taxonID	type	description	language	source
039E87DAFFD2A243E0C2FE170B05FBB4.taxon	materials_examined	Type species: Limnophila montana Osten Sacken, 1860 (monotypic).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A243E0C2FE170B05FBB4.taxon	diagnosis	Diagnosis. Head with eye dichoptic. Antenna 16 - segmented. Palpus 5 - segmented. Thorax with mesonotum bearing no pits. Wing venation: Sc ending near level of fork of Rs; crossvein sc-r situated 1 – 3 times length of itself proximal to tip of Sc; Rs branching into R 2, R 3, R 4, and R 5, origin of Rs situated near level of middle of M; R 2 + 3 + 4 short, 1 – 4 times length of crossvein r-m; R 2 + 3 strongly curved near fork of R 2 + 3 + 4; R 2 erect, 1 – 4 times length of apical section of R 1; M 1 + 2 forked, 0.5 – 2 times as long as cell m 1; cell dm present; crossvein m-cu situated at base of cell dm or very close to it. Legs with tibial spurs 1 + 2 + 2. Hypopygium not rotated; epandrium and hypandrium fused into continuous ring; proctiger large, lateral margin narrowly sclerotized, posterolateral corner with small lobe, pair of bacilliform sclerites present between these lobes; two pairs of gonostyli present, outer gonostylus fleshy; interbase with medial part fused. Female terminalia with tergites 8 to 10 distinctly separated; cercus broad; hypogynial valve stout, pointed at tip, with swelling at lateral side on basal part, each valve not distinctly separated except on distal part; genital frame roughly spoon-shaped with lateral arm oval, bearing small tubercle at center; two spermathecal ducts present, spermathecae indistinct.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	description	(Figs 1, 2, 6 A)	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	materials_examined	Type locality: Japan, Honshu, Gifu.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	materials_examined	Type material examined. HOLOTYPE, male, JAPAN, Honshu, Gifu, 20. IV. 1921, K. Takeuchi (USNM). Other examined material. JAPAN: [Honshu] 1 female, Aomori, Nishimeya-mura, Kawaratai, Ôkawa Path, altitude 130 m, N 40.5006 °, E 140.2040 °, 14. V. 2013, D. Kato (BLKU); 1 male, same data as previous except 7. V. 2014; 1 male, same data as previous except 13. V. 2014; 2 males, Aomori, Hirosaki-shi, Sôma Path, altitude 390 m, N 40.4947 °, E 140.4023 °, 24. V. 2013, D. Kato (BLKU); 1 male, Aomori, Fukaura-machi, Mt. Takanio-yama, altitude 140 m, N 40.6899 °, E 140.1028 °, 2. V. 2014, D. Kato (BLKU); 1 female, Niigata, Nagaoka-shi, Karuizawa, N 37.4417 °, E 138. 9342 °, 9. IV. 2016, G. Mashima (BLKU); 1 male, Niigata, Shibata-shi, Takidani, altitude 204 m, N 37.8155 °, E 139.4629 °, 11. V. 2022, D. Kato (BLKU); 2 males, 3 females, Niigata, Tokamachi-shi, Matsunoyama, Echigo-Matsunoyama Museum of Natural Science, ‘ Kyororo’, altitude 310 m, N 37.0995 °, E 138.6163 °, 4. IV. 2020, D. Kato (BLKU); 1 male, same data as previous except 9. V. 2020; 35 males, 19 females, same data as previous except 9 – 16. V. 2021, MT; 19 males, 37 females, same data as previous except 16 – 23. V. 2021, MT; 4 males, 5 females, same data as previous except 23 – 30. V. 2021, MT; 1 male, 2 females, Toyama, Nanto-shi, Tatenoharahigashi, Johana Service Area, altitude 215 m, N 36.5033 °, E 136.8729 °, 13. IV. 2014, D. Kato (BLKU); 1 male, Omi (Shiga), Mt. Ibuki, N 35.417 °, E 136.400 °, 13. IV. 1958, T. Mishima (USNM); 4 females, Ôsaka, Izumisano-shi, Inunakisan Spa, altitude 250 m, N 34.3398 °, E 135.3837 °, 16. IV. 2014, D. Kato (BLKU); 1 female, Tottori, Kurayoshi-shi, Sekiganechô-Nozoe, Mt. Karasuga-sen, altitude 1000 m, N 35.3535 °, E 133.5857 °, 