identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039F7945842BFFFCBC75F98B67CB20B7.text	039F7945842BFFFCBC75F98B67CB20B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria Tomlin 1930	<div><p>Genus  Plagigeyeria Tomlin, 1930</p><p>Type species:  Geyeria plagiostoma A. J. Wagner, 1914</p><p>Diagnosis</p><p>The genus is represented by solid shells with rather thick walls and expanded, outward reflexed labrum with mostly sinuated lateral profile and sinuated columellar margin, by teleoconch surface covered with weaker or coarser, sometimes sinuated regular axial ribs diagonally cross-connected by weaker or coarser crenate spiral lamellae, by cancellated nepionic whorl covered by fine, very close-set axial riblets cross-sected with closely set spiral ribs, by protoconch surface covered by fine irregularly crenated spiral riblets with crenated or malleated inter-rib spaces and malleated nucleus.</p><p>Anatomy</p><p>The anatomy of  Plagigeyeria montenegrina Bole, 1961 from Obodska Pećina, Rijeka Crnojevića, Montenegro was described by Bole 1970 (p. 105–106, fig. 7C).</p><p>Distribution</p><p>The distribution of  Plagigeyeria in the studied area represents a hotspot within the Neretva and South Bosnia River drainage basins ranging from Ilidža near Sarajevo southwards to Mostar, Tihaljina and Trebinje (Fig. 18). The closest adjacent distribution of the genus represents a second distribution hotspot at the basin of river Zeta, Kotor Bay and Skadar Lake Basin (as a part of Drin River drainage basin). A third smaller distribution area is isolated in northwest Kosovo (Peja) and Northeast Albania (Krumë) in the Drin drainage basin. Single species are known from, west and east Serbia, Bulgaria, as well as from South Albania. The single species from France likely represents a different genus, as well as the two species from Western Greater Caucasus assigned to  Plagigeyeria (Starobogatov 1962), which belong to a different unrelated new genus (Grego et al. in press).</p><p>Remarks</p><p>The current understanding of the genus gathers many similar morphotypes from a very wide geographical range from France to southwestern Caucasus. Herein we focus on the species of  Plagigeyeria from the studied area only, sharing the general shell morphology features with the type species so their assignment to the genus is convincing.</p></div>	https://treatment.plazi.org/id/039F7945842BFFFCBC75F98B67CB20B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458427FFFEBC25FBF16437200C.text	039F79458427FFFEBC25FBF16437200C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria plagiostoma (A. J. Wagner 1914)	<div><p>Plagigeyeria plagiostoma (A. J. Wagner, 1914)</p><p>Figs 5A, 6C</p><p>Geyeria plagiostoma A. J. Wagner, 1914: 123 (1): 46–47.</p><p>Plagigeyeria plagiostoma – Tomlin, 1930: 24.</p><p>Geyeria plagiostoma – A. J. Wagner 1928: 285, pl. 13. figs 70, 71, 73.</p><p>Plagiogeyeria  (sic) plagiostoma – Kuščer 1933: 62.</p><p>Plagigeyeria plagiostoma – Jaeckel, Klemm &amp; Meise 1957: 174. — Jaeckel 1967: 92. — Schütt 1972: 114, 119, pl. 6. fig. 1. — Willmann &amp; Pieper 1978: 126. — Radoman 1983: 107, 224, pl. VII. fig. 21 (p. 225). — Bole &amp; Velkovrh 1986: 202. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6. — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>Rather small shell (1.9 mm high) with faintly axially ribbed teleoconch, cancellate nepionic whorl and spirally ribbed protoconch with smoothly malleated nucleus, with open umbilicus, oval elongate, slightly expanded aperture.</p><p>Distribution</p><p>The species is known only from the spring of Bosna River (Vrelo Bosne) (581 m a.s.l.) near Ilidža, south of Sarajevo. This spring is draining the karst waters of the Igman Massif (1502 m a.s.l) and North Bjelašnica Massif (2067 m a.s.l.).</p><p>Remarks</p><p>The type species of the genus,  Plagigeyeria plagiostoma, with  P. inflata, were both originally described from the Bosna River Springs in Ilidža under the genus “  Geyeria ”, named after the famous German malacologist David Geyer (6 November 1855 – 6 November 1932). However the genus name was invalid due to homonymy, as it had been previously used by Buchecker in 1876 to name a moth in the family  Castniidae Boisduval, 1828, by Buckman in 1899 for a cephalopod, by Carapezzae &amp; Schopen in 1899 for a brachiopod, and by Fucini, 1901 for a cephalopod. Based on the homonymy, Tomlin in 1930 renamed the genus “  Geyeria ” as  Plagigeyeria with  Plagigeyeria plagiostoma as type species and the Vrelo Bosna (581 m a.s.l.) as type locality for the genus. The species inhabits karst conduits under the Igman Massif (1502 m a.s.l.) and northern Bjelašnica Mountains (2067 m a.s.l.), likely drained from Prečko Polje (1002 m a.s.l.).</p></div>	https://treatment.plazi.org/id/039F79458427FFFEBC25FBF16437200C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458425FFFEBC00FC7965C72486.text	039F79458425FFFEBC00FC7965C72486.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria inflata (A. J. Wagner 1928)	<div><p>Plagigeyeria inflata (A. J. Wagner, 1928)</p><p>Figs 5C, 6D</p><p>Geyeria plagiostoma inflata A. J. Wagner, 1928: 285, pl. 13. figs 68–69, 72.</p><p>Plagigeyeria plagiostoma f. inflata – Schütt 1972: 119, fig. 2.</p><p>Plagigeyeria plagiostoma – Bank 2013: Fauna Europaea v. 2.6 — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>The species can be separated from  Plagigeyeria plagiostoma by its larger shell (2.5 mm high against 1.9 mm) with a more robust and more conical shell shape with more open umbilicus and less blunt apex, by larger, more expanded aperture and by coarser ribs with different cancellate surface of the nepionic whorl.</p><p>Distribution</p><p>Only known from the type locality in Vrelo Bosne (581 m a.s.l.) near Ilidža, South of Sarajevo.</p><p>Remarks</p><p>Plagigeyeria inflata (A. J. Wagner, 1928) was described from the same spring where the empty shells of  P. plagiostoma had been present and, supposing their subspecific status and identical locality, the two species were synonymized by Schütt (1972) and then again in the Fauna Europaea (Bank 2013). Nevertheless, I have to note their conchological distinctness without intermediates, and that both species were found in the same locality only as empty shells washed together within the recent thanatocoenoses of the spring zone. We have no idea about the underground aquifer range and about its mutually separated cave branches. It is very common in the morphology of many spring caves, that the large isolated, hydrologically active cave passages converge their flow just upstream of the spring outlet. Thus we cannot exclude the presence of two separated species or populations existing within one cave system without being sympatric. The recent thanatocoenoses do not represent their true habitat. Both species are very well differentiated by their shell shape and size (Wagner 1928; Schütt 1972) without any intermediate and I prefer to treat them as two independent species.</p></div>	https://treatment.plazi.org/id/039F79458425FFFEBC00FC7965C72486	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458422FFF9BC46FEED64652486.text	039F79458422FFF9BC46FEED64652486.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria ljutaensis Grego 2020	<div><p>Plagigeyeria ljutaensis sp. nov.</p><p>urn:lsid:zoobank.org:act: A8D69E19-07EA-4432-9C5A-7FE2DA89BBC4</p><p>Figs 5D, 6A</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.029715&amp;materialsCitation.latitude=43.64461" title="Search Plazi for locations around (long 18.029715/lat 43.64461)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.029715&amp;materialsCitation.latitude=43.64461" title="Search Plazi for locations around (long 18.029715/lat 43.64461)">Herzegovina-Neretva Canton</a>, Konjic Municipality, Ljuta River spring (Fig. 2A); 43.644611° N, 18.029714° E; 383 m a.s.l; 29 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104180.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 37 shells; same data as for holotype; HNHM-MOLL-104181/1, SBMNH 626401 /6, coll. Grego /19, coll. Glöer /1 .</p><p>Dimensions (holotype)</p><p>H 2.09 mm; W 1.49 mm; BH 1.16 mm; BW 0.93 mm; AH 0.9 mm; AW 0.78 mm.</p><p>Etymology</p><p>Named after the type locality: Ljuta River spring (as called in material citation).</p><p>Description</p><p>The elongate-conical shell is 2.1 mm in height, with blunt elevated apex, milky-white colour and five inflated whorls separated by a deep suture. Shell surface finely axially ribbed with weak corrugated spiral ribs forming a cancellate structure. Prominent at the nepionic whorl. Wavy spiral cords continue towards the unribbed apex (18 cords) and to protoconch with strongly malleated nucleus. Umbilicus narrow slit-like. Excentric-oval aperture is attached to the body whorl. Peristome blunt and outward reflexed along the aperture outline. Outer lip lateral profile weakly sinuous, columellar lip is weakly concave. Columellar lip profile weakly concave. Expanded aperture conspicuously protruded against body whorl at frontal view.</p><p>Differentiating features</p><p>The new species differs from the geographically close  P. plagiostoma species from Vrelo Bosna in Ilidža (Fig. 5A) by its more conical shell with more inflated whorls, blunter apex, more narrow umbilicus and larger aperture.</p><p>Habitat</p><p>The shells of the new species were found in the sediments of a large karst spring of Ljuta River (383 m a.s.l.) (right tributary of Neretva River) rising from a small cave in a narrow gorge at the southwestern edge of Bjelašnica Massif (2067 m a.s.l.). The karst conduits are likely draining waters from northern Bjelašnica and from Džepi Polje (830 m a.s.l.). They are captured by a waterworks to supply potable water for Konjic.</p><p>Distribution</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458422FFF9BC46FEED64652486	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458423FFF8BC70FEED64652486.text	039F79458423FFF8BC70FEED64652486.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria konjicensis Grego 2020	<div><p>Plagigeyeria konjicensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 12169801-D468-422F-A065-0263523FB0B5</p><p>Figs 5E, 6B</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.020712&amp;materialsCitation.latitude=43.64413" title="Search Plazi for locations around (long 18.020712/lat 43.64413)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.020712&amp;materialsCitation.latitude=43.64413" title="Search Plazi for locations around (long 18.020712/lat 43.64413)">Herzegovina-Neretva Canton</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.020712&amp;materialsCitation.latitude=43.64413" title="Search Plazi for locations around (long 18.020712/lat 43.64413)">Konjic Municipality</a>, spring at left bank of Ljuta River 300 m west of Ljuta River spring (Fig. 2C); 43.644130° N, 18.020711° E; 345 m a.s.l.; 29 Mar. 2018; J. Grego, D. Angyal, G. Jakab, M. Olšavský leg.; HNHM-MOLL- 104174.