identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039E3278FFA2FFEFDDDA21F50B17900F.text	039E3278FFA2FFEFDDDA21F50B17900F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis camposi Navas 1930	<div><p>Callibaetis camposi Navás 1930</p><p>(Figs. 1 A–1G)</p><p>Callibaetis camposi Navás 1930d: 19; Gillies 1990: 22; Domínguez et al. 2006: 112.</p><p>Known stages: I ♀</p><p>Diagnosis. Female imago: 1) forewing with pigmented C, Sc and R1 areas (Fig. 1 B); 2) thick and black cross veins in C and Sc areas (Fig. 1 F); 3) marginal intercalary veins single; 4) hind wing hyaline (Fig. 1 G); 5) hind wing with thick cross veins; 6) hind wing with quadrangular costal process; 7) hind wing with intercalary marginal veins (Fig. 1 G); 8) body covered with brown spots (Figs. 1 B–1E); 9) anterolateral spot on abdominal terga and sterna present (Figs. 1 C and 1E).</p><p>Redescription. Female imago: Head (Fig. 1 E). Light brown, covered with brownish spots; compound eyes black. Antenna with brown scape apically; pedicel dark brown apically. Thorax (Fig. 1 F). Covered with brownish spots, metascutellar protuberance rounded. Leg III, anterior surface covered with brownish spots; tibia light brown, with one dark brown mark at base and another at apex; tarsi light brown, each segment brown apically. Wings. Forewing (Fig. 1 B) with pigmented C and Sc areas; some areas of Radial 1 pigmented; veins light brown; thick and black cross veins in C, Sc and R1 areas; stigmatic area with six cross veins touching Sc vein; marginal intercalary veins single; length of each intercalary vein 0.47 × distance between adjacent longitudinal veins; length of forewing about 2.4 × width. Hind wing (Fig. 1 G) hyaline, with 13 cross veins; one to three intercalary marginal veins; costal process quadrangular. Abdomen (Figs. 1 B–1E). Terga. Yellowish, covered with brown spots, brown medially, forming longitudinal pale line; one spot anterolaterally. Sterna. Yellowish, covered with brown spots, with one mark anterolaterally. Caudal filaments lost.</p><p>Comments. The only known specimen is in very poor condition; however, it is easy to observe the black and thick cross veins in the C, Sc and R1 areas, which are conspicuous but not exclusive to this species (also present in C. (C.) radiatus and C. viviparus). A number of characteristics are shared by these three species, such as anterolateral spots on the abdominal terga and sterna; pigmented C, Sc and R1 areas; hind wing with many thick cross veins; hind wing with quadrangular costal process; and abdomen covered with spots.</p><p>Based on re-evaluation of the morphology, we cannot distinguish between the female imagoes of C. camposi, C. (C.) radiatus and C. viviparus . Synonymizations are not proposed, because the delimitations of species are better investigated when nymphs and male imagoes are studied (see Cruz et al. 2016). Only C. (C.) radiatus is described based also on nymphs and male imago. In order to avoid unnecessary taxonomic acts, we consider it prudent to await the discovery and description of the unknown stages of C. viviparus and C. camposi before making any taxonomic hypotheses.</p><p>Although the species of the genus Callibaetis have a history of vague type locales, the collection of new specimens at the type locale of C. viviparus is achievable, and also of C. camposi, but with less probability.</p><p>We designate the unique known specimen of C. camposi as the lectotype. Material examined. Callibaetis camposi, female imago (lectotype by present designation, dry), ECUADOR, Guayaquil, i.1930, Campos leg, n° 75–5308, MZB.</p><p>Distribution. Ecuador: Guayaquil.</p></div>	https://treatment.plazi.org/id/039E3278FFA2FFEFDDDA21F50B17900F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFA0FFEFDDDA24540A4C97EC.text	039E3278FFA0FFEFDDDA24540A4C97EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Abaetetuba) capixaba Cruz, Salles & Hamada 2009	<div><p>Callibaetis (Abaetetuba) capixaba Cruz, Salles &amp; Hamada 2009</p><p>(Figs. 2 A–2G)</p><p>Callibaetis capixaba Cruz, Salles &amp; Hamada 2009: 31; Cruz et al. 2014:11.</p><p>Known stages: I ♀♂, N</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Fig. 2 A); 2) dorsal portion of turbinate eyes in lateral view without constriction; 3) dorsal portion of turbinate eyes stalk height 0.9 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 5) forewing hyaline (Fig. 29 in Cruz et al. 2009); 6) marginal intercalary veins paired (Fig. 29 in Cruz et al. 2009); 7) hind wing hyaline (Fig. 30 in Cruz et al. 2009); 8) costal process of hind wing rounded (Fig. 30 in Cruz et al. 2009); 9) marginal intercalary veins on hind wing absent (Fig. 30 in Cruz et al. 2009); 10) abdominal sterna with pair of medioanterior and medioposterior sigilla weak pigmented (Fig. 2 B); 11) abdominal sterna light brown with many red spots and two red marks, one lateral and other sublateral on anterior margin (Fig. 2 B); 12) forceps segment I wide at base (Fig. 2 F); 13) forceps segment III oval (Fig. 2 F).</p><p>Female imago: 1) forewing hyaline (Fig. 33 in Cruz et al. 2009); 2) marginal intercalary veins paired (Fig. 33 in Cruz et al. 2009); 3) hind wing hyaline (Fig. 34 in Cruz et al. 2009); 4) costal process of hind wing rounded (Fig. 34 in Cruz et al. 2009); 5) marginal intercalary veins on hind wing absent (Fig. 34 in Cruz et al. 2009); 6) abdominal sterna with pair of medioanterior and medioposterior sigilla weak pigmented (Fig. 2 D); 7) abdominal sterna reddish brown with many red spots and two red marks, one lateral and other sublateral on anterior margin (Fig. 2 D).</p><p>Mature nymph: 1) distal margin of labrum medially with long, fine and apically bifid setae (Fig. 2 G); 2) maxillary palp reaching apex of galea-lacinia (Fig. 41B in Cruz et al. 2009); 3) two rows of setae on basal part of inner-dorsal row of maxilla; 4) paraglossa subrectangular with apex truncated (Fig. 42A in Cruz et al. 2009); 5) paraglossa with three tufts of fine and simple; 6) metanotum with spines; 7) foretarsus anterior surface without spine-like setae (Fig. 43A in Cruz et al. 2009); 8) hind claw with minute denticles (Fig. 44B in Cruz et al. 2009).</p><p>Comments. Many species of the genus Callibaetis have a history of poor, inaccurate, or even absent, illustrations. In order to maximize the possibility of a reliable identification, here we present photographs of the complete dorsal and ventral habitus of the male and female imagoes (Figs. 2 A–2D), a photograph of the ventral view of the male genitalia, mounted on a slide (Fig. 2 F), photographs of the dorsal habitus of the nymphal exuviae (Fig. 2 E) and a detailed illustration of the labrum (dorsal and ventral views) (Fig. 2 G). For more, see morphological comments under C. (A.) pollens .</p><p>Material examined. Callibaetis capixaba, female imago with corresponding nymphal exuviae (holotype), BRAZIL, Espírito Santo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.53539&amp;materialsCitation.latitude=-19.87525" title="Search Plazi for locations around (long -40.53539/lat -19.87525)">Santa Teresa</a>, 19°52’30.9” S / 40°32’07.4” W, pool, sand, 26.x.2008, F.F. Salles coll., INPA ; one male imago (reared), 20 nymphs, BRAZIL, Espírito Santo State, Santa Teresa Municipality, 19° 52' 30.9'' S / 40° 32' 07.4' 'W, pool, sand, 26.x.2008, F.F. Salles coll., CZNC ; one male imago and one female imago (both reared), Espírito Santo, Santa Teresa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.556084&amp;materialsCitation.latitude=-19.925028" title="Search Plazi for locations around (long -40.556084/lat -19.925028)">Reserva Biológica Augusto Ruschi</a>, 19°55’30.1” S / 40°33’21.9” W, pool, sand, 26.ii.2009, F.F. Salles coll., INPA ; one female imago, Santa Catarina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.7034&amp;materialsCitation.latitude=-26.896742" title="Search Plazi for locations around (long -51.7034/lat -26.896742)">Vargem Bonita</a>, 26°53’48.27” S / 51°42'12.24” W, 2005, E. Raimundi coll., CZNC .