17. V. 2015, D. Kato (BLKU); 1 female, Okayama, Maniwa-shi, Hiruzen-Shimotokuyama, altitude 780 m, N 35.3293 °, E 133.5972 °, 1. V. 2016, D. Kato (BLKU); 1 male, Yamaguchi, Iwakuni-shi, Nishiki-machi-Ôno, Mt. Shôjô-yama, altitude 330 m, N 34.2999 °, E 131.9375 °, 2. V. 2016, D. Kato (BLKU). [Shikoku] 1 male, Ehime, Matsuyama, Sugitate, N 33.861 °, E 132.848 °, 18. IV. 1953, T. Mohri (USNM); 1 male, Ehime, Omogokei, 800 m, N 33.72 °, E 133.09 °, 10. V. 1952, Ishihara (USNM); 2 males, Ehime, Tôon-shi, Kamihayashi, near Shiraito Waterfall, altitude 560 m, N 33.7370 °, E 132.8969 °, 20. IV. 2014, D. Kato (BLKU); 1 female, Ehime, Tôon-shi, Kawanouchi, altitude 440 m, N 33.766 °, E 132.970 °, 20. IV. 2014, D. Kato (BLKU); 2 males, Ehime, Tôon-shi, Yamanouchi, Akako-keikoku Valley, N 33.876 °, E 132.945 °, 27. IV. 2018, R. Toda (BLKU); 4 males, Tokushima, Higashimiyoshi-chô, Higashiyama, Ogawadani-gawa River, altitude 340 m, N 34.0883 °, E 133.8799 °, 21. IV. 2014, D. Kato (BLKU); 2 males, Tokushima, Higashimiyoshi-chô, Nishisyô, Kamodani-gawa River, altitude 170 m, N 34.0153 °, E 133.9432 °, 21. IV. 2014, D. Kato (BLKU); 2 males, Tokushima, Miyoshi-shi, Higashiiya-Sugeoi, near Nagoro Dam, altitude 920 m, N 33.8518 °, E 134.0234 °, 29. IV. 2016, D. Kato (BLKU); 1 male, Miuma-shi, Koyadaira, altitude 1340 m, N 33.8754 °, E 134.0957 °, 30. IV. 2016, D. Kato (BLKU); 2 males, Kagawa, Mannô-chô, Myôjin-gawa River, altitude 480 m, N 34.0940 °, E 134.0143 °, 21. IV. 2014, D. Kato (BLKU). [Kyushu] 1 male, Fukuoka, Kitakyushu-shi, Kokuraminami-ku, Dôbaru, Takinoguchi-gawa River, N 33.790 °, E 130.817 °, 17. IV. 2019, R. Toda (BLKU); 3 males, Fukuoka, Miyako-machi, Saigawa-Hobashira, Notôge Pass, altitude 740 m, N 33.4956 °, E 130.9615 °, 22. IV. 2016, D. Kato (BLKU); 2 males, 4 females, Saga, Saga-shi, Fuji-machi-Sekiya, Kase-gawa River near Hokuzan Dam, altitude 320 m, N 33.4332 °, E 130.2321 °, 12. IV. 2015, D. Kato (BLKU). [Nansei Islands] Yakushima Is: 4 males, 2 females, Yakushima-chô, near Shirataniunsui-kyô Valley, altitude 600 m, N 30.3839 °, E 130.5728 °, 25. IV. 2018, D. Kato (BLKU).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	description	Redescription. Male. Body length 6.0 – 10.2 mm, wing length 7.5 – 11.4 mm. Head. Grey, tinged with ocherous, sparsely covered with yellow and brown setae. Anterior part of vertex with narrowest part as wide as eye in dorsal view; vertex posterior to eye slightly shorter than eye. Antenna generally brown to brownish black, about 2.5 times length of head. Scape weakly greyish by pruinosity, 3 – 4 times as long as wide. Pedicel twice longer than wide and half as long as scape, distal part distinctly broader. Flagellomeres long oval, about 1 / 3 as wide as long on segment 1, gradually decreased in size toward apical segment, base of flagellomere 1 often narrowly yellowish; each segment with about five verticils, at most as long as corresponding segment, and covered with pale pubescence, about 1 / 4 length of corresponding segment; pubescence on apical 1 – 2 flagellomere indistinct. Rostrum slightly darker than vertex by sparser pruinosity, 3 / 4 length of scape. Palpus dark brown to black; palpomere 1 small, roundish; palpomeres 2 – 5 almost same in length, each segment three times as long as palpomere 1; palpomere 4 slightly broader than other segments. Labellum dark brown to black. Thorax. Notum subnitidous by relatively sparse pruinosity, sparsely covered with relatively long, yellow to brown setae. Pronotum grey, weakly tinged with ocherous; postpronotum sometimes yellowish or brownish laterally. Mesonotum