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 21 shells; same data as for holotype; HNHM-MOLL-104175/1, SBMNH 626249 /2, coll. Grego /17, coll. Glöer /1 .</p><p>Dimensions (holotype)</p><p>H 2.09 mm; W 1.28 mm; BH 1.10 mm; BW 0.87 mm; AH 0.84 mm; AW 0.72 mm.</p><p>Etymology</p><p>Name derived from Konjic, the closest city to the type locality.</p><p>Description</p><p>Slender elongated, 2.1 mm-high shell slightly tapering towards the body whorl; milky-white colour with five convex whorls separated by a deep suture. Shell surface finely and densely axially ribbed. Very faint spiral cords are very close set, almost invisible at the teleoconch, and more prominent at nepionic whorl with 21 close-set spiral cords. Apex blunt and flat, protoconch with very faint spiral cords smoothly fading out towards the smooth, very weakly malleated nucleus. Umbilicus open. Almost round-shaped aperture weakly touching the body whorl in upper columellar part. Peristome blunt, margins reflexed along the aperture outline. Outer lip lateral profile strongly forward protruded and weakly sinuous as well as very weak sinuation present at the almost flat columellar lip profile. Expanded round aperture conspicuously protruding against the body whorl at basal view.</p><p>Differentiating features</p><p>The shells of  P. konjicensis sp. nov. differ from  P. plagiostoma (Fig 5A) and from  P. ljutaensis sp. nov. by having a significantly more elongated sub-cylindrical shell with a more blunt apex, a slightly sinuated columellar margin, a less prominent body whorl, by different protoconch structure and by a more rounded aperture.</p><p>Habitat</p><p>The empty shells were found in a smaller karst spring rising at the left bank of Ljuta River (345 m a.s.l.) among larger stone debris under the road to Ljuta village; the small rivulet from the spring joining the Ljuta River after 60 m, near the ruins of a water mill. It is likely draining the southernmost tip of Bjelašnica massif, south of the village Vrdolje (between Spiljani, Dubočani, Blace and Vrdolje).</p><p>Distribution</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458423FFF8BC70FEED64652486	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458421FFE4BC08FEED62002166.text	039F79458421FFE4BC08FEED62002166.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria mostarensis , Kuscer 1933	<div><p>Plagigeyeria mostarensis Kuščer, 1933 .</p><p>Figs 7 A–E, 8D</p><p>Plagiogeyeria  (sic) mostarensis Kuščer, 1933: 62, pl. 1 fig. 3.</p><p>Plagigeyeria mostarensis – Jaeckel, Klemm &amp; Meise 1957: 174. — Jaeckel 1967: 92. — Schütt 1972: 119, pl. 6. fig. 3. — Willmann &amp; Pieper 1978: 126. – Bole &amp; Velkovrh 1986: 202, fig. 37. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6.</p><p>Diagnosis</p><p>Shell elongate-conical with distinct regular ribbing, deep suture and very widely expanded, trumpetlike aperture stepwise pronounced at the basal view. The lateral and columellar profiles of the aperture are characteristically sinuated. Umbilicus slit-like. Nepionic whorl ribbed, only slight spiral ribs, and protoconch surface covered by irregular spiral ribs with inter-rib granulations, nucleus weakly malleated.</p><p>Distribution</p><p>Known from the springs of rivers Buna (36 m a.s.l.) and Bunica (55 m a.s.l.) southwest of Mostar. Springs are most likely draining water through karst conduits from Trusinsko Polje (866 m a.s.l.), Nevesinjsko Polje (817 m a.s.l.) and Rotimlja Polje (422 m a.s.l.).</p><p>Remarks</p><p>Kuščer (1933) described  P. mostarensis with the type locality as the vrelo “Buna” (36 m a.s.l.). However, the Buna spring hosts a different morphotype, while the typical morphotype can be found in the nearby Vrelo “Bunica” (55 m a.s.l.). He also reported the same type locality for  P. ovalis, which was not confirmed in Vrelo “Buna” but only in the Vrelo “Bunica” (see remark on  P. ovalis). Both species were also figured by Schütt as from the Vrelo “Bunica”. We suppose that Kuščer's specimens were actually collected from Vrelo “Bunica”, which most likely represents the true type locality of both taxa. It is probable that both springs were treated as the one well-known Vrelo “Buna” on the original collection label. The shells of  P. mostarensis known from the Vrelo “Buna” (Buna morphotype) (Fig. 5 D–E) differ from the typical form figured by Kuščer and known only from the Vrelo “Bunica” (Bunica morphotype) (Fig. 5 A–C) by their more pagoda shape of shell with more angled whorls with different suture and different shape of marginal sinuation. The Buna morphotype of  P. mostarensis is predominantly found in the deposits of the large main stream of Buna River. In all four small side streams at the left bank rising 300–600 m downstream from the main Buna resurgence the empty shells of  P. pseudocostellina sp. nov. (Fig. 5 G–H) can be found. These are separated by a straight, not sinuated, rounded aperture very similar to  Paladilhiopsis (Costellina) turrita (Kuščer, 1933) (known only from the small spring Izvor in Jadro Valley near Split, Croatia). The taxonomical importance of the Buna morphotype is still under investigation and for the time being, it is listed under  P. mostarensis .  Plagigeyeria reischuetzorum described herein from Vrelo “Vrijeka” in Dabarsko Polje (472 m a.s.l.) in the upper Bregava Basin, which is likely draining the karstwater from Slato Polje (1012 m a.s.l.) and Lukavačko Polje (895 m a.s.l.), is morphologically the closest relative of  P. mostarensis . The karst conduits of Bregava Basin indicate at least an occasional divergence of its waters into the Vrelo “Bunica” (55 m a.s.l.), thus their close relation is not surprising.</p></div>	https://treatment.plazi.org/id/039F79458421FFE4BC08FEED62002166	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945843FFFE6BC13FDAA64C427F1.text	039F7945843FFFE6BC13FDAA64C427F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria pseudocostellina Grego 2020	<div><p>Plagigeyeria pseudocostellina sp. nov.</p><p>urn:lsid:zoobank.org:act: 09A7B3DE-5B03-46E1-A0F9-A07BFEDBCA4B</p><p>Figs 7 G–H, 8 B</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Mostar Municipality</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Blagaj</a>, left bank springs of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Buna River</a> 400 m from main <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Buna River
Spring</a>, Vrelo “Buna”; 43.256090° N, 17.900073° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104164.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 15 shells; same data as for holotype; SBMNH 626414 /10, coll. Grego /9 •   26 shells; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Mostar Municipality</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.900072&amp;materialsCitation.latitude=43.25609" title="Search Plazi for locations around (long 17.900072/lat 43.25609)">Blagaj</a>, spring at left bank of Buna River, under road, 500 m from main Buna River Spring; 43.256090° N, 17.900073° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM MOLL 104177 /1, coll. Grego /24, coll. Glöer /1  •   5 shells; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.89738&amp;materialsCitation.latitude=43.25652" title="Search Plazi for locations around (long 17.89738/lat 43.25652)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.89738&amp;materialsCitation.latitude=43.25652" title="Search Plazi for locations around (long 17.89738/lat 43.25652)">Mostar Municipality</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.89738&amp;materialsCitation.latitude=43.25652" title="Search Plazi for locations around (long 17.89738/lat 43.25652)">Blagaj</a>, spring at left bank of Buna River, under road close to bridge towards mosque, 600 m from main Buna River Spring; 43.256518° N, 17.897381° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; coll. Grego /5  .</p><p>Dimensions</p><p>Holotype: H 2.34 mm; W 1.30 mm; BH 1.28 mm; BW 1.00 mm; AH 0.97 mm; AW 0.95 mm.</p><p>Paratype: H 2.10 mm; W 1.55 mm; BH 1.34 mm; BW 1.03 mm; AH 1.00 mm; AW 0.86 mm.</p><p>Etymology</p><p>Named after its resemblance to  Costellina turrita (Kuščer, 1933), known from a small spring named Izvor in the valley of the Jadro near Split, Croatia.</p><p>Description</p><p>Elongate-pyramidal shell 2.3 mm high, with rounded blunt apex; milky-yellowish corneous translucent colour with five convex whorls separated by a deep suture. The shell surface is densely covered by lamellar ribs, the ribbing is absent at the nepionic whorl, where it is replaced by 24 densely set spiral ribs cross-sected by fine axial growth lines forming a cancellate structure. Spiral ribs fading down adapically, in pre-nepionic whorl forming faint transversal intercostal structures. Wavy spiral ribs continue towards the nucleus with intercostal granulation. Umbilicus open. Aperture elongate oval to rounded, expanded, shortly attached to the body whorl. Peristome blunt, and outward reflexed along the aperture outline. Outer lip lateral profile straight and flat as well as flat along the columellar lip. Aperture slightly, sometimes conspicuously protruded against the teleoconch and body whorl profile at basal view.</p><p>Differentiating features</p><p>The new species differs from  P. mostarensis (Fig. 5 A–C) mainly by its straight labral and columellar margins and more open umbilicus. While possessing a similar shell morphology to  Costellina turrita (Kuščer, 1933) (Fig. 5F) from the family  Moitessieriidae, it differs by having a more open umbilicus and a more oval and more open aperture. Shell morphometry comparison with the related species is presented in the Table 1.</p><p>Habitat</p><p>The type specimens were found in sandy sediments washed out from the four small side springs at the right bank of Bunica River about 300–600 m from the main spring of Buna River (36 m a.s.l.). The position of sandy sediments and condition of the fragile shells suggest better adaptation of the new species to an interstitial habitat than we experienced in any other species of  Plagigeyeria, which, in contrast, are more specialised to open phreatic cave habitats. The small springs likely draining the low karst plateau situated at the southern foot of the mountain range west of Blagaj without interconnection to the main karst conduit of the Buna.