, Tamrit, Tassili n’Ajjer, 1700 m, 6 – 8 May 1987 (NMP).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509330FFC914C5571CFAE2B110.taxon	materials_examined	TURKEY: 1 ♀, Adiyaman, Halfeti, Birecik, 20 May 1998, Snizek leg. (MU). – IRAN: 2 ♂♂, Zagros Mt., prov. Fars, Shiraz – Guyom eoj., 6 Jun. 2011, J. Dalihod leg. (MU). – SYRIA: 3 ♂♂, environs of Palymra, 25 May 2009, M. Šárovec leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55528FB23B301.taxon	materials_examined	: 32 specimens of both sexes from MOROCCO (different localities in the regions of Souss-Massa, Marrakech-Safi, Rabat-Salé-Kenitra, Fes-Meknès).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55484FE3EB36B.taxon	materials_examined	: 65 specimens of both sexes from GREECE (different localities in the provinces of West Macedonia, East Macedonia, Thessaly, Peloponnese, Crete id., Kos id.) and TURKEY (different localities in the provinces of Izmir, Aydin, Isparta, Ankara, Nevsehir, Çankiri).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C553E0FE39B217.taxon	materials_examined	102 specimens of both sexes from TURKEY (different localities in the provinces of Isparta, Antalya, Kirsehir, Mersin, Sivas, Erzurm, Bingol, Hatay); IRAN (provinces of Golestan and Zanjan); SYRIA (Qal’at al Hisn; Djebel el-Druze); LEBANON (Laqlouq) and JORDAN (Zubya Nat. Park).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55573FCF2B3DE.taxon	materials_examined	32 specimens of both sexes from RUSSIA (Khasan reg., Vitjaz Bukhta) and JAPAN (Osaka, Tondabayashi).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55104FAD2B72D.taxon	materials_examined	UZBEKISTAN: 1 ♂, Buchara, Koron 15 km from Ekoc. Dzejran, 23 May 1992, Snizek leg. (MU); 2 ♂♂, 1 ♀, Buchara, Repetek, May 1900, coll. Hauser (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C5516EFBBEB7A5.taxon	materials_examined	UZBEKISTAN: 1 ♂, Kul’dzhuktau, Kizilkum desert, Haloxylon (saxaul) in the sand, 15 May 1961, Dlusskij leg. (MU). – TAJIKISTAN: 2 ♂♂, near Schaartuz [= Shahrtuz], Dzharkurgan env., 7 Apr. 2007, O. Pak leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C555DDFE76B032.taxon	materials_examined	74 specimens of both sexes from GERMANY (Gossenhain); AUSTRIA (Kaprum); ITALY (different localities in the regions of Lombardia, Trentino Alto Adige, Friuli Venezia Giulia, Veneto); ROMANIA (Hunedoara); ESTONIA (Palupohja; Saaremaa id.) and RUSSIA (Vladivostok).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55661FB5EB0E3.taxon	materials_examined	JAPAN: 4 ♂♂, 1 ♀, Nagano prov., Oshiha.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C551D5FE46B473.taxon	materials_examined	examination based on unpublished photos of the holotype provided by Denis Keith.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C55220FCFFB4D7.taxon	materials_examined	ALGERIA: 1 ♂, south of Bou Sàada, 1875, R. Oberthur leg. (GS); 10 ♂♂, 2 ♀♀, Bou Saada, 7 Apr. 1981, Pierotti leg. (MU). – TUNISIA: 2 ♂♂, 2 ♀♀, Kasserine, Magel Bel Abbes (Feriana-Gafsa), 30 Mar. 2013 G. Sabatinelli leg. (MU). – ISRAEL: 2 ♂♂, 1 ♀, Netanya (city, south), 17 Mar. 2016, G. Wizen leg. (MU).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509333FFCA14C552C4FDBBB52B.taxon	materials_examined	. “ AFRICA EQUAT. [= SOUTH SUDAN] ”: 1 ♂, syntypus, Laboré, Oct. [18] 92, E. Dabbene [leg.] (MSNG). – UGANDA: 5 ♂♂, 2 ♀♀, Bululo, Oct. 1909, Dr. e E. Bayon [leg.], det. E. Benderitter 1927 (MSNG); 2 ♂♂, Kabulamuliro, 1910, Dr. e E. Bayon [leg.], det. E. Benderitter 1927 (MSNG).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509332FFCB16DD525AFEC0B18F.taxon	description	urn: lsid: zoobank. org: act: BE 8189 F 2 - 2044 - 4 EB 2 - BAE 8 - 43 C 00824 B 437	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509332FFCB16DD525AFEC0B18F.taxon	type_taxon	Type species Araboplia lorisi gen. et sp. nov.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509332FFCB16DD525AFEC0B18F.taxon	diagnosis	Differential diagnosis Within Popilliina, it is most close to the genus Dicranoplia Reitter, 1903, from which it is distinct by the following characters states (on male): clypeus simple, not raised; peculiar shape of the anterior claw; spur of the anterior tibia short and blunt.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509332FFCB16DD525AFEC0B18F.taxon	description	Description Clypeus with anterior margin regular, poorly raised from lateral margins, not sinuate or notched in the middle. Basal margin of pronotum broadly rounded, with the three-segmented partition typical of Popilliina (median part appearing truncate) almost imperceptible. Elytra narrow, flattened, with maximum width just below the humeral region, then narrowing towards the apex. In dorsal view, the lateral margin is completely visible and straight in its medial part. In lateral view, the lateral margin is strongly sinuate, protruding in a large sub-humeral lobe and retreating in a broad concavity until the apical round. Fore legs: tibia with two well-developed apical teeth, the position of the third tooth is marked by a hardly noticeable knob, apical tooth acute but with blunt apex, proximal tooth obtuse, both non-spiniform; apical spur vestigial, not visible from above as almost