ocherous to brown; prescutum and presutural scutum with two indistinctly darker stripes at middle, often fused with each other, extending from near anterior end to near transverse suture; postsutural scutum with interspace between scutal lobes brighter by denser pruinosity. Mediotergite dark in dorsal view. Pleuron dark grey; dorsopleural membrane brighter or yellowish; laterotergite and around base of wing sometimes brownish. Wing (Fig. 1 B) tinged with brown, prearcular region yellowish. Obscure small brown spot present at origin of Rs, crossvein sc-r, tip of R 2, and fork of M 1 + 2; spot at tips of R 2 and fork of M 1 + 2 often indistinct; cord and outer end of cell dm vaguely seamed with brown. Rs weakly curved near base; cell dm 1.8 – 2.5 times as long as wide. Legs covered with short blackish setae. Coxae and trochanters yellow to ocherous, coxae dusted with grey in various degrees, sometimes entirely grey. Femora and tibiae dark brown on each distal part, gradually turning into yellow toward base of each segment in darker specimens, or tip of each segment narrowly dark in lighter colored specimens. Tarsi dark brown. Halter yellow to whitish yellow. Abdomen. Dark brownish grey, sparsely covered with yellowish setae. Hypopygium (Figs 1 C – I). Epandrium (Fig. 1 C) cylindrical, longer than wide. Hypandrium (Fig. 1 D) widely and slightly convex posteriorly at middle of posterior margin. Proctiger (Fig. 1 C) roughly squarish, lateral lobe (Fig. 1 E) long triangular, obtuse at tip, about 1.5 times as long as wide. Gonocoxite (Fig. 1 F) 1.5 – 1.7 times as long as wide, and 0.6 – 0.7 times as long as epandrium, widest on basal 1 / 3 and narrowest at middle. Outer gonostylus (Fig. 1 G) long finger-shaped, basally widened, about as long as gonocoxite and inner gonostylus, slightly wider than inner gonostylus; inner gonostylus (Fig. 1 G) slightly sinuous, tip pointed. Interbase (Figs 1 H – I) with middle part fused and forming narrow band, bearing pair of blade-shaped ventral lobes, distal 1 / 3 slightly broader and narrowed to tip; posterior margin of interbase with indistinct U-shaped notch at middle; lateral arm narrow and almost straight. Paramere (Figs 1 H – I) slender, weakly curved outward, outer margin with serrate area near base. Aedeagus (Figs 1 H – I) cylindrical, relatively stout, distal part slightly curved ventrally, tip ending at level of posterior end of interbase. Sperm pump (Figs 1 H – I) large, slightly shorter than and about three times as wide as aedeagus, lateral margin with fin-like extension directed anteriorly. Ejaculatory apodeme (Figs 1 H – I) with each arm large and wide, directed anteriorly. Female. Body length 7.0 – 8.5 mm, wing length 8.3 – 10.2 mm. Generally resembling male. Female terminalia (Figs 2 A – G). Generally dark brown. Tergite 10 brown, yellow apically. Cercus (Fig. 2 B) yellow, about as long as tergite 10, weakly constricted near middle in lateral view, densely covered with long hair-like setae; ventral margin at distal 1 / 5 roundly produced; distal 1 / 5 gradually narrowed to tip. Sternite 8 (Fig. 2 E) squarish, slightly longer than wide in ventral view. Hypogynial valve wide blade-shaped, about 1.8 times as long as sternite 8, apical 1 / 3 strongly narrowed in ventral view; tip ending near level of middle of cercus. Sternite 9 (Figs 2 C – D) with anterior margin deeply notched at middle, posterior margin with ventrally-directed lobe at center, posterolateral part extended into tongue-shaped lobe. Genital frame (Figs 2 F – G) with genital fork slender rod-shaped, arched in dorsoventral axis, about twice as long as remainder of genital frame, anterior part directed anteroventrally; lateral arm oval, central part distinctly produced; common spermathecal duct short, if bent anteriorly, furcation point situated near base of genital fork.