</p><p>Distribution. Brazil: Espírito Santo; Santa Catarina.</p></div>	https://treatment.plazi.org/id/039E3278FFA0FFEFDDDA24540A4C97EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFA0FFEADDDA226A0B6B930A.text	039E3278FFA0FFEADDDA226A0B6B930A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis gregarius Navas 1930	<div><p>Callibaetis gregarius Navás 1930 nomen dubium</p><p>(Figs. 3 A–3G)</p><p>Callibaetis gregarius Navás 1930a: 72; Gillies 1990: 24; Domínguez et al. 2006: 113; Cruz et al. 2014: 29.</p><p>Known stages. I ♂</p><p>Comments. The specimen studied at the Museu de Zoología de Barcelona (Figs. 3 A and 3B) is not a male imago, as indicated by Gillies (1990), but rather a male subimago in very poor condition. Other specimens are available in the Senckenberg Deutsches Entomologisches Institut Müncheberg (Figs. 3 C–3G), but they are also male subimagoes in poor condition. Even if additional specimens of this species could be obtained, the inadequate description, the very poor condition of the type and the vagueness of the type locality would probably minimize the possibility of a reliable identification. Based on our re-evaluation of the morphological and other evidence, we propose that Callibaetis gregarius be classified as a nomen dubium.</p><p>Material examined. Callibaetis gregarius, male subimago (labeled as type) and one male subimago, BRAZIL, Ypiranga, 19.iv.1910, MZB; three male subimagoes (labeled as syntypes) same data, Senckenberg.</p><p>Distribution. Brazil: São Paulo.</p></div>	https://treatment.plazi.org/id/039E3278FFA0FFEADDDA226A0B6B930A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFA5FFE4DDDA275A08D2978C.text	039E3278FFA5FFE4DDDA275A08D2978C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Callibaetis) guttatus Navas 1915	<div><p>Callibaetis (Callibaetis) guttatus Navás 1915</p><p>(Figs. 4 A–5D)</p><p>Callibaetis guttatus Navás 1915b: 120; Gillies 1990: 25; Da-Silva 1991: 346; Domínguez et al. 2006: 113; Nieto 2008: 232; Salles et al. 2010: 302; Lima et al. 2012: 306; Cruz et al. 2014: 30, Angeli et al. 2015: 199.</p><p>Callibaetis apicatus Navás 1917: 189 . (syn. by Gillies 1990)</p><p>Callibaetis bruchius Navás 1920b: 55 . (syn. by Gillies 1990)</p><p>Callibaetis zonatus Navás 1929: 224 . (syn. by Gillies 1990)</p><p>Known stages. I ♀♂, N</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval; 2) dorsal portion of turbinate eyes in lateral view without constriction; 3) dorsal portion of turbinate eyes stalk height 1.9× height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 3) forewing hyaline or with C, Sc, R1 and posterior margin areas with large marks; 4) marginal intercalary veins paired; 5) hind wing hyaline or with basal mark; 6) costal process of hind wing rounded; 7) marginal intercalary veins on hind wing present; 8) abdominal terga III, V and VII with lateral marks; 9) abdominal sterna with pair of medioanterior and medioposterior sigilla without pigment; 10) forceps segment I wide at base; 11) forceps segment III oval.</p><p>Female imago: 1) forewing at least with C, Sc, R1 and posterior margin areas with marks, maximal degree of pigmentation with seven complete and transversal bands (Fig. 5 C); 2) marginal intercalary veins paired (Fig. 5 C); 3) hind wing hyaline or with marks at base and middle (Fig. 5 D); 4) costal process of hind wing rounded; 5) marginal intercalary veins on hind wing present (Fig. 5 D); 6) abdominal terga II, III, V and VII with inverted V mark laterally (Fig. 5 A); 6) abdominal sterna with pair of medioanterior and medioposterior sigilla without pigment (Fig. 5 B); 7) abdominal terga and sterna with few or without spots (Figs. 5 A and 5B); 8) abdominal sterna medially on anterior margin without one large spot (Figs. 5 A and 5B); 9) abdominal terga without medial longitudinal mark (Figs. 5 A and 5B).</p><p>Mature nymph: 1) maxillary palp subequal in length than galea-lacinia; 2) below maxillary palp insertion on outer margin without tuft of robust spine-like setae (Fig. 29 in Nieto 2008); 3) paraglossa with row of spine-like setae on ventral surface (Fig. 30 in Nieto 2008); 4) segment III of labial apically rounded (Fig. 30 in Nieto 2008); 5) metanotum without spines; 6) foretarsus anterior surface without spine-like setae; 7) mid and hind claw denticles smaller than foreclaw denticles (Fig. 33 in Nieto 2008).</p><p>Comments. Nieto (2008) described variation in the fore- and hind wings of this species.</p><p>In the present work, the specimens illustrated in Figures 20, 21 a, 21b, 23, 24a and 24b in Nieto (2008) were examined. We conclude that they belong to C. (A.) fasciatus based on having forewing with three or four transverse bands of pigmentation, apical band transverse to posterior margin (see also Cruz et al. 2014 and comments for C. (A.) sellacki herein).</p><p>Callibaetis (C.) guttatus is similar to C. (C.) zonalis, C. (C.) willineri and C. (C.) jocosus . All four species share an inverted “V” mark on female abdominal terga III, V and VII and variable pigmentation on the female forewings. The female imago of C. (C.) guttatus can be differentiated from C. (C.) jocosus by the abdominal terga and sterna having few or no spots (Figs. 5 A and 5B); from C. (C.) willineri by the abdominal sterna lacking one large spot medially, on the anterior margins (Figs. 5 A and 5B); and from C. (C.) zonalis by abdominal terga without medial longitudinal mark (Figs. 5 A and 5B).</p><p>We present figures of the types of the two synonymized species (Figs. 4 A–4D) and newly collected specimens from Brazil (Figures 5 A–5D).</p><p>The specimens studied by Navás (1915b) from Prov. de Buenos Aires, Argentina, were not found, thus a lectotype was not designated.</p><p>Material examined. Male imago (type of Callibaetis apicatus), ARGENTINA, Buenos Aires, 15.v.1915, MZB ; one female imago (type of Callibaetis zonatus), ARGENTINA, Alta Gracia, 17.ii.1929, MZB ; one female imago ( Callibaetis guttatus), ARGENTINA, Alta Gracia, 27.vi.1926, C. Brush det., MZB ; one nymph, ARGENTINA: Tucumán, Depto. Tafí Viejo, Raco (km 19), A8 Palangana, 24.xi.2001 , C. Molineri coll.; one female imago reared, BRAZIL, Espírito Santo state, São Mateus, Rio São Mateus, 02.x.2007 , F. F. Salles coll., INPA.</p><p>Distribution. Argentina: Tucumán; Misiones; Buenos Aires; Alta Gracia. Brazil: Ceará; Espirito Santo; Rio de Janeiro; Pernambuco.</p></div>	https://treatment.plazi.org/id/039E3278FFA5FFE4DDDA275A08D2978C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFABFFE6DDDA23CA0B0D939A.text	039E3278FFABFFE6DDDA23CA0B0D939A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis jaffueli Navas 1918	<div><p>Callibaetis jaffueli Navás 1918 revalidated species</p><p>(Figs. 6 A–6E)</p><p>Callibaetis jaffueli Navás 1918: 214; Gillies 1990: 25; Domínguez et al. 2006: 115 (previously syn. with C. jocosus by Gillies 1990)</p><p>Known stages. I ♂.</p><p>Diagnosis. Male imago: 1) forewing with pigmentation overlapping R 1 in five regions, overlapping radial sector medially, without forming complete bands (Fig. 6 B); 2) marginal intercalary veins paired (Fig. 6 B); 3) hind wing with small brown mark at middle (Fig. 6 C); 4) hind wing with pointed costal process (Fig. 6 C); 5) marginal intercalary veins on hind wing present (Fig. 6 C); 6) abdominal terga brown, with few spots and dark longitudinal mark at middle (Fig. 6 E); 7) abdominal terga III, V and VI without lateral inverted V mark; 8) abdominal sterna covered with spots (Fig. 6 D); 9) abdominal sterna with strong pigmented medioanterior and medioposterior sigilla (Fig. 6 D).