</p><p>Distribution</p><p>The new species is so-far known only from the four small springs of the type locality at the left bank of river Buna in Blagaj, while a local Buna form of  P. mostarensis (Fig. 5 D–E) can be found in the sediment of the main spring of Buna River. The typical morphotype of  P. mostarensis is found in the nearby spring of river Bunica near Hodbina (Fig. 5 A–C) and we suppose this is the type locality, which was erroneously confused with the Buna spring by Kuščer (1933).</p><p>Remarks</p><p>The shell morphology similarity to the moitessieriid  Paladilhiopsis (Costellina) turrita could suggest an interlink towards  Plagigeyeria mostarensis and other members of the genus  Plagigeyeria, and its possible affiliation to the family  Moitessieriidae .</p></div>	https://treatment.plazi.org/id/039F7945843FFFE6BC13FDAA64C427F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945843DFFE0BC1AFB3D65BF27BB.text	039F7945843DFFE0BC1AFB3D65BF27BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria reischuetzorum Grego 2020	<div><p>Plagigeyeria reischuetzorum sp. nov.</p><p>urn:lsid:zoobank.org:act: A15894D4-DED0-4678-938A-FE39F32E0C51</p><p>Figs 8A, 9 G–J</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; Republika Srpska, Bileća Municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.239166&amp;materialsCitation.latitude=43.074722" title="Search Plazi for locations around (long 18.239166/lat 43.074722)">Berkovići</a>, Vrelo “Vrijeka” in Dabarsko Polje NW of Bjeljani (Fig. 2 B–G); 43.07472222° N, 18.23916667° E; 483 m a.s.l.; Aug. 2009; Alexander Reischütz, Nicole Steiner-Reischütz and Peter L. Reischütz leg.; NHMW 111678.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 49 shells; same data as for holotype; NHMW 111679/1, HNHM-MOLL-104165/1, coll. Reischütz /45, coll. Grego /2 •  21 shells; same locality as holotype; Mar. 2010; Alexander Reischütz, Nicole Steiner-Reischütz and Peter L. Reischütz leg.; coll. Reischütz/19, coll. Grego/2 •  3 shells; same locality as holotype; 31 Mar. 2018; J. Grego, D. Angyal, G. Jakaba and M. Olšavský leg.; SBMNH 626404 /1, coll. Grego /2 •  18 shells; same locality as holotype; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; coll. Grego /16 .</p><p>Other material</p><p>BOSNIA AND HERCEGOVINA • 3 shells; Republika Srpska, Bileća Municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.348642&amp;materialsCitation.latitude=43.018543" title="Search Plazi for locations around (long 18.348642/lat 43.018543)">Orahovice</a>, Estavela “Obod” in Fatničko Polje; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; 43.018543° N, 18.348643° E; 452 m a.s.l.; coll. Grego/3.</p><p>Dimensions</p><p>Holotype: H 2.73 mm; W 1.73 mm; BH 1.66 mm; BW 1.19 mm; AH 1.29 mm; AW 1.05 mm.</p><p>Paratypes:</p><p>H 2.51 mm; W 1.73 mm; BH 1.66 mm; BW 1.19 mm; AH 1.36 mm; AW 1.19 mm.</p><p>H 2.64 mm; W 1.68 mm; BH 1.64 mm; BW 1.22 mm; AH 1.27 mm; AW 1.15 mm.</p><p>H 2.42 mm; W 1.69 mm; BH 1.53 mm; BW 1.19 mm; AH 1.16 mm; AW 1.16 mm.</p><p>Etymology</p><p>Named after the Austrian malacologists Alexander and Peter L. Reischütz with Nicole Steiner-Reischütz, who were the first to collect the species in the type locality.</p><p>Description</p><p>Shell is narrow-conical, 2.7 mm high with rounded apex and yellowish corneous periostracum, with five convex pagoduline whorls with rather weak suture. Shell surface finely axially ribbed at late whorls of the teleoconch. Faintly cancellate nepionic whorls covered by 25 close-set spiral ribs. Density of spiral ribs apically decreasing at protoconch to 15 and fading out at malleate nucleus. Space among the protoconch ribs is covered by characteristically granulated surface. Umbilicus narrow slit-like and obscured by the strongly reflexed columellar marginal fold. Larger expanding reflexed trumpet-like aperture irregularly oval and attached to body whorl by a parietal callus. Peristome sharp, and strongly outward reflexed along the aperture outline. Outer lip lateral profile and columellar lip profile characteristically sinuous. Expanded aperture conspicuously protruded against shell outline at basal view.</p><p>Differentiating features</p><p>The shells of  P. reischuetzorum sp. nov. differ from the specimens of  P. mostarensis known from the spring of the Bunica River near Hodbina (Fig. 5 A–C) by their more inflated whorls and less sinuated labral and columellar margins, as well by a more prominent and a less declined aperture at the shell lateral outline. Shell morphometry comparison with the related species is presented in the Table 1.</p><p>Habitat</p><p>The type specimens were found in sediments washed out from the cave spring “Sušica Pećina” and its side permanent spring Vrelo “Vrijeka” (483 m a.s.l.) situated 100 m eastward from the cave and rising from debris under the road to Fatnica. It is a part of northern Bregava Basin, likely draining the karst waters from Lukavačko Polje (895 m a.s.l.) into Dabarsko Polje (472 m a.s.l.). During the sampling in March 2018 the adjacent parts of Dabarsko Polje were flooded by large spring outlets creating a vast lake.</p><p>Distribution</p><p>The new species is so-far known only from the type locality.</p></div>	https://treatment.plazi.org/id/039F7945843DFFE0BC1AFB3D65BF27BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945843BFFE2BC43FAF267AC2405.text	039F7945843BFFE2BC43FAF267AC2405.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria lewarnei Grego 2020	<div><p>Plagigeyeria lewarnei sp. nov.</p><p>urn:lsid:zoobank.org:pub: CC8487E0-169D-4B32-A2AB-F9B34DFB3F3B</p><p>Figs 9D, 11B</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.368095&amp;materialsCitation.latitude=42.713737" title="Search Plazi for locations around (long 18.368095/lat 42.713737)">Republika Srpska</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.368095&amp;materialsCitation.latitude=42.713737" title="Search Plazi for locations around (long 18.368095/lat 42.713737)">Trebinje Municipality</a>, Gorica, Vrelo “Vruljak 1”, sandy sediment taken by SCUBA dive ca 200 m from the cave entrance (Fig. 2F); 42.713736° N; 18.368096° E; 280 m a.s.l.; 6 Sep. 2009; Gergely Balázs. leg.; HNHM MOLL 104176.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 1 fragmented specimen as for holotype; same data; HNHM MOLL 105099 /; Bosnia and Hercegovina •   7 shells; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.30331&amp;materialsCitation.latitude=42.714256" title="Search Plazi for locations around (long 18.30331/lat 42.714256)">Republika Srpska</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.30331&amp;materialsCitation.latitude=42.714256" title="Search Plazi for locations around (long 18.30331/lat 42.714256)">Trebinje Municipality</a>, Vrelo “Tučevac” at right bank of Trebišnjica River, 42.714257° N; 18.303311° E; 280 m a.s.l.; 31 Mar. 2018; J. Grego, M. Olšavský, G. Jakab and M. Kováčiková leg.; SBMNH 626406 /1 fragmented specimen, JG F1336 /6 fragmented specimens  .</p><p>Dimensions</p><p>Holotype: H 1.95 mm; W 1.32 mm; BH 1.21mm; BW 1.05 mm; AH 0.93 mm; AW 1.05 mm.</p><p>Etymology</p><p>Named after Brian Lewarne from The Devon Karst Research Society, Plymouth UK, who leads the “Proteus Project” hypogean species conservation project in Trebinje and supported this study by extensive consultancy about karst groundwater circulation on the studied area.</p><p>Description</p><p>Elongate conical shell 1.95 mm high with four convex whorls (apex of holotype missing) separated with rather weak suture. Shell surface covered by slightly sinuated lamellate axial ribs cross-sected by slightly crenated fine spiral cords. Twelve spiral cords are more prominent at the nepionic whorl forming a cancellate surface. Faint spiral ribs continue through the apex where the axial ribs are missing and slowly fade out at the protoconch covered by malleate structure. Umbilicus narrow slit-like and obscured by the strongly reflexed columellar marginal fold. Expanding trumpet-like reflexed aperture with typically sinuated labral margin adapically forward protruding. The ventral shape of the aperture is oval, protruded against the shell outline and slightly labrally depressed, attached to the body whorl at its columellar side. Peristome blunt and strongly outward reflexed along aperture outline.</p><p>Differentiating features</p><p>The shell morphology and surface sculpture of the new species are closely related to those of  P. mostarensis, however the new species differs significantly by its much smaller shell with different shape, by more blunt-rounded apex and by the differently shaped aperture, more protruding at its lower margin. Furthermore the radial malleated cord structure of the first protoconch whorl is weaker in the new species. The new species represents an intermediate between  P. mostarensis and  P. olsavskyi by its protruding lateral labral profile, however, it differs from the latter by a more closed umbilicus and different shell surface sculpture and by a more smooth protoconch surface.</p><p>Habitat</p><p>Only scarce, eroded or fragmented shells have been recovered from the spring and cave sediments of the spring at the right bank of Trebišnjica in Trebinje. This suggests a deep cave habitat remote from the spring head or from the hitherto known cave passages. The cave in the type locality drains the karst aquifers north of Trebinje likely from Ljubomirsko Polje (506 m a.s.l.).</p><p>Distribution</p><p>The new species is so-far known only from the type locality and from the Vrelo “Tučevac” in Trebinje. Fragments resembling this species were found in sediments of “Sušicka Jama” Estavelle at the border with Montenegro.</p></div>	https://treatment.plazi.org/id/039F7945843BFFE2BC43FAF267AC2405	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458439FFEDBC0BF86865A4266E.text	039F79458439FFEDBC0BF86865A4266E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria zetatridyma (Schutt 1960)	<div><p>Plagigeyeria zetatridyma (Schütt, 1960)</p><p>Plagigeyeria zetaprotogona zetatridyma Schütt, 1960: 149, fig. 4.</p><p>Plagigeyeria zetaprotogona zetatridyma – Schütt 1972: 121, pl. 7. figs 14–16.