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu, and Nansei Islands (Yakushima Island )) (Fig. 6 A) (Oosterbroek 2023).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD2A247E0C2F8A80BABFBCF.taxon	discussion	Remarks. This species is similar to a West Palaearctic species D. (D.) transversa (Meigen, 1804) based on the general appearance and hypopygium, but is differentiated from it by the following characters: dark markings on wing obscure, distal part of CuA without dark area (Fig. 1 B) (distinct, borders of markings clearer, distal part of CuP with distinctly dark area in D. transversa); hypopygium with epandrium cylindrical, longer than wide (Fig. 1 D) (roughly trapezoid in dorsal view, shorter than wide in D. transversa); female terminalia with cercus wider on basal half (Fig. 2 B) (wider on distal half in D. transversa).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	description	(Figs 3, 6 A)	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	diagnosis	Diagnosis. Grey to dark brownish grey species with sparsely patterned wing. Antenna generally brownish black, 1.7 – 2 times length of head. Thorax generally grey to dark grey, with four darker thoracic stripes. Wing brownish tinged, yellowish on base, with small or narrow darkening at origin of Rs, R 2, along cord and CuA. Legs with femora to tarsi dark brown except yellow femoral bases. Hypopygium with epandrium cylindrical, about as long as wide. Gonocoxite 2.2 times as long as wide and about as long as epandrium. Outer gonostylus roughly finger-shaped, slightly shorter than gonocoxite and inner gonostylus. Interbase with middle part broadened into squarish plate, bearing distinct U-shaped notch at middle of posterior margin. Aedeagus cylindrical, straight, tip extending slightly beyond posterior end of interbase.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	materials_examined	Type material. HOLOTYPE, male, JAPAN, Honshu, Niigata, Tokamachi-shi, Matsunoyama, Echigo-Matsunoyama Museum of Natural Science, ‘ Kyororo’, altitude 310 m, N 37.0995 °, E 138.6163 °, 16 – 23. V. 2021, D. Kato, MT (BLKU). PARATYPES: JAPAN: [Honshu] 1 male, same data as holotype except 14. V. 2023; 1 male, Niigata, Shibata-shi, Takidani, altitude 204 m, N 37.8155 °, E 139.4629 °, 11. V. 2022, D. Kato (BLKU).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	description	Description. Male. Body length 8.1 – 9.0 mm, wing length 8.0 – 9.3 mm. Head. Grey, weakly tinged with brown on posterolateral part of vertex, sparsely covered with yellow and dark brown setae. Anterior part of vertex with narrowest part relatively broad, 4 / 3 as wide as eye in dorsal view; vertex posterior to eye slightly longer than eye. Antenna generally brownish black, 1.7 – 2 times length of head. Scape dark grey, 3.5 times as long as wide. Pedicel slightly greyish, twice longer than wide and about half as long as scape, distal part distinctly broader. Flagellomere oval to long oval, 2 – 3 times length of width on flagellomere 1, gradually decreased in size toward apical segment; each segment with about seven verticils, at most as long as corresponding segment, and covered with pale pubescence, about 1 / 4 length of corresponding segment; pubescence on apical flagellomere indistinct. Rostrum 3 / 4 length of scape. Palpus dark brown; palpomere 1 small, roundish; palpomeres 2 – 5 almost same in length, each