</p><p>Redescription. Male imago. Length: body, 6.5 mm; forewing, 7 mm; cerci 8.6 mm. Head. Compound eyes with brown turbinate portion. Antenna with brown scape, pedicel and flagellum. Thorax. Anteronotal protuberance brown medially; posterior scutal protuberance and scutellum brown. Anteronotal and metascutellar protuberance rounded. Legs: Tibia I with brown marks apically. Wings. Forewing with pigmented C and Sc areas, overlapping R 1 in five regions, third region overlapping Radial sector (Fig. 6 B); veins light brown; marginal intercalary veins paired; length of each intercalary vein 0.7 × distance between adjacent longitudinal veins; length of forewing about 2.6 × width. Hind wing with small brown mark at middle (Fig. 6 C); costal process pointed; three marginal intercalary veins. Abdomen. Terga. Brown, with some spots and dark longitudinal mark in middle (Fig. 6 E); terga III, V and VI without lateral inverted V mark. Sterna. Light brown, covered with spots and with well pigmented medioanterior and medioposterior sigilla (Fig. 6 D). Genitalia. Forceps light brown and badly damaged.</p><p>Comments. Callibaetis jaffueli revalidated species can be differentiated from C. (C.) jocosus, C. (C.) zonalis, C. (C.) willineri and C. (C.) guttatus by the abdominal sterna medially, on anterior margin, without one large spot (in contrast to C. (C.) willineri); marginal intercalary veins on hind wing present (in contrast to C. (C.) zonalis); terga III, V and VI without lateral inverted V mark (Fig. 6 E) (in contrast to C. (C.) guttatus and C. (C.) jocosus); and the forewing with pigmentation overlapping R 1 in five regions, overlapping radial sector medially, without forming complete bands (Fig. 6 B) (in contrast to C. (C.) jocosus).</p><p>Based on the re-evaluation of the morphological evidence, we propose the revalidation of Callibaetis jaffueli revalidated species Navás and designate the unique known specimen as lectotype.</p><p>Material examined. Callibaetis jaffueli, male imago (lectotype by present designation, dry), CHILE, Los Perales, Marga-Marga, i.1918, P. Jaffuel coll., MZB.</p><p>Distribution. Chile: Marga-Marga.</p></div>	https://treatment.plazi.org/id/039E3278FFABFFE6DDDA23CA0B0D939A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFA9FFE7DDDA27E108D996E0.text	039E3278FFA9FFE7DDDA27E108D996E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Callibaetis) jocosus Navas 1912	<div><p>Callibaetis (Callibaetis) jocosus Navás 1912</p><p>(Figs. 7 A–7B)</p><p>Callibaetis jocosa Navás 1912: 195 . (name emended by Ulmer 1920)</p><p>Callibaetis jocosus Ulmer 1920: 126; Gillies 1990: 26; Domínguez et al. 2006: 115; Cruz et al. 2014: 40. Callibaetis spegazzinus Navás 1920c: 36 . (syn. by Gillies 1990)</p><p>Callibaetis rimatus Navás 1933: 113 . (syn. by Gillies 1990)</p><p>Known stages. I ♀♂, N.</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Fig. 7 A); 2) dorsal portion of turbinate eyes in lateral view without constriction; 3) dorsal portion of turbinate eyes stalk height 1.9 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins subparallel; 5) forewing with three transversal bands on apical third, base without pigmentation (Fig. 111 in Cruz et al. 2014); 6) marginal intercalary veins paired (Fig. 111 in Cruz et al. 2014); 7) hind wing hyaline (Fig. 112 in Cruz et al. 2014); 8) costal process of hind wing rounded (Fig. 112 in Cruz et al. 2014); 9) marginal intercalary veins on hind wing present (Fig. 112 in Cruz et al. 2014); 10) abdominal terga covered with spots and terga III, V and VII with lateral, inverted V mark (Fig. 7 A); 11) abdominal sterna covered with spots and with one anterolateral mark (Fig. 7 B); 12) abdominal sterna with pair of medioanterior and medioposterior sigilla weakly pigmented (Fig. 7 B); 13) forceps segment I wide at base (Fig. 110 in Cruz et al. 2014); 14) forceps segment III oval (Fig. 110 in Cruz et al. 2014).</p><p>Female imago: 1) forewing with pigment in C and Sc areas, overpassing R1, and after R2 pigmentation lighter (Fig. 114 in Cruz et al. 2014); 2) marginal intercalary veins paired (Fig. 114 in Cruz et al. 2014); 3) hind wing with mark near costal process (Fig. 115 in Cruz et al. 2014); 4) costal process of hind wing rounded (Fig. 1151 in Cruz et al. 2014); 5) marginal intercalary veins on hind wing present (Fig. 115 in Cruz et al. 2014); 6) abdominal terga covered with spots, medially with longitudinal mark, and terga III, V and VII with lateral inverted V mark (Fig. 113 in Cruz et al. 2014); 7) abdominal sterna covered with spots (Fig. 116 in Cruz et al. 2014); 8) abdominal sterna with pair of medioanterior sigilla and medioposterior sigilla weak pigmented (Fig. 116 in Cruz et al. 2014); 9) abdominal sterna medially, on anterior margin, without one large spot.</p><p>Mature nymph: 1) maxillary palp subequal in length of galea-lacinia (Fig. 122 in Cruz et al. 2014); 2) below maxillary palp insertion on outer margin with tuft of robust spine-like setae (Fig. 122 in Cruz et al. 2014); 3) paraglossa with row of spine-like setae on ventral surface (Fig. 123 in Cruz et al. 2014); 4) segment III of labium apically rounded (Fig. 123 in Cruz et al. 2014); 5) metanotum without spines; 6) foretarsus anterior surface without spine-like setae; 7) hind claw denticles smaller than foreclaw denticles (Fig. 125D in Cruz et al. 2014).</p><p>Comments. The female imago of C. (C.) jocosus is similar to C. (C.) zonalis, C. (C.) willineri and C. (C.) guttatus (see C. (C.) guttatus comments). The female imago of C. (C.) jocosus can be differentiated from these three species by the abdominal sterna covered with spots (Fig. 116 in Cruz et al. 2014) (in contrast to C. (C.) guttatus); abdominal sterna medially, on anterior margin, without one large spot (in contrast to C. (C.) willineri); and marginal intercalary veins on hind wing present (in contrast to C. (C.) zonalis).</p><p>Callibaetis jaffueli Navás 1918 and Callibaetis stictogaster Navás 1915, previouly considered synonyms of C. (C.) jocosus by Gillies (1990), have here their species status revalidated. For the diagnoses, redescriptions and comments, see their respective species sections.</p><p>The specimens studied by Navás (1912) from São Paulo, Brazil, were not found, thus the lectotype was not designated.</p><p>Material examined. Female imago (reared), two male imagoes (reared), BRAZIL, São Paulo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.940613&amp;materialsCitation.latitude=-23.237528" title="Search Plazi for locations around (long -46.940613/lat -23.237528)">Jundiaí</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.940613&amp;materialsCitation.latitude=-23.237528" title="Search Plazi for locations around (long -46.940613/lat -23.237528)">Serra do Japií</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.940613&amp;materialsCitation.latitude=-23.237528" title="Search Plazi for locations around (long -46.940613/lat -23.237528)">Lake</a> near two houses, 23°14’15.1” S / 046°56’ 26.2” W, 14.vii.2009, 1043 m, P. Brito coll., INPA ; female imago, photograph of the type of Callibaetis stictogaster, ARGENTINA, Buenos Aires, 15.x.1913, Museo de Ciencias Naturales de La Plata, C. Bruch coll.; one male imago, BRAZIL, São Paulo, i.1915, L. Navás det., MZB ; one female imago (type of Callibaetis spegazzinus), PARAGUAY, Asunción, 10.x.1919, Spegazinni coll., L. Navás det., MZB ; one female imago reared, BRAZIL, Santa Catarina state, Dionisio Cerqueira, Linha Toldo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-53.61711&amp;materialsCitation.latitude=-26.303165" title="Search Plazi for locations around (long -53.61711/lat -26.303165)">Cachoeira do Toldo</a>, 26°18’11.4” S / 53°37’01.6” W, 649 m alt., 14–15.ix.2011, P.V. Cruz, R. Boldrini, N. Hamada and A. M. O. Pes cols., INPA ; one female imago, BRAZIL, Paraná state, Balsas Novas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.744583&amp;materialsCitation.latitude=-25.44161" title="Search Plazi for locations around (long -49.744583/lat -25.44161)">Rio Pomba</a>, 25°26’29.8” S / 49°44’40.5” W, 911 m alt., 11.ix.2011, P.V. Cruz, R. Boldrini, N. Hamada and A. M. O. Pes cols., INPA .</p><p>Distribution. Argentina: Buenos Aires. Brazil: São Paulo; Mato Grosso do Sul; Santa Catarina; Paraná. Paraguay: Asunción.