</p><p>Plagigeyeria zetatridyma – Bole &amp; Velkovrh 1986: 202, fig. 37. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13.</p><p>Diagnosis</p><p>Large species (max 3.1 mm) with turreted spire and inflated body whorl, very faint and close-set axial striae at shell surface and corrugated varix, faint spiral ribs at early whorls and very expanded trumpetlike aperture with distinctly broadly folded adapical notch groove as well as smoothly impressed posterior channel. Columellar labral fold covering the umbilicus.</p><p>Distribution</p><p>Plagigeyeria zetatridyma has its main distribution range over Upper Nikšićko Polje (647 m a.s.l.), Krupac Polje (615 m a.s.l.), Slano Polje (613 m a.s.l.), Slivlje Polje (601 m a.s.l.) and in the main spring of Zeta River in Glava Zete (88 m a.s.l.) as well as in the left side springs of the Zeta River Valley (Viska Vrela, 47 m a.s.l.) in Montenegro. The empty shells of the species were detected by H. Schütt at Nikšićko Vrelo (342 m a.s.l.) at Miruše, in the basin of the Trebišnjica River, indicating the karstwater and its species divergence between basins of river Drin (Zeta) and Trebišnjica.</p><p>Remarks</p><p>Specimens of  P. zetatridyma detected by Schütt (1963 and 1972) in one of the main springs of Trebišnjica (Nikšićko Vrelo, 342 m a.s.l.), show extraordinarily high shell morphology resemblance to the morphotype of the same taxon known from springs (spring Močila) and estavelles (Gornjepoljski Vir, 643 m a.s.l.) at the north-western edge of Upper Nikšićko Polje (Montenegro). Its presence in a spring supplying the Trebišnjica River Basin represents additional proof of the transboundary groundwater communication with the Zeta River Basin at Nikšičko Polje. It is also possible, that the empty shells are washed from their Zeta Basin habitats into the neighbouring Trebišnjica aquifer only by the interim high water levels. Unfortunately the localities have been flooded by a hydropower dam so, we have lost the chance to get genetical proof of such groundwater divergence.</p></div>	https://treatment.plazi.org/id/039F79458439FFEDBC0BF86865A4266E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458436FFECBC50FAA46465251D.text	039F79458436FFECBC50FAA46465251D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria erossi Grego 2020	<div><p>Plagigeyeria erossi sp. nov.</p><p>urn:lsid:zoobank.org:act: 405BAA2B-296D-4B18-9D62-343C35A6963E</p><p>Figs 9 A–C, 10A</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.741081&amp;materialsCitation.latitude=43.6173" title="Search Plazi for locations around (long 17.741081/lat 43.6173)">Bosnian Federation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.741081&amp;materialsCitation.latitude=43.6173" title="Search Plazi for locations around (long 17.741081/lat 43.6173)">Herzegovina-Neretva Canton</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.741081&amp;materialsCitation.latitude=43.6173" title="Search Plazi for locations around (long 17.741081/lat 43.6173)">Donja Jablanica</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.741081&amp;materialsCitation.latitude=43.6173" title="Search Plazi for locations around (long 17.741081/lat 43.6173)">Komadinovo Vrelo</a>, 2 km SW of, and under the road from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.741081&amp;materialsCitation.latitude=43.6173" title="Search Plazi for locations around (long 17.741081/lat 43.6173)">Mostar</a> to Jablanica, side outlet of main spring at left bank of Neretva (Fig. 2D); 43.617298° N, 17.741081° E; 180 m a.s.l.; 30 Jul. 2016; J. Grego and Z. P. Erőss leg.; HNHM-MOLL-104166.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 3 shells; same data as for holotype; HNHM-MOLL-104167/1, coll. Grego /1, coll. Erőss /1 . •  1 shell; same locality as holotype; 29 Mar. 2018; Jozef Grego leg; coll. Grego /1 .</p><p>Dimensions</p><p>Holotype: H 2.10 mm; W 1.13 mm; BH 1.25 mm; BW 0.90 mm; AH 0.87 mm; AW 0.72 mm.</p><p>Paratypes:</p><p>H 2.01 mm; W 1.28 mm; BH 1.12 mm; BW 0.93 mm; AH 0.88 mm; AW 0.77 mm.</p><p>H 1.79 mm; W 1.09 mm; BH 1.12 mm; BW 0.88 mm; AH 0.79 mm; AW 0.70 mm.</p><p>Etymology</p><p>Named after my friend Zoltán Péter Erőss, president of the Hungarian Malacological Society, Budapest, who participated in the field trip and brought my attention to the locality of the new species.</p><p>Description</p><p>Solid, narrow, conical shell, 2.1 mm in height, with blunt apex, milky-white colour and four convex whorls separated by a deep suture. Shell surface very finely axially ribbed especially at the late teleoconch and ribs cross-sected by irregular wavy faint spiral structures. Nepionic whorl covered by 18 close-set spiral ribs, continuing apically into fine irregular granulose cord-like structure at protoconch and smoothly malleate nucleus. Umbilicus open, partly obscured by the strongly reflexed columellar margin. Larger expanding reflexed aperture oval, attached to the body whorl in its upper columellar part. Peristome blunt, reflexed along aperture outline. Outer lip lateral profile characteristically sinuous adapically as well as having a characteristic sinuation which is present at the columellar lip. Expanded aperture stepwise protruded against body whorl at frontal view.</p><p>Differentiating features</p><p>Compared with the geographically closest  Plagigeyeria mostarensis Kuščer, 1933 (Hercegovina, Blagaj) (Fig. 7 A–E), the new species differs by its smaller cylindrical and less conical shell shape, blunter rounded apex and less prominent protoconch, more open umbilicus, as well as by a proportionally smaller and more oval aperture. The new species has a characteristic sinuation at outer and inner lips and a less prominent aperture within the shell peripheral outline. The shell morphology of the new species is closely related to  Plagigeyeria plagiostoma (A. J. Wagner, 1914) from the spring of the river Bosna near Ilidža, which differs by its more slender shell, cylindrical shape, more open umbilicus, different sculpture of the protoconch, proportionally smaller aperture and a weaker marginal sinuation. Shell morphometry comparison with the related species is presented in the Table 2.</p><p>Habitat</p><p>The type locality is a large karst spring rising at the right bank of Neretva River among large stone blocks under highway (Sarajevo-Mostar) and draining the limestone massif of Mt Prenj (2115 m a.s.l.) in Čvrsnica Massif (highest peak Pločno, 2228 m a.s.l.). Due to a strong current in the spring zone, there was not enough fine sediment in the main spring zone, but empty shells had been found in two smaller side springs at the left side of the main outlet. This species most likely inhabits an undiscovered karst conduit upstream the spring inside the Čvrsnica Massif.</p><p>Distribution</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458436FFECBC50FAA46465251D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458437FFE9BC45F94B649B2215.text	039F79458437FFE9BC45F94B649B2215.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria olsavskyi Grego 2020	<div><p>Plagigeyeria olsavskyi sp. nov.</p><p>urn:lsid:zoobank.org:act: 24E627A4-A78B-4447-BB44-B34BB9D0F943</p><p>Figs 5 F–I, 10C–D</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; Herzegovina-Neretva Canton, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.628004&amp;materialsCitation.latitude=43.169697" title="Search Plazi for locations around (long 17.628004/lat 43.169697)">Studenci</a>, Vrelo “Kajtazovina” in Studeničko Polje (Fig. 3D); 43.169696° N, 17.628004° E; 53 m a.s.l.; 1 Apr. 2018; J. Grego, D. Angyal, G. Jakab, M. Olšavský leg; HNHM-MOLL-104178.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 3 shells; same data; coll. Grego/3 •  45; same locality as holotype; 22 Apr. 2019; J. Grego, M. Olšavský, G. Jakab and M. Kováčiková leg.; HNHM-MOLL-105100/1, coll. Grego /44 •  41 shells; West Herzegovina Canton, Ljubuški district, Donji Proboj, Vrelo “Mali Prokop” 400 m north of road M6; 43.223905° N, 17.514440° E; 98 m a.s.l. (Fig. 4C); 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; SBMNH 626260 /7, coll. Grego /32, coll. Reischütz /1 .</p><p>Dimensions</p><p>Holotype: H 2.01 mm; W 1.52 mm; BH 1.22 mm; BW 1.07 mm; AH 0.9 mm; AW 0.93 mm.</p><p>Paratypes:</p><p>H 2.00 mm; W 1.32 mm; BH 1.25 mm; BW 0.99 mm; AH 0.94 mm; AW 0.79 mm.</p><p>H 2.03 mm; W 1.42 mm; BH 1.25 mm; BW 1.01 mm; AH 0.85 mm; AW 1.07 mm.</p><p>H 1.87 mm; W 1.34 mm; BH 1.19 mm; BW 0.96 mm; AH 0.86 mm; AW 0.69 mm.</p><p>Etymology</p><p>Named after my friend Mário Olšavský, speleologist and geologist from Banská Bystrica, Slovakia, who actively helped with the sampling of springs in our field trips.</p><p>Description</p><p>Elongate-pyramidal 2 mm-high shell with flat blunt apex and four convex whorls separated by a deep suture; milky-white colour and distinctly open umbilicus. Shell surface covered by fine regular axial ribs at last two teleoconch whorls, nepionic whorl with cancellate surface and dominating axial ribs, embryonal whorls covered by irregular faint spiral rib-like structure with raised malleate structures at the interspaces. Rounded or slightly oval aperture with slightly outward reflexed margins. Sharp peristome slightly expanded along aperture outline. Outer lip lateral profile forward protruding and very weakly sinuous adapically and straight to very weakly sinuated at the columellar lip. The aperture is conspicuously stepwise protruded against the shell outline at the basal view.</p><p>Differentiating features</p><p>The shell morphology of the new species closely resembles  P. inflata (A. J. Wagner, 1928), differing by its smaller and less conical shell, less open umbilicus, more blunt apex and a more rounded shape of the aperture. From  P. plagiostoma it differs by having a more robust shell shape with a more prominent body whorl and broader umbilicus. Shell morphometry comparison with the related species is presented in the Table 2.</p><p>Habitat</p><p>The type lot was found in the sediments of two springs at a left tributary of the Tihaljina (Trebižat) River:</p><p>1. Vrelo “Kajtazovina” (53 m a.s.l.) in Studeničko Polje (LT): a large karst spring with a small cave and ruins of a water mill with a bridge at its mouth. Spring likely draining the water from Čitlučko Polje (217 m a.s.l.).