non-emerging from its socket; internal claw broader than the external one, asymmetrically separated at the apex, the internal margin bearing a broad and low expansion defining a deep and narrow basal notch, external claw simple. Mid legs: external claw separated at the apex, longer and much more curved than the internal one, which is not separated. Posterior legs: tibia enlarged from the base to the apex, not sinuated, with two lateral carinae; the proximal carina vestigial, placed at about mid length, only developed near the dorsal edge, and bearing 4 – 5 long setae, thick but not spiniform; the distal carina complete (covering the whole width of the tibia), placed near the apex, at about 5 / 6 of the whole length, and bearing short spiniform setae; both claws not separated, the external larger than the other but similar in shape.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509332FFCB16DD525AFEC0B18F.taxon	etymology	Etymology The name of the genus is derived from the area of occurrence of the species (the Arabian Peninsula) and the suffix – oplia, for assonance with other genera of Rutelinae including Dicranoplia Reitter, 1903, the closest genus.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509335FFCF16275104FE5EB006.taxon	description	urn: lsid: zoobank. org: act: 15 AF 1535 - 9 CC 8 - 44 A 4 - A 6 DA-FCBF 9 B 869739 Figs 1 – 9, 14	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509335FFCF16275104FE5EB006.taxon	diagnosis	Diagnosis Same as for genus.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509335FFCF16275104FE5EB006.taxon	etymology	Etymology The specific epithet is dedicated by MU to his father, Loris Uliana, with thanks for his constant encouragement and support of his interest in natural sciences and love for insects during the years of his youth. In addition, the two subjects are alike in being covered with white hairs.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509335FFCF16275104FE5EB006.taxon	materials_examined	Material examined Holotype SAUDI ARABIA: 1 ♂, near Ar Riyad, 1989, [Ex] Collection David Král Praha (NHMB).	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
039AB4509335FFCF16275104FE5EB006.taxon	description	Description Characters mentioned in the diagnosis and the description of the genus are not repeated. Colour of integument Body black, shining, without coloured metallic reflections. Abdominal sternites dark brown. Elytra uniformly pale yellow. Appendages rusty orange except the apex of the first antennal article, the anterior and the median femora, which are slightly darkened, and the posterior femora, which are dark brown. Vestiture The whole body, especially the head and pronotum, are densely covered with long, fine white hair-like setae, except tibia (fine setae present, but sparse and mixed with spiniform setae), tarsi (only spiniform setae are present), antennal articles 5 – 10 (only short sensillar setae are present on antennal club), scutellum (bearing few short hairs), and elytra (glabrous). Morphology HEAD. Coarsely and densely punctate, punctures confluent and producing irregular transverse wrinkles on the clypeus, less densely confluent and not forming transverse wrinkles on the rest of the head. PRONOTUM. Densely punctured, punctures much finer than those of the head, confluent, well impressed along their anterior border, evanescent along the posterior one. Punctures evenly distributed except for a very narrow, poorly defined, medial stripe extending along the basal third. Lateral sides of pronotum finely bordered, base bordered except in front of the scutellum. ELYTRA. Leaving the propygidium uncovered for about half of its length. Each elytron with seven recognizable striations, all punctate. Striations 1 – 4 well impressed along their whole length, 5 very weak, only visible in its distal portion, 6 – 7 well impressed only in the basal portion. Intervals with sparse weak punctures, the whole surface with smooth and irregular transverse corrugations. EPIPLEURA. Poorly separated from the rest of the elytron, enlarged and faintly swollen at the sub-humeral lobe, narrow on the rest of its length, progressively fading in the apical part. Epipleural membrane present only in the apical quarter. A tiny spine present at the sutural angle of each elytron. Pygidium gibbose, with a strongly protruding convexity, covered with dense, sharp and evenly distributed punctures. ANTENNA. 9 - segmented, article 5, measured along its posterior border, about as long as articles 3 – 4. Club about 1.4 × as long as the rest of the antenna. LEGS. Medial legs normally developed, posterior legs comparatively much broader, in particular the tibia, which is about 1.8 × as long as wide (maximum width at the distal end) and about 3.4 × the maximum width of the median tibia. Apex of posterior tibia pitted with 13 – 14 large, deep sockets, hosting spiniform setae similar to those of the apical carina. PARAMERES. As in Figs 6 – 7.	en	Uliana, Marco, Sabatinelli, Guido (2017): Araboplia lorisi new genus and species of Rutelinae from Saudi Arabia (Coleoptera, Scarabaeidae), with comments on the subtribe Popilliina. European Journal of Taxonomy 373: 1-12, DOI: 10.5852/ejt.2017.373