</p></div>	https://treatment.plazi.org/id/039E3278FFA9FFE7DDDA27E108D996E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFAFFFE1DDDA25AC0B7891EA.text	039E3278FFAFFFE1DDDA25AC0B7891EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis nigrivenosus Banks 1918	<div><p>Callibaetis nigrivenosus Banks 1918 nomen dubium</p><p>(Figs. 8 A–8E)</p><p>Callibaetis nigrivenosa Banks 1918: 11 (name emended by McCafferty 1996) Callibaetis nigrivenosus McCafferty 1996: 231; Domínguez et al. 2006: 115.</p><p>Known stages. I ♀.</p><p>Comments. We studied the type specimen from photographs (Figs. 8 A–8E). It is fragile, badly damaged and lacks the hind wings. The only reliable structure is the forewing, which allowed us to identify the specimen to the family level ( Baetidae). The absence of hind wings in the type specimen and its description lead us to believe that these structures were always absent. The manuscript is very brief and is not illustrated. The type locality presented in the manuscript and shown on the label is not precise, referring to a broad geographical area. Even if additional specimens of this species were collected, there would be little possibility of a reliable identification using the types or the original description. Based on the lack of information in the description, the damaged type and the improbability of assigning specimens to this species, we propose Callibaetis nigrivenosus Banks as a nomen dubium.</p><p>Material examined. Callibaetis nigrivenosa, male imago (type), Ecuador, Huigar, 4,500 ft., 17 June, H. S. Parish coll., Museum of Comparative Zoology, Harvard University, MCZ-ENT 0 0 0 10057. Distribution . Ecuador: Huigar</p></div>	https://treatment.plazi.org/id/039E3278FFAFFFE1DDDA25AC0B7891EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFAEFFE3DDDA2477089B971A.text	039E3278FFAEFFE3DDDA2477089B971A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Abaetetuba) pollens Needham & Murphy 1924	<div><p>Callibaetis (Abaetetuba) pollens Needham &amp; Murphy 1924</p><p>(Figs. 9 A–10H)</p><p>Callibaetis pollens Needham &amp; Murphy 1924: 51; Nieto 2008: 235; Domínguez et al. 2006: 116; Boldrini et al. 2012: 92; Lima et al. 2012: 306; Cruz et al. 2014: 56; Boldrini &amp; Cruz, 2014: 4.</p><p>Known stages. I ♀♂, N.</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Figs. 9 A, 10A); 2) dorsal portion of turbinate eyes in lateral view with constriction (Fig. 10 B); 3) dorsal portion of turbinate eyes stalk height 1.3 × height of dorsal portion (Fig. 10 B); 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 5) forewing hyaline (Fig. 9 B); 6) marginal intercalary veins single (Fig. 9 B); 7) hind wing hyaline (Fig. 9 B); 8) costal process of hind wing rounded (Fig. 9 B); 9) marginal intercalary veins on hind wing present; 10) abdominal terga on anterior margin with a transversal mark of pigment (Fig. 10 A); 11) abdominal sterna with pair of medioanterior and medioposterior sigilla weak pigmented (Fig. 10 C); 12) abdominal sterna with black anterolateral mark (Fig. 10 C); 13) forceps segment I wide at base, with setae on inner margin (Figs. 9 E, 10G); 14) forceps segment III oval (Figs. 9 E, 10G).</p><p>Female imago: 1) head with a Y mark (Fig. 10 D); 2) forewings hyaline (Fig. 161 in Cruz et al. 2014); 3) marginal intercalary veins single (Fig. 161 in Cruz et al. 2014); 4) hind wing hyaline (Fig. 162 in Cruz et al. 2014); 5) costal process of hind wing rounded (Fig. 162 in Cruz et al. 2014); 6) marginal intercalary veins on hind wing present (Fig. 162 in Cruz et al. 2014); 7) abdominal terga reddish brown (Fig. 10 D); 8) abdominal sterna with pair of medioanterior and medioposterior sigilla weak pigmented (Fig. 10 E); 9) abdominal sterna II to IX with black mark (Fig. 10 E).</p><p>Mature nymph: 1) distal margin of labrum, medially with row of long and bifid setae (Fig. 43B in Nieto 2008); 2) maxillary palp 1.5 × length of galea-lacinia (Fig. 47 in Nieto 2008); 3) crown of galea-lacinia with three rows of setae; 4) paraglossa with rounded apex (Fig. 48A in Nieto 2008); 5) paraglossa with one tuft of fine and simple setae; 6) metanotum with spines (Fig. 10 H); 7) foretarsus anterior surface without spine-like setae (Fig. 49 in Nieto 2008); 8) hind claw with minute denticles (Fig. 51 in Nieto 2008).</p><p>Comments. Cruz et al. (2009, 2014) differentiate C. (A.) pollens and C. (A.) capixaba by the setae on the femur of the nymph (trifid setae absent on the femur of C. (A.) pollens). We studied the same specimens analyzed by Cruz et al. (2009, 2014) and some of those analyzed by Nieto (2008), and observed trifid setae on the femur of C. (A.) pollens . However, the nymphs of both species can be differentiated by the shape of the apex of paraglossa, being round in C. (A.) pollens (Fig. 48A in Nieto 2008) and truncate in C. (A.) capixaba (Fig. 42A in Cruz et al. 2009); and by the number of tufts of fine and simple setae on the apex of the paraglossa, having one in C. (A.) pollens and three in C. (A.) capixaba .</p><p>We present figures of the holotype (Figs. 9 A–9E) and more recently collected specimens assigned to C. (A.) pollens (Figs. 10 A–10H) from Brazil.</p><p>Material examined. Callibaetis pollens, photographs of the slide of the male imago (holotype), BRAZIL, Mato Grosso do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.46219&amp;materialsCitation.latitude=-27.11664" title="Search Plazi for locations around (long -65.46219/lat -27.11664)">Corumbá</a>, 14.xii.1919, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.46219&amp;materialsCitation.latitude=-27.11664" title="Search Plazi for locations around (long -65.46219/lat -27.11664)">Cornell University</a>, n° 644; two female imagoes, ARGENTINA, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.46219&amp;materialsCitation.latitude=-27.11664" title="Search Plazi for locations around (long -65.46219/lat -27.11664)">Acheral</a>, Tucumán, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.46219&amp;materialsCitation.latitude=-27.11664" title="Search Plazi for locations around (long -65.46219/lat -27.11664)">Aranillas River</a>, Cruce RP 38, 366 m, 27°06’59.9” S / 65°27’43.9” W, 13.ix.2007, C. Nieto and C. Molineri cols., CZNC ; five nymphs, ARGENTINA, Acheral, Tucumán, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.46219&amp;materialsCitation.latitude=-27.11664" title="Search Plazi for locations around (long -65.46219/lat -27.11664)">Aranillas River</a>, Cruce RP 38, 366 m, 27°06’59.9” S / 65°27’43.9” W, 28.vii.2006, C. Nieto and C. Molineri cols., CZNC ; one female imago, one male imago, eight nymphs, BRAZIL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.382385&amp;materialsCitation.latitude=-6.071" title="Search Plazi for locations around (long -47.382385/lat -6.071)">Lajeado River</a>, BR 010, under the bridge, Maranhão, 06°04’15.6” S / 47°22’56.6” W, 21.vii.2010, N. Hamada, P.V. Cruz and R. Boldrini cols., INPA ; two male imagoes, five nymphs, BRAZIL, Maranhão, Riachão, Riacho Cocal, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.456833&amp;materialsCitation.latitude=-7.2295" title="Search Plazi for locations around (long -46.456833/lat -7.2295)">Cachoeira Santa Bárbara</a>, 07°13’46.2” S / 46°27’24.6” W, 23.vii.2010, N. Hamada, P.V. Cruz &amp; R. Boldrini cols., INPA ; two female imagoes, BRAZIL, Maranhão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.469833&amp;materialsCitation.latitude=-6.529805" title="Search Plazi for locations around (long -47.469833/lat -6.529805)">Farinha River</a>, BR 010, under the bridge, 06°31’47.3” S / 47°28’11.4” W, 22.vii.2010, N. Hamada, P.V. Cruz &amp; R. Boldrini cols., INPA ; five male imagoes, seven female imagoes, 12 nymphs, BRAZIL, Ceará, Jaburu River, Ubajara, Distrito da Cachoeira do Boi Morto, 03°52’36.2” S /41°0.1’0.08” W, 26.vii.2010, N. Hamada, P.V. Cruz &amp; R. Boldrini cols., INPA ; four nymphs, BRAZIL, Brejinho, near Posto Cachoeira, Bahia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.393917&amp;materialsCitation.latitude=-13.731527" title="Search Plazi for locations around (long -45.393917/lat -13.731527)">Correntina</a>, 13°43’53.5” S / 45°23’38.1” W, 08.viii.2010, N. Hamada, R. B. Querino &amp; R. Boldrini cols., INPA ; eight nymphs, BRAZIL, Bahia, Côcos, Formoso River, highway to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.825638&amp;materialsCitation.latitude=-14.666834" title="Search Plazi for locations around (long -45.825638/lat -14.666834)">Mambaí</a> municipality, 14°40’00.6” S / 45°49’32.3” W, 11.viii.2010, N. Hamada, R. B. Querino &amp; R. Boldrini cols., INPA ; five nymphs, 2 female and 1 male (reared), BRAZIL, Mato Grosso do Sul, Bonito, Balneário municipal de Jardim <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.390003&amp;materialsCitation.latitude=-21.41786" title="Search Plazi for locations around (long -56.390003/lat -21.41786)">Rio Prata</a>, 21°25’04.3” S / 56°23’24.0” W, 15/iii/2012, P. V. Cruz coll., INPA .