</p><p>2. Vrelo “ Mali Prokop” (98 m a.s.l): A small spring rising from a crack in the limestone massif just beside the road to the northernmost houses of Donji Proboj Village. The small amount of sandy sediment is accumulated at the bottom of the shallow spring basin. The spring outlet looks to be almost constant during the year and not influenced by the spring maxima in the other neighbouring springs. It is likely draining the water from Rašanjsko Polje (344 m a.s.l.).</p><p>Distribution</p><p>Only known from the type locality and the nearby Vrelo “ Mali Prokop” in Donji Proboj in the left tributary of the Tihaljina (Trebižat) River.</p></div>	https://treatment.plazi.org/id/039F79458437FFE9BC45F94B649B2215	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458432FFEBBC72FE50656A2239.text	039F79458432FFEBBC72FE50656A2239.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria listicaensis Grego 2020	<div><p>Plagigeyeria listicaensis,  sp. nov.</p><p>urn:lsid:zoobank.org:act: BBEACB84-641D-4E59-B97A-C748FF43816F</p><p>Figs 7 E–F, 8B</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; Bosnian Federation, West Herzegovina Canton, Široki Brijeg, Vrelo Lištica 2 (BiloVrilo) 1 km north of the city (Fig. 4B); 43.396392° N, 17.596437° E; 319 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104168.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 37 shells; same data as for holotype; HNHM-MOLL-104169/1, SBMNH 626250/4, coll. Grego /31, coll. Glöer /1 .</p><p>Dimensions</p><p>Holotype: H 2.00 mm; W 1.30 mm; BH 1.23 mm; BW 1.03 mm; AH 0.91 mm; AW 0.81 mm.</p><p>Paratype: H 2.21 mm; W 1.47 mm; BH 1.28 mm; BW 1.09 mm; AH 0.98 mm; AW 0.88 mm.</p><p>Etymology</p><p>Named after the type locality in the springs of Lištica River in the vicinity of Široki Brijeg.</p><p>Description</p><p>Solid elongate-conical shell, 2.0 mm high, apex blunt, slightly elevated protoconch, with four convex whorls and deep suture. Shell colour is translucent-whitish. Teleoconch surface covered by fine axial ribs and fine, close-set spiral cords forming a distinct cancellate structure. With dominating axial ribs. Nepionic whorl covered by 25 wavy granulose close-set spiral ribs connected by granulose irregular structures fusing at malleate apex. Open umbilicus, partly obscured by reflexed columellar margin. The elongate-oval reflexed aperture attached to body whorl by a narrow sulcus. Peristome reflexed along the aperture outline. Outer lip weakly sinuous, straight at its columellar side and slightly protruded adapically. Columellar lip with straight lateral profile. Aperture at basal view stepwise protruded against body whorl.</p><p>Differentiating features</p><p>Compared to  P. olsavskyi sp. nov., differs by its broader, more conical shell, less blunt protoconch, a proportionally larger aperture, more closed umbilicus and more coarsely sculptured protoconch. Shell morphometry comparison with the related species is presented in the Table 2.</p><p>Habitat</p><p>The type locality consists of two large karst spring risings at Lištica valley 1 km north of Široki Brijeg at 319 m a.s.l. The southern spring is captured for a water supply, while the northern one has a deep spring lake with free outflows through old water mill ruins, where the sandy sediment samples were taken. The spring is most likely getting its water through karst conduits from Rakitno Polje (890 m a.s.l.) and from the sinkholes around the upper Ugrovača River.</p><p>Distribution</p><p>The species is known only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458432FFEBBC72FE50656A2239	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458430FFEABC4AFE746465213D.text	039F79458430FFEABC4AFE746465213D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria ozimeci Grego 2020	<div><p>Plagigeyeria ozimeci sp. nov.</p><p>urn:lsid:zoobank.org:act: D4D9C7F0-9C1D-44EF-9933-C27CB712A994</p><p>Figs 11A, 12 F–K</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; West Herzegovina Canton, Vitina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.485964&amp;materialsCitation.latitude=43.237595" title="Search Plazi for locations around (long 17.485964/lat 43.237595)">Vrelo “Vrioštica”</a> (Fig. 3E); 43.237595° N, 17.485963° E; 98 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg; HNHM-MOLL-104173.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 8 shells; same data as for holotype; HNHM-MOLL-104179/1, SBMNH 626245/2, coll. Grego /5 •  2 shells; same locality as for holotype; 22 Apr. 2019; J. Grego, M. Olšavský, G. Jakab and M. Kováčiková leg.; coll. Grego /2;</p><p>Dimensions</p><p>Holotype: H 1.97 mm; W 1.45 mm; BH 1.28 mm; BW 1.03 mm; AH 1.03 mm; AW 0.83 mm.</p><p>Paratypes:</p><p>H 1.82 mm; W 1.38 mm; BH 1.17 mm; BW 1.00 mm; AH 0.98 mm; AW 0.79 mm.</p><p>H 1.90 mm; W 1.28 mm; BH 1.24 mm; BW 0.93 mm; AH 1.00 mm; AW 0.86 mm.</p><p>Etymology</p><p>Named after my friend Roman Ozimec from Zagreb, Croatia who has contributed much to the speleobiology and cave exploration of the Dinarides.</p><p>Description</p><p>Narrow conical shell 1.97 mm high, with blunt globose apex and four convex whorls and semi-deep suture. Shell yellowish corneous colour with rusty-brown inorganic incrustations. Teleoconch surface covered by fine regular axial ribbing cross-sected by irregular spiral cord-like structures forming a distinct cancellated surface. The embryonal whorl surface has 16 irregular spiral cords with malleated interstices and a number of cords gradually decreasing towards the malleate apex. The body whorl could bear 2–4 distinct sharp varices. Umbilicus open. Aperture elongate-rhomboid shaped, labrally depressed. Margins outward reflexed. Outer lip lateral profile has sinuous protrusion at depressed part. Columellar lip lateral profile straight with slight elevation at its central part. Expanded aperture stepwise protruding against the teleoconch basal periphery outline.</p><p>Differentiating features</p><p>Shells of  P.ozimeci sp. nov. can be found sympatrically with  P.vriosticaensis sp. nov. and can be distinguished by its smaller, more conical shell shape with lower numbers of whorls and more robust shell shape with depressed labral margin. Shells of  P. ozimeci frequently have on their body whorl one or more varices with sharp, winged frame not seen in other species of  Plagigeyeria . From other similar sized members of the genus, it can be differentiated by its proportionally larger and more elongated aperture with labrum elevated and depressed at its middle part. The columellar peristome has a characteristic double sinuation not seen in other species. Shell morphometry comparison with the related species is presented in the Table 2.</p><p>Habitat</p><p>The type locality is a large karst spring (98 m a.s.l.) rising from a cave under a limestone cliff in the town of Vitina. A stone wall holding a pond directs the springwater to a small disused powerplant at the left side of spring zone. A significant amount of water is also leaking under the stone wall to a lower, shallow ornamental duckpond at the northern edge of the town park. Vrelo “Vrioštica” is likely supplied by water drained in Rakitno Polje (890 m a.s.l) and through Kočerinsko Polje (302 m a.s.l.).</p><p>Distribution</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458430FFEABC4AFE746465213D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458431FFD5BC52FD6864652533.text	039F79458431FFD5BC52FD6864652533.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria jakabi Grego 2020	<div><p>Plagigeyeria jakabi sp. nov.</p><p>urn:lsid:zoobank.org:act: 8FFFE2C7-2DB1-4F0E-9FAD-D8027A4DD709</p><p>Fig. 12 A–B</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.602928&amp;materialsCitation.latitude=43.181473" title="Search Plazi for locations around (long 17.602928/lat 43.181473)">Herzegovina-Neretva Canton</a>, Studenci-Guljevina, Vrilo “Guljevina” (Fig. 4A); 43.181472° N, 17.602929° E; 77 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104172.</p><p>Paratype</p><p>BOSNIA AND HERCEGOVINA •  1 shell; same data as for holotype; coll. Grego /1 .</p><p>Dimensions</p><p>Holotype: H 2.52 mm; W 1.58 mm; BH 1.45 mm; BW 1.14 mm; AH 1.17 mm; AW 1.00 mm.</p><p>Paratype: H 2.45 mm; W 1.40 mm; BH 1.45 mm; BW 1.14 mm; AH 1.13 mm; AW 1.07 mm.</p><p>Etymology</p><p>Named after my friend Gabriel Jakab, a speleologist from Plešivec, Slovakia, who supported all our Balkan field trips and helped to explore the type locality of the new species.</p><p>Description</p><p>Solid elongate-conical 2.5 mm-high shell with blunt rounded apex, milky-yellowish colour and 4.5 convex inflated whorls separated by a deeply cut suture. Shell surface regularly axially ribbed with faintly granulose interstices. Nepionic whorl almost smooth with faint spiral rib-like structures. Umbilicus narrow, slit-like, partly obscured by the reflexed columellar marginal fold. The larger expanding reflexed aperture is tear-shaped with a sinulus-like extension at its upper-right end. Peristome blunt and trumpetlike reflexed along aperture outline, especially at upper marginal part. Outer lip lateral profile very weakly sinuous and characteristic sinuation present at columellar lip profile. Expanded aperture stepwise and conspicuously protruded against body whorl at basal view.</p><p>Differentiating features</p><p>P. jakabi sp. nov. differs from all related species like  P. angyaldorkae sp. nov. by a typically tear-shaped aperture with a sharply conspicuous indication of a posterior channel and an oval rounded bottom part of the aperture and, by a less sinuated labral margin and by a more prominent body whorl. Shell morphometry comparison with the related species is presented in the Table 3.</p><p>Habitat</p><p>The shells of the new species were washed out from subterranean aquifers through a large karst spring “Vrilo” in Guljevina (77 m a.s.l.). The spring rising under the thick vertical Neogene limestone cliff with a smaller cave left from the outlet and water mill ruins downstream. It is likely draining the water from Mokro Polje (230 m a.s.l.).</p><p>Distribution</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458431FFD5BC52FD6864652533	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945840EFFD6BC61F97864682238.text	039F7945840EFFD6BC61F97864682238.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria angyaldorkae Grego 2020	<div><p>Plagigeyeria angyaldorkae sp. nov.</p><p>urn:lsid:zoobank.org:act: 6BC5A082-0F63-4116-92B3-3BD320D5D1FA</p><p>Fig. 12E</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • West Herzegovina Canton, Tihaljina District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.373026&amp;materialsCitation.latitude=43.3229" title="Search Plazi for locations around (long 17.373026/lat 43.3229)">Lukende, Jakšenica Village, “Vrelo Jakšenica”</a> (Fig. 3F); 43.322897° N, 17.373026° E; 173 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104163.