segment three times as long as palpomere 1; palpomere 4 slightly broader than other segments. Labellum dark brown. Thorax. Notum weakly subnitidous by relatively sparse pruinosity, sparsely covered with short yellow to brown setae. Pronotum grey, with or without brownish longitudinal line at middle; postpronotum vaguely brownish laterally. Mesonotum generally grey; prescutum and presutural scutum tinged with ocherous except lateral margin, with four darker stripes, middle pair well-separated on anterior part in anterodorsal view, extending from near anterior end to near transverse suture; lateral pair broader and shorter, about half length of middle one, reaching at transverse suture and joining dark marking on scutal lobes; scutal lobe with large brown area at middle, narrow posteriorly. Scutellum weakly brownish. Mediotergite distinctly dark on posterior half in dorsal view. Pleuron dark grey, dorsopleural membrane pale yellowish. Wing (Fig. 3 B) tinged with brown, prearcular region yellowish. Obscure small darkening present at origin of Rs and R 2, one at crossvein sc-r and near fork of R 2 + 3 + 4 absent in one paratype; CuA and cord seamed with brown, one on CuA more distinct, one on A 1 and outer end of cell dm indistinctly dark. Rs strongly curved near base; cell dm 2.8 times as long as wide. Legs covered with short blackish setae. Coxae grey, posteroapical surface of fore coxa brownish. Trochanters brown to dark brown. Succeeding segments dark brown, basal 1 / 3 of femora gradually turning into yellow toward bases. Halter whitish yellow. Abdomen. Dark brownish grey, sparsely covered with yellowish setae. Hypopygium (Figs 3 C – I). Epandrium (Fig. 3 C) cylindrical, about as long as wide. Hypandrium (Fig. 3 D) strongly and widely convex posteriorly at middle of posterior margin. Proctiger (Fig. 3 C) roughly squarish, lateral lobe (Fig. 3 E) roughly tongue-shaped, weakly pointed at tip, twice as long as wide. Gonocoxite (Fig. 3 F) 2.2 times as long as wide, and 0.9 – 1 times as long as epandrium, slender, widest on basal and distal 1 / 3. Outer gonostylus (Fig. 3 G) roughly finger-shaped, gradually narrowing to tip, slightly shorter than gonocoxite and inner gonostylus, slightly wider than inner gonostylus; inner gonostylus (Fig. 3 G) curved and flattened on distal 1 / 3, tip rounded. Interbase (Figs 3 H – I) with middle part fused and forming wide, squarish plate, bearing pair of small triangular ventral lobes, gradually narrowed to tip; posterior margin of interbase with distinct U-shaped notch at middle; lateral arm curved medially and abruptly narrowed on distal 1 / 3. Paramere (Figs 3 H – I) slender, weakly curved outward. Aedeagus (Figs 3 H – I) cylindrical, relatively slender, straight, tip extending slightly beyond posterior end of interbase. Sperm pump (Figs 3 H – I) large, slightly shorter than and about five times as wide as aedeagus, with small nail-shaped plate covering middle of ventral end. Ejaculatory apodeme (Figs 3 H – I) with each arm large and wide, directing anteroventrally. Female. Unknown.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	distribution	Distribution. Japan (Honshu) (Fig. 6 A).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	etymology	Etymology. This new species is named after the type locality, Echigo-Matsunoyama Museum of Natural Science, ‘ Kyororo’, where it was found for the first time. The specific name is a noun in nominative singular.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD6A245E0C2FB6F0A3FFA83.taxon	discussion	Remarks. This species is generally somewhat similar to the previous species, D. (D.) diluta, and the morphological differences are shown in the key to species above.