</p><p>Distribution. Argentina: Tucumán. Brazil: Ceará; Bahia; Goiás; Maranhão; Mato Grosso do Sul; Pernambuco; Piauí; Rondônia; Roraima.</p></div>	https://treatment.plazi.org/id/039E3278FFAEFFE3DDDA2477089B971A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFACFFF9DDDA23670DDE90E7.text	039E3278FFACFFF9DDDA23670DDE90E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Callibaetis) radiatus Navas 1920	<div><p>Callibaetis (Callibaetis) radiatus Navás 1920</p><p>(Figs. 11 A–14D)</p><p>Callibaetis radiatus Navás 1920a: 132; Gillies 1990: 26; Salles et al. 2003: 13; Domínguez et al. 2006: 116; Cruz et al. 2014: 58; García 2014: 41; Lima et al. 2016: 214; Vinasco-Mondragón &amp; Zúñiga 2016: 92.</p><p>Callibaetis venulosus Navás 1933: 114 . (syn. by Gillies 1990)</p><p>Known stages. I ♀♂, N.</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eye oval (Fig. 21 in Salles et al. 2003); 2) dorsal portion of turbinate eyes without constriction; 3) height of dorsal portion of turbinate eye stalk 0.85 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with divergent anterior and posterior margins; 5) forewings hyaline (Fig. 167 in Cruz et al. 2014); 6) thick cross veins in C and Sc areas (Fig. 167 in Cruz et al.</p><p>2014); 7) marginal intercalary vein single (Fig. 167 in Cruz et al. 2014); 8) hind wing hyaline (Fig. 168 in Cruz et al. 2014); 9) hind wing with quadrangular costal process (Fig. 168 in Cruz et al. 2014); 10) hind wing with thick cross veins (Fig. 168 in Cruz et al. 2014); 11) marginal intercalary veins present on hind wing (Fig. 168 in Cruz et al. 2014); 12) abdominal terga with black mark anterolaterally (Fig. 169 in Cruz et al. 2014); 13) abdominal sterna with a weakly pigmented medioanterior and medioposterior sigilla (Fig. 172 in Cruz et al. 2014); 14) segment I of forceps wide at base (Fig. 166 in Cruz et al. 2014); 15) segment III of forceps truncated apex (Fig. 166 in Cruz et al. 2014).</p><p>Female imago: 1) forewing with brown pigmented C, Sc and R1 areas, extending beyond R1 but not forming bands (Fig. 11 C); 2) forewing with thick and black cross veins in C and Sc areas (Fig. 11 B); 3) marginal intercalary veins single (Fig. 11 C); 4) hind wing hyaline (Fig. 171 in Cruz et al. 2014); 5) hind wing with quadrangular costal process (Fig. 171 in Cruz et al. 2014); 6) marginal intercalary veins on hind wing present (Fig. 171 in Cruz et al. 2014); 7) hind wing with thick cross veins (Fig. 171 in Cruz et al. 2014); 8) abdomen with brown spots; 9) anterolateral spot on abdominal terga and sterna present (Figs. 169 and 172 in Cruz et al. 2014); 10) abdominal sterna with a weakly pigmented medioanterior and medioposterior sigilla (Fig. 172 in Cruz et al. 2014).</p><p>Mature nymph: 1) maxillary palp 1.2 × length of galea-lacinia (Fig. 12 F); 2) tuft of robust spine-like setae below insertion of maxillary palp absent (Fig. 12 F); 3) ventral surface of paraglossa with row of spine-like setae (Fig. 12 G); 4) segment III of labial palp rounded apically (Fig. 12 G); 5) metanotum without spines; 6) anterior surface of foretarsus without spine-like setae (Fig. 13 A).</p><p>Redescription. Mature nymph: Length: body, 7.2–8.0 mm; broken cerci and terminal filament (n=1). Head. Coloration: faded in alcohol. Turbinate portion of male compound eyes yellowish brown. Antenna with small spines and fine, simple setae (Fig. 12 A). Labrum (Fig. 12 B) maximum length about 1.4 × maximum width; anterolateral margins with long spine-like setae; distal margin with spine-like setae medially; dorsal surface with many, long, fine and simple setae; distal margin with one row of fine spine-like setae ventrally; ventral surface with short, spine-like setae near lateral margin. Right mandible (Fig. 12 C) with 4 + 3 denticles; margin between prostheca and mola convex; basal half with short, fine, simple setae and pores scattered over dorsal surface. Left mandible (Fig. 12 D) with 4 + 2 denticles; margin between prostheca and mola straight; basal half with short, fine and simple setae and pores scattered over dorsal surface. Hypopharynx (Fig. 12 E). Lingua with lobe covered with small simple setae; short, fine and simple setae scattered over distal margin of superlingua. Maxilla (Fig. 12 F). Medial protuberance of galea with 1+ 3 spine-like setae. Maxillary palp short, 1.2 × length of galea-lacinia; palp segment II 1.2 × length of segment I; outer margin of segment I scattered with long, fine, simple setae; inner margin of segment II with few spine-like setae. Labium (Fig. 12 G). Glossa subequal in length to paraglossa; inner margin with 11 spine-like setae; apex with 3 – 5 long spine-like setae; outer margin with row of long spine-like setae; dorsal surface with one row of long spine-like setae near outer margin and one tuft of setae at apex. Paraglossa. Ventral surface with one row of fine and simple setae; dorsal surface with three rows of long spine-like setae. Labial palp with segment I 1.1 × length of segments II and III combined; segment I covered with micropores; inner and outer margin of segment II with six spine-like setae; dorsal surface with row of six short spine-like setae; segment III with rounded apex. Thorax. Foreleg (Figs. 13 A). Ratio of foreleg segments 0.9:1:0.6:0.3. Forefemur. Length about 4.0 × maximum width; dorsal margin with row of short, spine-like setae; apex with two robust spinelike setae; length of setae about 0.1 × maximum width of femur; anterior surface near dorsal margin with one row of spine-like setae. Tibia. One row of short spine-like setae ventrally; anterior surface with few fine and simple setae. Tarsus. Anterior and posterior surface without spine-like setae. Tarsal claw 0.5 × length of tarsus (Figs. 13 B). Femur of hind leg without trifid setae; tarsal claw with small denticles (Fig. 13 C). Abdomen. Terga. Posterior margin with regular spines (Fig. 14 A). Sterna. Surface with fine, simple setae. Paraproct (Fig. 14 B) with 24 marginal spines; surface with micropores and short, fine, simple setae. Cercus and terminal filament at base as in Figures 14 C and 14D respectively.</p><p>Comments. The female imagoes of C. (C.) radiatus, C. viviparus and C. camposi are similar and could not be distinguished based on re-evaluation of the morphological evidence. However, as stated in the comments of C. camposi, the type locality of C. viviparus can be accessed and new specimens can be collected in the future.</p><p>García (2014) recorded C. (Callibaetis) radiatus in Venezuela based on reared specimens (female, male and its nymphal exuviae). Taking into account the similarity between C. (C.) radiatus, C. viviparus and C. camposi and, in order to enable the comparisons to confirm this record, we re-describe and present new diagnosis and illustrations of the nymph described by Salles et al. (2003), as well figures of the specimen originally described by Navás (1920), here designated as lectotype (Figs. 11 A–11D).</p><p>Recently, Lima et al. (2016) and Vinasco-Mondragón &amp; Zúñiga (2016) recorded C. (C.) radiatus respectively from Bahia state (Brazil) and Colombia, all based on female imagoes. Taking into account the morphological evidence here presented, both 2016 records of C. radiatus could be applicable to C. viviparus or C. camposi . Associations between stages should be made in order to confirm or refute these records.