</p><p>Dimensions</p><p>Holotype: H 3.07 mm; W 1.86 mm; BH 1.45 mm; BW 1.31 mm; AH 1.21 mm; AW 1.21 mm.</p><p>Etymology</p><p>Named after my friend Dorottya Angyal from the Budapest Natural History Museum, who substantially helped with the samplings during our field trip.</p><p>Description</p><p>Solid, elongate-conical shell, 3.1 mm-high, with 5.5 regularly tapering narrow inflated whorls, deep suture and blunt rounded apex, milky-white colour and fine regular sinuated axial ribs. Late protoconch faintly irregularly spirally ribbed. Umbilicus open, obscured by the outward reflexed flaring columellar margin. Expanding reflexed aperture with irregular elongate-oval shape and a slightly depressed and inward-bent labral margin. The upper labral margin is elongate-flaring. Labral lateral profile protruded and bent backward at the body whorl, columellar margin profile sinuated. Expanded upper tip of aperture is conspicuously protruded, flaring against the frontal shell profile.</p><p>Differentiating features</p><p>P. angyaldorkae sp. nov. differs from all other known species of the region by its elongate-conical shell shape with 6 whorls, by more convex whorls and proportionally smaller body whorls and by a characteristically elongated labrally depressed aperture with a slightly depressed labral margin. From  Lanzaia ephantota (Megerle von Mühlfeld, 1824) it differs by larger more elongate shell wit finer, more close-set ribbs and more depressed aperture. Shell morphometry comparison with the related species is presented in the Table 3.</p><p>Habitat</p><p>The empty shell of this subterranean species was washed out by a middle-sized karst spring (173 m a.s.l.) rising from a 1.5 m-broad limestone crack among houses in the village. The strong stream flows under the main road, depositing the sand at a pond just before the old watermill ruins. The spring is likely getting its water from sinkholes north of village Drinovci (245 m a.s.l.) in Imotsko Polje.</p><p>Distribution</p><p>Only known from the type locality</p></div>	https://treatment.plazi.org/id/039F7945840EFFD6BC61F97864682238	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945840DFFD6BC6EFE7264C32486.text	039F7945840DFFD6BC6EFE7264C32486.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagigeyeria vriosticaensis Grego 2020	<div><p>Plagigeyeria vriosticaensis sp. nov.</p><p>urn:lsid:zoobank.org:act: BE0E3ADC-7F3F-4866-9BCD-2769746A2A81</p><p>Fig. 12 C–D</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; West Herzegovina Canton, Vitina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.485964&amp;materialsCitation.latitude=43.237595" title="Search Plazi for locations around (long 17.485964/lat 43.237595)">Vrelo “Vrioštica”</a> (Fig. 3E); 43.237595° N, 17.485963° E; 98 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104182.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 4 shells; same data as for holotype; SBMNH 626400 /2, coll. Grego /2 .</p><p>Dimensions</p><p>Holotype: H 2.82 mm; W 1.76 mm; BH 1.66 mm; BW 1.31 mm; AH 1.21 mm; AW 1.17 mm.</p><p>Paratype: H 3.00 mm; W 1.45 mm; BH 1.72 mm; BW 1.28 mm; AH 1.24 mm; AW 1.28 mm.</p><p>Etymology</p><p>Named after the type locality: spring of Vrioštica River.</p><p>Description</p><p>Elongate oval shell with 4.5 convex whorls and deep suture, 2.8 mm high, with blunt rounded apex, milky-white colour with rusty incrustations. Shell surface shiny and finely axially ribbed at the teleoconch. Nepionic whorl faintly spirally ribbed. Umbilicus narrow, slit-like. Aperture rhomboid-oval with outward reflex, posteriorly and anteriorly more expanded. Peristome blunt, and reflexed along the aperture outline. Outer lip lateral profile straight and sinuous posteriorly. Sinuation present at columellar marginal profile. Aperture protruded against teleoconch outline from frontal view.</p><p>Differentiating features</p><p>The species is closely related to  P. angyaldorkae sp. nov., from which it differs by having a smaller number of whorls and a more prominent body whorl and, by a more narrow umbilicus and different shape of aperture. Conchologically, it represents an intermediate shell morphology between  P. angyaldorkae sp. nov. and  P. jakabi sp. nov., from which it can be distinguished by its more robust shell shape and different shape of the aperture with much broader posterior channel and by a smaller umbilicus. From sympatric  P. ozimeci sp. nov. it differs by a larger shell with more whorls, a less conical shell shape and by a different shape of the aperture. Shell morphometry comparison with the related species is presented in the Table 3.</p><p>Habitat</p><p>See the habitat of  P. ozimeci sp. nov.</p><p>Distribution</p><p>The species is only known from the type locality.</p></div>	https://treatment.plazi.org/id/039F7945840DFFD6BC6EFE7264C32486	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945840BFFD0BC03FEED675925F7.text	039F7945840BFFD0BC03FEED675925F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana Grego & Gloer 2019	<div><p>Genus  Travunijana Grego &amp; Glöer, 2019</p><p>Until now, this recently described genus was recognised only from the type locality of its type species (Vrelo “Goricki Studenac” in Gorica near Trebinje). However, closer investigation of the protoconch structure as well as of the general shell morphology supported the transferral of several species formerly assigned to  Plagigeyeria to the genus  Travunijana .</p><p>Differential diagnosis</p><p>The genus  Travunijana can be clearly distinguished from the genus  Plagigeyeria by its overall pitted surface sculpture of the protoconch without any spiral cords.  Plagigeyeria has a faintly malleated and radially structured protoconch with spiral cords, malleated or granulated inter-rib spaces without a pitted surface. Furthermore, the shells of  Travunijana are more solid with no flaring and less expanded apertures and with straight lateral labral profile and a characteristic smooth sinuation at the columellar margin. The lateral labral profile of  Plagigeyeria is mostly sinuated, especially around the anterior and posterior canals and the part of the columellar margin adjacent to columella is straight or weakly sinuated. The peristome of  Plagigeyeria is more reflexed and more widely expanded. The shell surface of  Travunijana is lacking the inter-rib axially malleated surface structures of  Plagigeyeria . The genus  Belgrandia Bourguignat, 1870 with its Balkan representatives has also a pitted protoconch, however the present pitting is deeper and coarser than that of  Travunijana . The genus  Belgrandia can be distinguished by a distinct callous varix at the labrum or at the labral part of the body whorl.</p><p>Anatomy</p><p>The anatomy of the male genitals of the type species is described by Grego &amp; Glöer (2019) (p. 88: fig. 6).</p><p>Distribution</p><p>The genus  Travunijana is known from the karst regions adjacent to the South Dalmatian coastline from Gradac and Ploče to Dubrovnik, and about 32 km inland from the Adriatic coastline to Prolog, Vrgorac, Svitava, Veličani and to Trebinje. Further northwards its distribution is known from Trebinje to Bileća and towards Gatačko Polje and Dabarsko Polje to Vrelo “Bunica” in Hodbina (Fig. 19). The distribution ranges of  Travunijana and  Plagigeyeria overlap only in the area of Hodbina, Dabarsko Polje and north of Trebinje, where both genera could occur sympatrically.</p><p>Type species</p><p>Travunijana vruljakensis Grego &amp; Glöer, 2019, holotype HNHM MOLL 104416.</p><p>Other taxa assigned to the genus</p><p>Plagigeyeria ovalis Kuščer, 1933,  Plagigeyeria robusta Schütt, 1959,  Plagigeyeria robusta asculpta Schütt, 1972,  Plagigeyeria klemmi Schütt, 1961,  Plagigeyeria nitida Schütt, 1963,  Plagigeyeria angelovi Schütt, 1972,  Plagigeyeria edlaueri Schütt, 1961,  Plagigeyeria tribunicae Schütt, 1963 and  Travunijana gloeri sp. nov.</p></div>	https://treatment.plazi.org/id/039F7945840BFFD0BC03FEED675925F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F7945840BFFD3BC27F931634F206B.text	039F7945840BFFD3BC27F931634F206B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana vruljakensis Grego & Gloer 2019	<div><p>Travunijana vruljakensis Grego &amp; Glöer, 2019</p><p>Figs 5B, 14A</p><p>Travunijana vruljakensis Grego &amp; Glöer, 2019: 88, figs 2–6.</p><p>Diagnosis</p><p>Robust elongate-conical shell (2.7–2.9 mm) with prominent body whorl, fine axial growth lines, blunt apex, closed umbilicus, unprotruding aperture with straight labral profile and sinuated columellar aperture margin. Penis morphology characterised by a small typical outgrowth at the left side, near the blunt penis tip. Teleoconch surface with irregular growth lines, nepionic whorl with irregular flatnodulous malleation and protoconch surface weakly regularly pitted.</p><p>Distribution</p><p>The species is known only from its type locality at Vrelo “Goricki Studenac” at Gorica near Trebinje and in the nearby Vrelo “Vruljak 1”.</p><p>Remarks</p><p>The type species of the genus can be classified as crenobiont, however the white animal and reduced eyes could suggest an adaptation to deeper crenobiont zone at the border of stygobiont habitat. The species was not found inside the deep cave zone of the adjacent cave Vrelo “Vruljak 1”. The sexual dimorphism reported for this species (Grego &amp; Glöer 2019) can be very likely generalised to the whole genus and could explain the shell morphology variability of other species assigned herein to the genus.</p></div>	https://treatment.plazi.org/id/039F7945840BFFD3BC27F931634F206B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458408FFD3BC2AFCA2637025A8.text	039F79458408FFD3BC2AFCA2637025A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana nitida (Schutt 1963) Grego 2020	<div><p>Travunijana nitida (Schütt, 1963) comb. nov.</p><p>Fig. 13F</p><p>Plagigeyeria nitida Schütt, 1963: 210–211, fig. 8.</p><p>Plagigeyeria nitida – Schütt 1972: 116, 119, pl. 6 fig. 8. — Bole &amp; Velkovrh 1986: 202 — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6 — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>Elongate-oval hard-shelled species (3.6 mm) with smooth shiny shell surface and weak umbilical keel present close to the umbilicus. Elongate oval aperture obscuring the umbilicus and only slightly protruding at the basal view. Lateral labral profile is straight and columellar labrum possesses a weak fold.