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD4A24EE0C2FA230D2BFED7.taxon	description	(Figs 4, 5, 6 B)	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD4A24EE0C2FA230D2BFED7.taxon	materials_examined	Type material examined. HOLOTYPE, male, JAPAN, Nansei Islands, Kagoshima, Yakushima, Kosugidani, altitude 2,500 feet, N 30.34 °, E 130.55 °, 29. IV. 1929, S. Issiki (as Dactylolabis longicauda megastylata Alexander, 1930) (USNM). PARATYPES: JAPAN: [Honshu] 1 male, Ôsaka, Mount Minomo, N 34.85 °, E 135.46 °, 4. V. 1921, K. Takeuchi (as D. longicauda longicauda Alexander, 1922) (USNM); 1 male, same data as holotype of D. l. megastylata (as D. l. megastylata). Other examined material. JAPAN: [Honshu] 1 male, Aomori, Nishimeya-mura, Kawaratai, Ôkawa-rindô Path, altitude 300 m, N 40.5006 °, E 140.2040 °, 30. V. 2014, D. Kato (BLKU); 1 male, Niigata, Tokamachi-shi, Matsunoyama, Echigo-Matsunoyama Museum of Natural Science, ‘ Kyororo’, altitude 310 m, N 37.0995 °, E 138.6163 °, 9. V. 2020, D. Kato (BLKU); 1 male, Niigata, Itoigawa-shi, Kajiyama, near Amakazari Spa, altitude 861 m, N 36.9213 °, E 137.9539 °, 14. VI. 2022, D. Kato (BLKU); 1 male, 1 female, Toyama, Toyamashi, Arimine, Inone-dani Valley, altitude 1134 m, N 36.4905 °, E 137.4656 °, 15. VI. 2022, D. Kato (BLKU); 1 male, Gifu, Sakauchi, N 35.59 °, E 136.42 °, 4. V. 1958, Mishima (USNM); 1 male, 1 female, Aichi, Seto-shi, Katakusachô, altitude 400 m, N 35.26 °, E 137.16 °, 5. V. 2012, D. Kato (BLKU); 4 males, Aichi, Toyota-shi, Kawashimo-chô, tributary of Yahagi-gawa River, altitude 140 m, N 35.2037 °, E 137.3012 °, 4. V. 2014, D. Kato (BLKU); 2 males, Yamanashi, Kôshu-shi, Enzankamihagihara, altitude 1360 m, N 35.7092 °, E 138.8274 °, 7. VII. 2014, D. Kato (BLKU); 1 female, Mie, Matsuzaka-shi, Iitaka-chô-Kuwabara, Tsukide-gawa River, altitude 320 m, N 34.425 °, E 136.163 °, 15. IV. 2014, D. Kato (BLKU); 4 males, 1 female, Mie, Matsuzaka-shi, Okawachi-chô, altitude 73 m, N 34.5197 °, E 136.4727 °, 15. V. 2014, D. Kato (BLKU); 1 male, Yamaguchi, Iwakuni-shi, Nishiki-machi-Ôno, Mt. Shôjô-yama, altitude 330 m, N 34.2999 °, E 131.9375 °, 2. V. 2016, D. Kato (BLKU). [Shikoku] 1 male, Iyo (Ehime), Omogokei, altitude 800 m, N 33.72 °, E 133.09 °, 10. V. 1952, Ishihara (USNM); 1 male, 1 female, Iyo (Ehime), Mt. Ishizuchi, 9. VI. 1950, Issiki-Ito (USNM); 1 male, same data as previous except 8. VI. 1950; 1 male, 1 female, Kagawa, Mannô-chô, Katsuura, altitude 480 m, N 34.0940 °, E 134.0143 °, 16. V. 2015, D. Kato (BLKU); 1 male, Awa (Tokushima), Mt. Tsurugi, 1400 – 1955 m, N 33.85 °, E 134.09 °, 1. VI. 1950, Issiki-Ito (USNM); 2 males, 1 female, Tokushima, Higashimiyoshi-chô, Nishisyô, Kamodani-gawa River, altitude 170 m, N 34.0153 °, E 133.9432 °, 21. IV. 2014, D. Kato (BLKU); 1 male, Tokushima, Miyoshi-shi, Higashiiya-Oshima, Oshimadani-gawa River, altitude 550 m, N 33.8870 °, E 133.8716 °, 29. IV. 2016, D. Kato (BLKU); 1 male, Ehime, Kumakôgen-chô, Hinoura, near Prefectural road 328, altitude 1490 m, N 33.5703 °, E 132.9381 °, 17. VI. 2019, D. Kato (BLKU); 1 female, Ehime, Shikokuchuôshi, Sangawa-chô, Hase-gawa River, altitude 260 m, N 33.9487 °, E 133.5055 °, 20. IV. 2014, D. Kato (BLKU); 1 male, 2 females, Ehime, Mima-shi, Waki-machi, Shimizu, Soedani-gawa River, N 34.15 °, E 134.15 °, 5. V. 2016, M. Aoyagi (BLKU); 6 males, 2 females, Ehime, Saijô-shi, Nishinokawa-Tei, Mt. Ishizuchi-san, altitude 1530 m, N 33.7547 °, E 133.1530 °, 16. VI. 2019, D. Kato (BLKU); 8 males, 2 