</p><p>Material examined. Callibaetis radiatus, photographs of one female imago (lectotype by present designation), Argentina, Santa Fé, 1.iii.1918, MNHN ; one female imago, Argentina, Formosa, Reserva EL, Bagual, xi-2004, Serochi coll.; one female imago, Paraguay, Asunción, 15.x.1919, MZB ; male imago, female imago (reared) and two nymphs, BRAZIL, Minas Gerais, Viçosa, Ranário—Universidade Federal de Viçosa, 27.i.1997, E.R. Silva coll.</p><p>Distribution. Argentina: Santa Fé. Paraguay: Asunción. Brazil: Bahia; Minas Gerais.</p></div>	https://treatment.plazi.org/id/039E3278FFACFFF9DDDA23670DDE90E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFB6FFFBDDDA27060C47922A.text	039E3278FFB6FFFBDDDA27060C47922A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Abaetetuba) sellacki (Weyenbergh 1883) Weyenbergh 1883	<div><p>Callibaetis (Abaetetuba) sellacki (Weyenbergh 1883)</p><p>(Figs. 15 A–16B)</p><p>Cloe sellacki Weyenbergh 1883: 164 . (comb. by Eaton 1883)</p><p>Cloe lorentzii Weyenbergh 1883: 167 . (syn. by Gillies 1990)</p><p>Callibaetis sellacki Eaton 1883: 198; Gillies 1990: 27; Nieto 2008: 237; Domínguez et al. 2006: 116; Cruz et al. 2014: 60. Callibaetis trifasciatus (partim) Navás 1915b: 120. (misidentification, the name is not syn. Gillies 1990; McCafferty 1996) Callibaetis fasciatus Ulmer 1921: 246 . (misidentification, the name is not syn. Gillies 1990) Callibaetis lineatus Navás 1932: 82 . (syn. by Gillies 1990)</p><p>Baetis gloriosus Navás 1923: 2 . syn. nov.</p><p>Callibaetis gloriosus Navás 1930b: 360 . syn. nov. (comb. by Navás 1930b)</p><p>Known stages. I ♀♂, N</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Fig. 173 in Cruz et al. 2014); 2) dorsal portion of turbinate eyes in lateral view without constriction; 3) dorsal portion of turbinate eyes stalk height 0.21 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 5) forewing with two brown bands, apical band parallel to posterior margin (Fig. 176 in Cruz et al. 2014); 6) marginal intercalary veins paired (Fig. 176 in Cruz et al. 2014); 7) hind wing with small marks of pigmentation (Fig. 177 in Cruz et al. 2014); 8) costal process of hind wing pointed (Fig. 177 in Cruz et al. 2014); 9) marginal intercalary veins on hind wing present (Fig. 177 in Cruz et al. 2014); 10) abdominal sterna with pair of medioanterior and medioposterior sigilla strongly pigmented; 11) abdominal sterna washed with red spots and with one anterolateral mark; 12) forceps segment I wide at base (Fig. 175 in Cruz et al. 2014); 13) forceps segment III elongate (Fig. 175 in Cruz et al. 2014).</p><p>Female imago: 1) forewing with two bands, apical band parallel to posterior margin (Fig. 179 in Cruz et al. 2014); 2) marginal intercalary veins paired (Fig. 179 in Cruz et al. 2014); 3) hind wing with small marks of pigmentation (Fig. 180 in Cruz et al. 2014); 4) costal process of hind wing pointed (Fig. 180 in Cruz et al. 2014); 5) marginal intercalary veins on hind wing present (Fig. 180 in Cruz et al. 2014); 6) abdomen covered by red spots (Figs. 178 and 181 in Cruz et al. 2014); 7) abdominal sterna with pair of medioanterior and medioposterior sigilla strongly pigmented (Fig. 181 in Cruz et al. 2014).</p><p>Mature nymph: 1) distal margin of labrum medially with simple spine-setae (Fig. 61B in Nieto 2008); 2) maxillary palp 1.5 × the length of galea-lacinia (Fig. 65 in Nieto 2008); 3) crown of galea-lacinia with two rows of setae; 4) paraglossa with rounded apex (Fig. 66A in Nieto 2008); 5) paraglossa with one tuft of fine and simple setae on apex; 6) metanotum with spines; 7) foretarsus anterior surface with spine-like setae (Fig. 67 in Nieto 2008); 8) hind claw denticles smaller than foreclaw denticles (Fig. 69 in Nieto 2008).</p><p>Comments. Until now, C. gloriosus is considered to be a junior synonym of Callibaetis (A.) fasciatus . After studying the type of C. gloriosus, we observed that the most conspicuous diagnostic character of C. (A.) sellacki is present in C. gloriosus: an apical band on the female forewing, parallel to the posterior margin. Thereby, C. gloriosus is proposed here as junior synonym of C. (A.) sellacki . The type specimens of C. gloriosus possess damaged forewings (Fig. 16 B), but the apical band is visible and comparable with those presented by Cruz et al. (2014: Figs. 27D, 27G).</p><p>We present figures of the dorsal and ventral habitus of the nymphs of C. (A.) sellacki (Figs. 15 A–15B) and the type of C. gloriosus (Figs. 16 A–16B).</p><p>Material examined. Callibaetis gloriosus, female subimago (type), CHILE, Marga-Marga, Ian 1919, MZB ; Callibaetis lineatus, female imago (type), CHILE, Marga Marga, iii.1931, MZB ; Callibaetis trifasciatus, male imago (type), ARGENTINA, Prov. Buenos Aires, C. Bruch coll., MZB ; two nymphs (one mounted on slides), ARGENTINA, Buenos Aires, Tandil, A ◦ Quequén Chico, 6.i.1983, E. Domínguez coll., INPA ; one male subimago, BRAZIL, Rio Grande do Sul state, Pelotas municipality, 10.xi.1959, Biezanko, M. 1960-3, NHM .</p><p>Distribution. Argentina: Bueno Aires. Chile: Marga-Marga. Brazil: Rio Grande do Sul; São Paulo.</p></div>	https://treatment.plazi.org/id/039E3278FFB6FFFBDDDA27060C47922A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFB4FFF5DDDA26450D6A91E5.text	039E3278FFB4FFF5DDDA26450D6A91E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis stictogaster Navas 1915	<div><p>Callibaetis stictogaster Navás 1915 revalidated species</p><p>(Figs. 17 A–18E)</p><p>Callibaetis stictogaster Navás 1915b: 121; Gillies 1990: 28. (previously syn. with C. jocosus by Gillies 1990)</p><p>Known stages. I ♀♂.</p><p>Diagnosis. Male imago: 1) costal margin of forewing with pigmentation extending beyond R1 and less intense after R2 (Fig. 17 D); 2) marginal intercalary veins paired (Fig. 17 D); 3) hind wing with pointed costal process; 4) marginal intercalary veins present on hind wing.</p><p>Female imago: 1) forewing completely brown (Figs. 17 F, 18C, 18D); 2) marginal intercalary veins paired (Figs. 17 F, 18C, 18D); 3) hind wing with brown marks (Fig. 18 E); 4) hind wing with pointed costal process (Fig. 18 E); 5) marginal intercalary veins on hind wing present (Fig. 18 E); 6) abdominal terga and sterna covered with brown spots (Figs. 17 B and 18A); 7) abdominal terga with longitudinal mark medially, near anterior margin; 8) anterior margin of terga II–IX with one pale triangle, inside of it with spots (Fig. 18 A); 8) abdominal sterna with pigmented medioanterior and medioposterior sigilla (Fig. 18 B); 9) abdominal sterna with two large brown marks on anterior margin, one medially and other anterolaterally (Fig. 18 B).</p><p>Redescription. Female imago: Head. Yellowish with red “M” mark (Fig. 18 A). Thorax. Anteronotal protuberance rounded. Metascutellar protuberance pointed. Wings. Forewing completely brown, with hyaline cross veins (Figs. 18 C and 18D); stigmatic area with 12 cross veins touching Sc vein; marginal intercalaries paired; length of each intercalary vein 0.9 × distance between adjacent longitudinal veins; length of forewing about 2.8× width. Hind wing (Fig. 18 E) with some brown marks; with two intercalary marginal veins; costal process pointed. Abdomen. Terga (Fig. 18 A) with longitudinal mark medially, near anterior margin, and small light brown spots; anterior margins of terga II-IX each with one white triangular mark, inside with spots. Sterna (Fig. 18 B) covered with reddish spots; with one longitudinal line medially; two large brown marks medially and anterolaterally, medial mark formed by spots. Caudal filaments light brown, segments brown at base and apex.</p><p>Comments. The female imago of C. stictogaster revalidated species can be distinguished from C. (C.) jocosus, C. (C.) zonalis, C. (C.) willineri, C. (C.) guttatus and C. (A.) dominguezi, as well other species of the genus, by the following characteristics: forewing completely brown (Figs. 17 F, 18C, 18D); marginal intercalary veins on hind wing present (Fig. 18 E); abdominal terga and sterna covered with brown spots (Figs. 17 B and 18A); anterior margin of terga II–IX with one white triangular mark, inside of it with brown spots (Fig. 18 A); and abdominal sterna with two large brown marks on anterior margin, one medially and other anterolaterally (Fig. 18 B).