</p><p>Distribution</p><p>The empty shells are known only from springs at the southern end of Hutovo (Svitavsko) Blato (Vrelo “Sopot Mlin” and others at Sjekoše and Bajovci 2 m a.s.l.) (Schütt 1972). These springs receive their karst water from the main swallow holes of the River Trebišnjica at the end of Popovo Polje (227 m a.s.l.).</p><p>Remarks</p><p>The species is typical with its carinated umbilicus and elongated smooth shell surface. It is well separable from the sympatric  T. edlaueri (Schütt, 1961), which has a larger, more conical shell with a more inflated aperture, but both still keeping the straight labral profile and weak sinulus at the columellar margin.</p></div>	https://treatment.plazi.org/id/039F79458408FFD3BC2AFCA2637025A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458408FFD2BFDAF8E267F820AF.text	039F79458408FFD2BFDAF8E267F820AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana angelovi (Schutt 1972) Grego 2020	<div><p>Travunijana angelovi (Schütt, 1972) comb. nov.</p><p>Figs 13E, 14B</p><p>Plagigeyeria nitida angelovi Schütt, 1972: 116, 119, tab. 6, fig. 9.</p><p>Plagigeyeria angelovi – Bole &amp; Velkovrh 1986: 202 – Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13.  Plagigeyeria nitida angelovi – Bank 2013: Fauna Europaea v. 2.6 – Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>Hard shell (3.3 mm) with conical spire and broader body whorl and with distinct broad and blunt umbilical keel at lower part of the body whorl. Aperture irregularly broad lens shaped and more protruding at the basal view. Lateral labral profile straight and columellar labrum possesses a weak sinuation. Protoconch surface regularly pitted.</p><p>Distribution</p><p>Empty shells are known from the Vrelo “Ombla” (2.5 m a.s.l.) in Komolac near Dubrovnik (Schütt 1972; Schütt 2000), supplied by groundwater mainly from Popovo Polje (269 m a.s.l.) and its hinterland.</p><p>Remarks</p><p>The second carinated species of the Western Trebišnjica aquifer,  T. angelovi (Schütt, 1972) has an elongate shape with a smooth surface. Originally placed by Schütt (1972) as a subspecies of  T. nitida (Schütt, 1963), it is known from springs in Hutovo (Svitavsko) Blato. However nothing common can be found on the shell morphology of both taxa except for the carinated bottom of the body whorl; so their treatment as independent species is preferred herein.  Travunijana angelovi can be scarcely found in the accessible thanatocoenoses of the Vrelo “Ombla” near Komolac. Only a few usually worn shells within each sample suggest that its remote habitat is most likely situated deep inside the mountain, far away from the spring outlet.</p></div>	https://treatment.plazi.org/id/039F79458408FFD2BFDAF8E267F820AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458409FFDCBFD7FBE764B82127.text	039F79458409FFDCBFD7FBE764B82127.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana tribunicae (Schutt 1963) Grego 2020	<div><p>Travunijana tribunicae (Schütt, 1963) comb. nov.</p><p>Figs 13C, 14 C–D</p><p>Plagigeyeria tribunicae Schütt, 1963: 208–209, figs 4–7.</p><p>Plagigeyeria tribunicae – Schütt 1972: 116, 123, pl. 8. figs 21–22. — Willmann &amp; Pieper 1978: 126 — Bole &amp; Velkovrh 1986: 202 — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013. Fauna Europaea v. 2.6 — Bank &amp; Neubert 2017: 25</p><p>Diagnosis</p><p>Flat discoid shell 1.9–2.5 mm high with prominent body whorl having angled basal keel at its lower part and an ear-shaped aperture. Spire height variable, from entirely flat to elevated broad conical shape. Shell shape unique within the genera  Travunijana and  Plagigeyeria . Lateral labral profile straight and columellar labrum possesses a weak sinuation. Protoconch surface regularly and deeply pitted.</p><p>Distribution</p><p>Only known from the springs of Trebišnjica River (342 m a.s.l.) in Bileća (Schütt 1972), currently flooded under the lake behind the Grančarevo Dam (Bilećko Jezero). The Trebišnjica River springs represent a rather complicated hydrological drainage pattern. Most of the springs receive their water supplies from Gatačko Polje (936 m a.s.l.), Cerničko Polje (816 m a.s.l.) and partly through Fatničko Polje (452 m a.s.l.), but at least two of the springs, Vrelo “Oko” and Nikšićko Vrelo, collecting also the interim divergent groundwaters from Northern Nikšićko Polje (643 m a.s.l.).</p><p>Remarks</p><p>The distinct shells of  T. tribunicae (Schütt, 1963) detected in the subsidiary spring of the Trebišnjica (Nikšićko Vrelo, 342 m a.s.l.), show a remarkable variability in spire height from flat to pyramidal form (Schütt 1972). Unfortunately the type locality is currently flooded by the Grančarevo Dam, and it is questionable whether the lake had altered the subterranean aquifer water streams in a negative way leading to the species’ extinction, or if the species might still have survived in some other part of the aquifer. The shell morphology with dominant body whorl and keeled umbilicus significantly resembles the  Plagigeyeria sp. morphotypes found at the north-western edge of Nikšićko Polje (Montenegro, 643 m a.s.l.), confirming the transboundary communication of aquifers at least during high groundwater periods. However, the pitted protoconch structure is more similar to that of the genus  Travunijana . The slightly reflexed margin is unique in the genus  Travunijana and its position within this genus is only provisional. Based on the conditions of empty shells, the habitat is most likely upstream inside the undiscovered cave system, far from the actual spring water outlet (recently under the Bilećko Jezero) where the empty shells were collected.</p></div>	https://treatment.plazi.org/id/039F79458409FFDCBFD7FBE764B82127	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458407FFDCBC2EFD6C67EB2486.text	039F79458407FFDCBC2EFD6C67EB2486.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana ovalis (Kuscer 1933) Grego 2020	<div><p>Travunijana ovalis (Kuščer, 1933) comb. nov.</p><p>Fig. 7 I–K</p><p>Plagiogeyeria  (sic) ovalis Kuščer, 1933: 62–63, pl. 1, fig. 4.</p><p>Plagigeyeria ovalis – Jaeckel, Klemm &amp; Meise 1957: 174. — Jaeckel 1967: 92. — Bole &amp; Velkovrh 1986: 202.</p><p>Plagigeyeria mostarensis ovalis – Bank 2013: Fauna Europaea v. 2.6 — Bank &amp; Neubert 2017: 25.</p><p>non  Plagigeyeria mostarensis ovalis – Schütt 1972: 119, tab. 6, fig. 4.</p><p>Diagnosis</p><p>Robust oval shell (3 mm) with prominent body whorl and conical spire, smooth or finely striated shell surface, blunt apex, narrow umbilicus and large callous irregularly oval aperture only slightly protruding at basal view. Lateral labral profile straight and columellar labrum possesses a weak-cut sinuation. Protoconch very faintly pitted.</p><p>Distribution</p><p>Known only from Vrelo “Bunica” (55 m a.s.l.) near Hodbina south of Mostar. The records of the species from springs at lower Neretva (Kuščer 1935) were not supported since and need further confirmation.</p><p>Remarks</p><p>Kuščer (1933) described Plagiogeyeria  (sic) ovalis as a subspecies of  P. mostarensis with the type locality in the Buna spring. However, many sampling attempts in the type locality (Vrelo “Buna”) have not proven the presence of the species. The species was repeatedly detected in the sediments of the Vrelo “Bunica”, a name very similar to the one given as the type locality. We suppose the true type locality of  P. ovalis is the Vrelo “Bunica”, and the original labels were likely misinterpreted in same manner as for  P. mostarensis described from the same collection at the same time from the same locality. The subspecific status of  P. ovalis was misleading for most of the following authors as Schütt (1972), figured a specimen of  P. mostarensis from Vrelo “Buna” with a less elevated and broken spire as  P. mostarensis ovalis (p. 119, pl. 6, fig. 4.). The understanding of  P. ovalis as a valid species was therefore overlooked for several decades. Nevertheless,  P. ovalis can be clearly distinguished from  P. mostarensis not only by its more robust-oval shell with different aperture shape, but also by its less conspicuous ribbing, representing rather axial growth lines, and by a substantially different straight lateral and weakly sinuated columellar marginal profile, against the strongly sinuated ones in  P. mostarensis . Furthermore, the substantially different protoconch morphology separate both taxa also on the genus level.</p></div>	https://treatment.plazi.org/id/039F79458407FFDCBC2EFD6C67EB2486	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458404FFDFBFF6FEED62EA26D1.text	039F79458404FFDFBFF6FEED62EA26D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana robusta subsp. robusta (Schütt 1959) robusta (Schutt 1959	<div><p>Travunijana robusta robusta (Schütt, 1959) comb. nov.</p><p>Figs 7 L–M, 13A, 15A–B</p><p>Plagigeyeria robusta Schütt, 1959: 186, fig. 1.</p><p>Plagigeyeria robusta – Schütt 1972: 115, 123, pl. 8 figs 23–24. — Willmann &amp; Pieper 1978: 126. — Bole &amp; Velkovrh 1986: 202. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6 — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>Robust oval-conical shell (2.1 mm) with thick walls, rounded apex, smooth or finely ribbed shell surface, elongate round-rhomboidal aperture with slightly protruded columellar fold, open umbilicus and not protruded aperture against basal shell profile. Lateral labral profile straight and columellar labrum possesses weak negative sinuation. Nepionic whorl with weakly cancellated surface and protoconch with distinct pitting. Pitting depth variable according to population: Vrelo “Vruljak 1” population with regular shallow pitting concentrating around the suture and very weak at central part of whorl; population from Vrelo “Tučevac” with overall regular deep pitting over the whole protoconch surface.</p><p>Distribution</p><p>Travunijana robusta is known from the spring group of Trebišnjica near Bileća and Vrelo Čepo near Čepelica (currently flooded by the Bilećko Jezero), and from Estavela “Obod” in Fatničko Polje (452 m a.s.l.), spring Dobričevo near Stepen in Gatacko Polje (936 m a.s.l.) (Schütt 1972), and from Vrelo “Goricki Studenac” (280 m a.s.l.), Vrelo “Vruljak 1” (282 m a.s.l.) and Vrelo “Tučevac” (275 m. a.s.l.) in Trebinje.