</p><p>Based on the re-evaluation of the morphological evidence, we propose the revalidation of Callibaetis stictogaster revalidated species and designate the female imago housed in MZLP as lectotype (Figs. 17 A and 17B), the male imago housed in MZLP as paralectotype (Figs. 17 C and 17D) and the female imago housed in MZB as paralectotype (Figs. 17 E and 17F).</p><p>Females collected from Brazil possess egg–nymph development inside their abdomens, indicating a possible ovoviviparous mode of reproduction.</p><p>Material examined. Callibaetis stictogaster, one female (lectotype by present designation), ARGENTINA, Prov. de Buenos Aires, 15.x.1913, C. Bruch coll., MZLP . Callibaetis stictogaster, one male imago (paralectotype by present designation), ARGENTINA, Prov. de Buenos Aires, 15.x.1913, C. Bruch coll., MZLP . Callibaetis stictogaster, one female imago (paralectotype by present designation), ARGENTINA, Prov. de Buenos Aires, 15.x.1913, C. Bruch coll., MZB . Four female imagoes, BRAZIL, Rio Grande do Sul state, Bossoroca, Barra do Angico, Piratini River, 112m alt., 28°032’06.2” S / 54°057’29.9” W, light trap, 20.x.2013 - 03.xi.2013, A. M. O. Pes coll., INPA.</p><p>Distribution. Argentina: Prov. Buenos Aires, Brazil: Rio Grande do Sul state.</p></div>	https://treatment.plazi.org/id/039E3278FFB4FFF5DDDA26450D6A91E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFBAFFF7DDDA23A40BFA91EA.text	039E3278FFBAFFF7DDDA23A40BFA91EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis viviparus Needham & Murphy 1924	<div><p>Callibaetis viviparus Needham &amp; Murphy 1924</p><p>(Figs. 19 A–19C)</p><p>Callibaetis viviparus Needham and Murphy 1924: 50; Domínguez et al. 2006, p. 117; Cruz et al. 2014: 62; Vinasco- Mondragón &amp; Zúñiga 2016: 93.</p><p>Known stages. I ♀</p><p>Diagnosis. Female imago: 1) forewing with pigmented C, Sc and R1 areas, extending beyond R1 but not forming bands (Fig. 19 C); 2) forewing with thick cross veins in C and Sc areas (Fig. 19 C); 3) marginal intercalary veins single (Fig. 19 C); 4) hind wing with one intercalary marginal vein (Fig. 19 C); 5) hind wing with quadrangular costal process; 6) body covered with brown spots (Fig. 19 B); 7) anterolateral spots on abdominal terga and sterna present (Fig. 19 B).</p><p>Complementary description. Female imago: Thorax. Covered with brownish spots. Wings. Forewing (Fig. 19 C) pigmentation extending beyond R1 but not forming bands; veins light brown; C, Sc and R1 areas with thick cross veins; stigmatic area with 2–3 cross veins touching and 3–4 not touching subcostal vein; marginal intercalary veins single; length of each intercalary vein 0.41 × distance between adjacent longitudinal veins; length of forewing about 2.4 × width. Hind wing hyaline (Fig. 19 C) and with 14 cross veins; one intercalary marginal vein present. Abdomen. Terga covered with brown spots, with spot anterolaterally (Fig. 19 B). Sterna with spot anterolaterally (Fig. 19 B). Caudal filaments lost.</p><p>Comments. Vinasco-Mondragón &amp; Zúñiga (2016) recorded C. viviparus and C. (C.) radiatus in Colombia based on female imagoes. The morphological evidence to differentiate both species was the shape of the costal process of the hind wing (round in C. viviparus and truncante or quadrangular in C. (C.) radiatus). Studying the holotype of C. viviparus (mounted on a slide), we observed that one peak of the costal process is turned inward, appearing to be rounded. Thus, the morphological evidence presented by Vinasco-Mondragón &amp; Zúñiga (2016) and Cruz et al. (2014) is not corroborated, not allowing the differentiation of both species. Taking into account the new evidence, association between stages should be made in order to confirm or refute the records from Colombia.</p><p>Material examined. Callibaetis viviparus, female imago (holotype, slide), Brazil, Mato Grosso state, Corumbá municipality, 14.xii.1919, Cornell University n° 643.</p><p>Distribution. Brazil: Mato Grosso do Sul.</p></div>	https://treatment.plazi.org/id/039E3278FFBAFFF7DDDA23A40BFA91EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFB8FFF7DDDA247A0CD296FF.text	039E3278FFB8FFF7DDDA247A0CD296FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Callibaetis) willineri Navas 1933	<div><p>Callibaetis (Callibaetis) willineri Navás 1933</p><p>(Figs. 20 A– 21I)</p><p>Callibaetis willineri Navás 1933: 115; Gillies 1990: 30; Domínguez et al. 2006: 117; Nieto 2008: 240; Cruz et al. 2014: 63. Callibaetis alegre Traver 1944: 45 . (syn. by Gillies 1990)</p><p>Known stages. I ♀♂, N</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Fig. 20 H); 2) dorsal portion of turbinate eyes in lateral view without constriction; 3) dorsal portion of turbinate eyes stalk height 0.75 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 5) forewing hyaline (Fig. 21 G); 6) marginal intercalary veins paired (Fig. 21 G); 7) hind wing hyaline (Fig. 21 H); 8) costal process of hind wing rounded; 9) marginal intercalary veins on hind wing absent (Fig. 21 H); 10) abdominal terga VII – X darker (Fig. 20 H); 11) abdominal sterna covered with spots, all sterna with one medial mark on anterior margin (Fig. 20 G); 12) abdominal sterna with pair of medioanterior and medioposterior sigilla weak pigmented (Fig. 20 G); 13) forceps segment I wide at base (Fig. 21 I); 14) forceps segment III oval (Fig. 21 I).</p><p>Female imago: 1) forewing with C and Sc areas pigmented overpassing R1, after R2 pigmentation with large degree of intensity (Figs. 21 A, 21C, 21E); 2) marginal intercalary veins paired (Figs. 21 A, 21C, 21E); 3) hind wing usually with one brown mark near costal process (Figs. 21 B, 21D); 4) costal process of hind wing rounded (Figs. 21 B, 21D); 5) marginal intercalary veins on hind wing absent (Figs. 21 B, 21D, 21F); 6) abdominal terga I –VIII with anterolateral spot, terga II – IX with medial longitudinal mark, terga III, V and VII laterally with inverted V mark (Figs. 20 A, 20C, 20E); 7) abdominal sterna covered with spots, with one medial mark on anterior margin (Figs. 20 B, 20D, 20F); 8) abdominal terga with medial longitudinal mark (Figs. 20 B, 20D, 20F).</p><p>Mature nymph: 1) maxillary palp shorter than apex of galea-lacinia (Fig. 80 in Nieto 2008); 2) below maxillary palp insertion on outer margin without tuft of robust spine-like setae (Fig. 80 in Nieto 2008); 3) paraglossa with row of spine-like setae on ventral surface (Fig. 81 in Nieto 2008); 4) segment III of labial palp apically rounded (Fig. 81 in Nieto 2008); 5) metanotum without spines; 6) foretarsus anterior surface without spine-like setae (Fig. 82 in Nieto 2008); 7) hind claw denticles smaller than foreclaw denticles (Fig. 84 in Nieto 2008).</p><p>Comments. The nymphs of C. (C.) willineri and C. (C.) zonalis are similar, but they can be differentiated by the length of the maxillary palp, which is shorter than the galea-lacinia in C. (C.) willineri and longer in C. (C.) zonalis . Cruz et al. (2014) equivocally presented a female C. (C.) willineri labeled as C. (C.) guttatus (see more comments about similar species and its differentiation in C. (C.) guttatus and C. (C.) jocosus).</p><p>We present photographs of the female imago and male imago (Figs. 20 A–20H).</p><p>The specimens studied by Navás (1915) from San Miguel, Buenos Aires, Argentina, were not found, thus a lectotype was not designated.</p><p>Material examined. One female imago, URUGUAY, San Gregorio, Ar. Orillo Rio Uruguay, 29.xi.1959, A. Mesa y San Martin, C. S. Carbonell coll., MZB ; one nymph and one female imago, URUGUAY, Flores, Ruta 14, Km 235, Gruta del Palacio, 16.v.2009, E. Morelli and C. Molineri cols.; one female imago, ARGENTINA, La Plata city, light of living room, N. Hamada coll., INPA ; two females and one male imago, BRAZIL, Rio Grande do Sul, Pelotas, 14.vi.2011, INPA ; four female reared, BRAZIL, Santa Catarina, Iraní - Ponte Serrada (Lake next to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-53.61711&amp;materialsCitation.latitude=-26.303165" title="Search Plazi for locations around (long -53.61711/lat -26.303165)">Gas station</a>), BR 282, 26°18’11.4” S / 53°37’01.6” W, 649 m alt., 17.ix.2011, P.V. Cruz and N. Hamada cols., INPA ; one female reared, BRAZIL, lake in Valcir Rodrigues farm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-53.82139&amp;materialsCitation.latitude=-27.281221" title="Search Plazi for locations around (long -53.82139/lat -27.281221)">Derrubadas</a>, 27°16’52.4” S / 53°49’17.0” W, 29.ix. 2011, 429m, P.V. Cruz and R. Boldrini cols., INPA ; one male and female reared, BRAZIL, Rio Grande do Sul, Balneário das fontes, lake next to swimming pool, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-53.875942&amp;materialsCitation.latitude=-27.257889" title="Search Plazi for locations around (long -53.875942/lat -27.257889)">Derrubadas</a>, 27°15’28.4” S / 53°52’33.4” W, 29.ix. 2011, 421 m. alt., P.V. Cruz and R. Boldrini cols., INPA ; one female reared, BRAZIL, Paraná, PR 170,estrada de terra, Rondon farm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.37264&amp;materialsCitation.latitude=-26.358" title="Search Plazi for locations around (long -51.37264/lat -26.358)">General Carneiro Pedra</a>, 26°21’28.8” S / 51°22’21.5” W, 04.x.2011, 1059 m. alt., P.V. Cruz and R. Boldrini cols., INPA .</p><p>Distribution. Argentina: Buenos Aires. Brazil: Paraná; Santa Catarina; Rio Grande do Sul. Uruguay: Flores.</p></div>	https://treatment.plazi.org/id/039E3278FFB8FFF7DDDA247A0CD296FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
039E3278FFBEFFCCDDDA22760BF291C7.text	039E3278FFBEFFCCDDDA22760BF291C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callibaetis (Callibaetis) zonalis Navas 1915	<div><p>Callibaetis (Callibaetis) zonalis Navás 1915</p><p>(Figs. 22 A–22E)</p><p>Callibaetis zonalis Navás 1915a: 13; Gillies 1990: 31; Domínguez et al. 2006: 117; Cruz et al. 2014: 63. Callibaetis vitreus Navás 1915b: 121 (syn. by Gillies 1990)</p><p>Callibaetis vitreus Navás, 1919: 81 (syn. with C. vitreus Navás 1915 by Hubbard &amp; Edmunds 1977; note that this is a hom. syn.)</p><p>Baetis opacus Navás 1915a: 12 . (syn. by Gillies 1990)</p><p>Callibaetis sobrius Navás 1916: 61 . (syn. by Gillies 1990)</p><p>Baetis virellus Navás 1915b: 119 . (syn. by Gillies 1990)</p><p>Callibaetis apertus Navás 1917: 190 . (syn. by Gillies 1990)</p><p>Callibaetis depressus Navás 1922: 59 . (syn. by Gillies 1990)</p><p>Callibaetis amoenus Navás 1930c: 131 . (syn. by Gillies 1990)</p><p>Callibaetis fluminensis Cruz, Salles &amp; Hamada 2009 syn. nov.</p><p>Known stages. I ♀♂, N</p><p>Diagnosis. Male imago: 1) dorsal portion of turbinate eyes oval (Fig. 1 in Cruz et al. 2009); 2) dorsal portion of turbinate eyes in lateral view with constriction; 3) dorsal portion of turbinate eyes stalk height 2.6 × height of dorsal portion; 4) turbinate portion of compound eyes (in lateral view) with anterior and posterior margins divergent; 5) forewing with brownish stigmatic area and small brownish spots along costal vein (Fig. 3 in Cruz et al. 2009); 6) marginal intercalary veins paired (Fig. 3 in Cruz et al. 2009); 7) hind wing hyaline (Fig. 4 A in Cruz et al. 2009); 8) costal process of hind wing rounded (Fig. 4 B in Cruz et al. 2009); 9) marginal intercalary veins on hind wing absent (Fig. 4 B in Cruz et al. 2009); 10) abdominal terga III, V and VII laterally with inverted V mark; 11) abdominal sterna covered with spots and with pair of medioanterior and medioposterior sigilla weak pigmented; 12) forceps segment I wide at base (Fig. 5 in Cruz et al. 2009); 13) forceps segment III oval (Fig. 5 in Cruz et al. 2009).</p><p>Female imago: 1) forewing with C and Sc areas pigmented overlapping R1, after R2 with lighter pigmentation (Fig. 7 in Cruz et al. 2009); 2) marginal intercalary veins paired (Fig. 7 in Cruz et al. 2009); 3) hind wing with brown mark near costar process (Fig. 8 A in Cruz et al. 2009); 4) costal process of hind wing rounded (Fig. 8 B in Cruz et al. 2009); 5) marginal intercalary veins on hind wing absent (Fig. 8 B in Cruz et al. 2009); 6) abdominal terga III, V and VII laterally with inverted V mark (Fig. 6 in Cruz et al. 2009); 7) abdominal sterna covered by spots (Fig. 9 in Cruz et al. 2009); 8) abdominal terga with medial longitudinal mark (Figs. 22 B–22D); 9) abdominal sterna medially, on anterior margin, without one large spot.</p><p>Mature nymph: 1) maxillary palp 1.3 × the length of galea-lacinia (Fig. 15 in Cruz et al. 2009); 2) below maxillary palp insertion on outer margin with tuft of robust spine-like setae (Fig. 15 in Cruz et al. 2009); 3) paraglossa with row of spine-like setae on ventral surface (Fig. 16 A in Cruz et al. 2009); 4) segment III of labial apically rounded (Fig. 16 A in Cruz et al. 2009); 5) metanotum without spines; 6) foretarsus anterior surface without spine-like setae (Fig. 17 A in Cruz et al. 2009); 7) hind claw denticles smaller than foreclaw denticles (Fig. 18 E in Cruz et al. 2009).</p><p>Comments. After analyzing the types, it is clear that the pigmentation pattern and morphology of C. (C.) fluminensis are identical to those of C. (C.) zonalis . Both species possess forewings with C and Sc areas pigmented overlapping R1, after R2 with lighter pigmentation (Fig. 7 in Cruz et al. 2009); marginal intercalary veins paired (Fig. 7 in Cruz et al. 2009); costal processes of hind wings rounded (Fig. 8 B in Cruz et al. 2009); marginal intercalary veins on hind wings absent (Fig. 8 B in Cruz et al. 2009); abdominal terga III, V and VII laterally with inverted V marks (Figs. 22 C–22E and Fig. 6 in Cruz et al. 2009); abdominal sterna covered by spots (Figs. 22 C– 22E and Fig. 9 in Cruz et al. 2009); abdominal terga with medial longitudinal marks (Figs. 22 B–22D); 9) abdominal sterna medially, on anterior margin, without one large spot. Thereby, C. (C.) fluminensis is considered a junior synonym of C. (C.) zonalis .</p><p>The specimen studied by Navás (1915) from La Plata, Argentina, is damaged, and we analyzed it only through photographs. Further studies are necessary to determinate if the specimen should be designated as lectotype or if a neotype should be designated.</p><p>Material examined. Callibaetis apertus, female subimago (type), ARGENTINA, Prov. Buenos Aires, 4.x.1915, C. Bruch coll., MZB ; Callibaetis vitreus, male subimago (type) ARGENTINA, La Plata, 12.iv.1915, MZB ; Callibaetis depressus, female subimago (type), ARGENTINA, Santiago del Esterno, 1920, MZB ; Callibaetis zonalis, photograph of female imago (type), ARGENTINA, La Plata, iii.1913, C. Bruch coll., MZLP ; one female imago, BRAZIL, São Paulo, i.1926, MZB; one female imago, ARGENTINA, Buenos Aires, 6.x.1915, C. Bruch coll., MZB; Callibaetis fluminensis, female imago (holotype), one nymph and one male imago (paratypes), BRAZIL, Rio de Janeiro state, Nova Friburgo Municipality, Lumiar, 22°23’ 27.2” S / 42° 20’ 03.6” W, 3rd order tributary of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.33433&amp;materialsCitation.latitude=-22.390888" title="Search Plazi for locations around (long -42.33433/lat -22.390888)">Rio Bonito</a>, pool, v.2008, M.R. Souza coll., INPA .</p><p>Distribution. Argentina: Buenos Aires; La Plata; Santiago del Esterno. Brazil: São Paulo; Rio de Janeiro. Paraguay: Rio Paraguay. Uruguay: Maldonado.</p></div>	https://treatment.plazi.org/id/039E3278FFBEFFCCDDDA22760BF291C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cruz, Paulo Vilela;Salles, Frederico Falcão;Hamada, Neusa	Cruz, Paulo Vilela, Salles, Frederico Falcão, Hamada, Neusa (2017): Additions and corrections to the systematics of mayfly species assigned to the genus Callibaetis Eaton 1881 (Ephemeroptera: Baetidae) from South America. Zootaxa 4231 (4): 500-534, DOI: 10.11646/zootaxa.4231.4.2