</p><p>Remarks</p><p>Most of the robust  Travunijana specimens found in the Trebišnjica River Basin are assigned to  Travunijana robusta (Schütt, 1959) (Fig. 7 L–M) divided into two populations or maybe subspecies (Schütt 1972). The karst conduits north of the Trebišnjica valley (north of Popovo Polje) host the nominotypical subspecies. The karst conduits south of Popovo and Mokro Poljes are populated by  T. robusta asculpta (Schütt, 1972) (Fig. 7N). Furthermore, each particular locality can have specific morphotypes (e.g., population in Vrelo “Tučevac” near Trebinje hosts its specific smooth-shelled morphotype with different aperture), which are for time being regarded as falling within one morphologically variable species.</p></div>	https://treatment.plazi.org/id/039F79458404FFDFBFF6FEED62EA26D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458404FFDEBFEDFA1A622920FB.text	039F79458404FFDEBFEDFA1A622920FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana robusta subsp. asculpta (Schutt 1972)	<div><p>Travunijana robusta asculpta (Schütt, 1972) comb. nov.</p><p>Figs 7N, 15C</p><p>Plagigeyeria robusta asculpta Schütt, 1972: 115, 122, pl. 8, figs 25–26.</p><p>Plagigeyeria robusta asculpta – Bole &amp; Velkovrh 1986: 202. — Bank 2013: Fauna Europaea v. 2.6. — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>Shell very similar to nominotypical subspecies but having in general a more elevated, more conical spire with blunter apex. Surface can be smooth or finely ribbed as well as in the nominotypical morphotype. Aperture ear-shaped and missing the slightly protruded columellar fold of the nominotypical subspecies. Lateral labral profile straight and columellar labrum possesses a weak cut sinuation. Protoconch surface pitted.</p><p>Distribution</p><p>Travunijana robusta asculpta can be found in the thanatocoenoses of the Vrelo “Ombla” near Komolac (7 m a.s.l.) and Vrelo “Mlin” (2 m a.s.l.) at Mali Zaton north of Dubrovnik. Springs are draining the karst area of upper-to-central Popovo Polje (227 m a.s.l).</p><p>Remarks</p><p>The morphological variability of nominotypical subspecies and  T. robusta asculpta overlap significantly and even Schütt (2000) assigned some of specimens of  P. robusta asculpta from the type locality (Vrelo “Ombla”) to the nominotypical subspecies based upon the presence of ribs, and conversely, the few specimens collected from Trebišnjica Springs in Bileća were assigned to  T. robusta asculpta just for their lack of ribs. However the presence or absence of the fine shell ribs on the eroded shells found in the thanatocoenoses has to be carefully evaluated by their remains around the suture, as upon first sight both ribbed and smooth forms can be found in the thanatocoenoses within the springs draining the whole Eastern Trebišnjica aquifer. The main distinguishing feature of  T. robusta asculpta is in general the more elevated and more conical spire and slightly more elongated shell shape, but even this feature possesses a high variability within each locality. Until live material becomes available and larger shells material can be examined, I prefer to separate both in a provisional way on a subspecies level, based only on their zoogeographical data. It is possible that further investigation could reveal several related species in the  T. robusta complex.</p></div>	https://treatment.plazi.org/id/039F79458404FFDEBFEDFA1A622920FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458405FFDEBC24FC3362A72441.text	039F79458405FFDEBC24FC3362A72441.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana klemmi (Schutt 1961) Grego 2020	<div><p>Travunijana klemmi (Schütt, 1961) comb. nov.</p><p>Figs 13B, 16 A–B</p><p>Plagigeyeria klemmi Schütt, 1961: 133–134, fig. 2.</p><p>Plagigeyeria klemmi – Schütt 1972: 115, 119 pl. 6 figs 5–6. — Willmann &amp; Pieper 1978: 126. — Bole &amp; Velkovrh 1986: 202. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6. — Bank &amp; Neubert 2017: 25.</p><p>Diagnosis</p><p>T. klemmi has regularly narrow-conical shells (2.2 mm) with blunt apex, deep suture and widely spaced coarse ribs on shell surface, rounded callous irregular-oval shaped aperture and closed umbilicus. Lateral labral profile straight and columellar labrum only weakly sinuated. Protoconch surface overall covered by regular and deep pitting.</p><p>Distribution</p><p>T. klemmi is known from Vrelo “Stenjevac” and Vrelo “Baja” in Vrgorsko Polje (20 m a.s.l.) near Vrgorac and from Vrelo “Kutac” and other springs around Baćinska Lakes (0.8 m a.s.l.) near Ploče. The aquifer also receives its water from Rastok Polje (72 m a.s.l.).</p><p>Remark s</p><p>Travunijana klemmi represents the northernmost species of the genus. Its shell morphology has some features of the genus  Belgrandia (e.g.,  Belgrandia torifera Schütt, 1961) known from the region, however its shell is more conical and lacks the typical callous varix near the labrum.</p></div>	https://treatment.plazi.org/id/039F79458405FFDEBC24FC3362A72441	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458405FFD8BFDBF88D6290273D.text	039F79458405FFD8BFDBF88D6290273D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana edlaueri (Schutt 1961) Grego 2020	<div><p>Travunijana edlaueri (Schütt, 1961) comb. nov.</p><p>Figs 13D, 16 C–D</p><p>Plagigeyeria edlaueri Schütt, 1961:132–133, fig. 1.</p><p>Plagigeyeria edlaueri – Schütt 1972: 115, 119, pl. 6 fig. 7. — Willmann &amp; Pieper 1978: 126. — Bole &amp; Velkovrh 1986: 202. — Bodon, Manganelli &amp; Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6. — Bank &amp; Neubert 2017: 24.</p><p>Diagnosis</p><p>Shell large for the genus (3.7 mm), conical spire with regularly tapering whorls, weak suture and rather sharp apex, body whorl triangular, centrally bumped, shell surface smooth shiny, aperture wide, callous, but not expanded and conspicuously protruding at the labral view. Lateral labral profile is straight and columellar labrum possesses a weak negative sinuation at the slightly protruding columellar fold. Protoconch surface overall covered by regular, large and very deep pitting.</p><p>Distribution</p><p>Empty eroded shells of  T. edlaueri appear in the sediment of springs at the side of the Hutovo (Svitavsko) Blato depression (2 m a.s.l.) (Vrelo “Sopot Mlin” and at Sjekoše) and in Vrelo “Glušči” (4 m a.s.l.) near Metković in Croatia. The springs are draining karst water from main swallow holes of the River Trebišnjica at the end of Popovo Polje (227 m a.s.l.) and from Gradac Polje (88 m a.s.l.), and are connected to the alluvial delta of the Neretva River.</p><p>Remark s</p><p>Travunijana edlaueri is one of the largest representatives of the genus, frequently reaching over 3 mm (max. 3.7 mm). Its general shell shape is most similar to  Plagigeyeria of all the species of  Travunijana, however the pitted protoconch structure and straight labral / weakly sinuated columellar labrum indicates its position within the genus  Travunijana or maybe within a new unrecognized genus.</p></div>	https://treatment.plazi.org/id/039F79458405FFD8BFDBF88D6290273D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
039F79458403FFC6BC51FB68650B2239.text	039F79458403FFC6BC51FB68650B2239.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travunijana gloeri Grego 2020	<div><p>Travunijana gloeri sp. nov.</p><p>urn:lsid:zoobank.org:act: 2A411831-BA56-490B-8754-80A0A0EE048F</p><p>Figs 9 K–L,15D</p><p>Type material</p><p>Holotype</p><p>BOSNIA AND HERCEGOVINA • 1 shell; Republika Srpska, Bileća Municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.239056&amp;materialsCitation.latitude=43.07469" title="Search Plazi for locations around (long 18.239056/lat 43.07469)">Berkovići, Vrelo “Vrijeka” in Dabarsko Polje NW of Bjeljani</a> (Fig. 3A); 43°04′28.9″ N, 18°14′20.6″ E; 498 m a.s.l.; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; HNHM-MOLL-104418/1.</p><p>Paratypes</p><p>BOSNIA AND HERCEGOVINA • 10 shells; same data; HNHM-MOLL-104419/1, SBMNH 632722/1, coll. Grego /8, coll. Glöer /1 •  2 shells; same locality as for holotype; D. Angyal, J. Grego, G. Jakab and M. Olšavský leg.; coll. Grego /2 .</p><p>Other material</p><p>BOSNIA AND HERCEGOVINA • 6 shells; Republika Srpska, Bileća Municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.348642&amp;materialsCitation.latitude=43.018543" title="Search Plazi for locations around (long 18.348642/lat 43.018543)">Orahovice, Estavela “Obod” in Fatničko Polje</a>; leg. 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková; 43.018543° N, 18.348643° E; coll. Grego /9.</p><p>Dimensions</p><p>Holotype: H 3.38 mm; W 1.96 mm; BH 2.23 mm; BW 1.56 mm; AH 1.51 mm; AW 1.29 mm.</p><p>Paratype: H 3.38 mm; W 1,87 mm; BH 2.31 mm; BW 1.60 mm; AH 1.42 mm; AW 1.24 mm.</p><p>Etymology</p><p>Named after my friend and renowned malacologist Peter Glöer from Hetlingen, Germany, who largely contributed to the freshwater gastropod fauna of the Balkans.</p><p>Description</p><p>Inflated conical 3.38 mm-high shell with dominant convex body whorl. Light yellowish corneous shell with smooth shiny surface, weak close-set axial ribs and 4.5 convex whorls separated by a weak suture. Protoconch surface smooth, weakly and distantly pitted with larger weak malleation over the pitting. Apex blunt and flat. Umbilicus closed. Elongate asymmetric-oval aperture attached to the body whorl by a weak sulcus. Peristome callous and blunt, not reflexed. Outer lip almost flat at and weakly sinuated at columellar profile. Aperture profile aligned with the body whorl and with the evenly tapering whorls at basal view.</p><p>Differentiating features</p><p>The shells of  T. gloeri sp. nov. differ from  T. ovalis (Fig. 7 I–K) from spring of Bunica River near Hodbina (but not found in Vrelo “Buna” as declared by Kuščer (1933) as its type locality) by its more slender shell shape, closed umbilicus and more flat protoconch. From the type species  T. vruljakensis it differs by adapically more protruded and differently shaped aperture, by more coarsely ribbed teleoconch surface and by more smooth protoconch surface. Shell morphometry comparison with the related  Travunijana species is presented in the Table 4.</p><p>Habitat</p><p>See habitat of  P. reischuetzorum sp. nov.</p><p>Distribution</p><p>The new species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/039F79458403FFC6BC51FB68650B2239	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Grego, Jozef	Grego, Jozef (2020): Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions. European Journal of Taxonomy 691: 1-56, DOI: 10.5852/ejt.2020.691