females, Ehime, Tôon-shi, Kamihayashi, altitude 400 m, N 33.741 °, E 132.891 °, 20. IV. 2014, D. Kato (BLKU). [Kyushu] 1 male, Fukuoka, Kitakyushu-shi, Kokuraminami-ku, Dôbaru, Takinoguchi-gawa River, N 33.790 °, E 130.817 °, 17. IV. 2019, R. Toda (BLKU); 1 female, Fukuoka, Fukuoka-shi, Nishi-ku, Myôbaru, Myôbaru-gawa River, altitude 160 m, N 33.5618 °, E 130.2638 °, 26. III. 2016, D. Kato (BLKU); 1 male, same data as previous except 15. IV. 2016; 1 female, Fukuoka, Fukuoka-shi, Nishi-ku, Kuwabara, Biodiversity Conservation Zone of Kyushu Univ., altitude 38 m, N 33.5980 °, E 130.2161 °, 15. IV. 2016, T. Mishima (BLKU); 1 male, 1 female, Fukuoka, Miyako-machi, Saigawa-Hobashira, Notôge Pass, altitude 740 m, N 33.4956 °, E 130.9615 °, 22. IV. 2016, D. Kato (BLKU); 3 males, Saga, Saga-shi, Fuji-machi-Sekiya, Kase-gawa River near Hokuzan Dam, altitude 320 m, N 33.4332 °, E 130.2321 °, 12. IV. 2015, D. Kato (BLKU); 2 males, same data as previous except 23. IV. 2015; 2 males, 1 female, Saga, Karatsu-shi, Kyuragi-machi-Hirano, Mt. Sakurei-zan, altitude 860 m, N 33.3570 °, E 130.0703 °, 26. IV. 2015, D. Kato (BLKU); 1 female, Saga, Karatsu-shi, Hamatamamachi-Torisu, Tsubakiyama Pond, altitude 630 m, N 33.4041 °, E 130.1064 °, 26. IV. 2015, D. Kato (BLKU); 1 male, Kumamoto, Yatsushiro-shi, Izumi-machi-Momiki, altitude 1350 m, N 32.4812 °, E 131.0123 °, 11. V. 2016, D. Kato (BLKU); 1 male, Kumamoto, Yatsushiro-shi, Futamisuguchimachi, altitude 7 m, N 32.4106 °, E 130.5388 °, 10. IV. 2019, R. Toda (BLKU); 2 males, 1 female, Kirishima, altitude 2500 feet, 3. V. 1929, S. Issiki (USNM). [Nansei Islands] Yakushima Island: 1 male, 1 female, Onoaida, Ara-kawa River, altitude 1200 m, N 30.299 °, E 130.556 °, 4 – 7. VI. 2007, CT, Yamauchi et al. (BLKU); 1 male, 1 female, same data as previous except 7 – 28. VI. 2007, MT.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD4A24EE0C2FA230D2BFED7.taxon	description	Redescription. Male. Body length 6.0 – 9.8 mm, wing length 6.2 – 9.0 mm. Head. Grey, sparsely clothed with brown and yellow setae. Anterior part of vertex with narrowest part slightly wider than eye in dorsal view; vertex posterior to eye as long as eye. Antenna generally yellowish brown to brownish black, about 2.5 times length of head. Scape greyish by pruinosity, 3 – 4 times as long as wide. Pedicel often brownish, twice longer than wide and half as long as scape, distal part distinctly broader. Flagellomere oval, twice as long as wide on flagellomere 1, gradually decreased in size toward apical segment; each segment with about four verticils, at most as long as corresponding segment, and covered with pale pubescence, about 1 / 4 length of corresponding segment; pubescence on terminal flagellomere indistinct. Rostrum grey, 3 / 4 as long as scape. Palpus dark brown to black; palpomere 1 small, roundish; palpomeres 2 – 5 almost same in length, each segment three times as long as palpomere 1; palpomere 4 slightly broader than other segments. Labellum dark brown to black. Thorax. Generally grey, notum sparsely covered with short yellow setae. Pronotum sometimes bearing brown longitudinal line at middle of antepronotum; postpronotum sometimes yellowish or brownish laterally. Prescutum and presutural scutum often widely tinged with ocherous on center, with four brown stripes, more evident in anterodorsal view; middle pair distinct, running from anterior end of mesonotum to slightly before transverse suture, often fused at posterior end; lateral pair broader and shorter, about half length of middle one, reaching to transverse suture and joining dark marking on scutal lobes; scutal lobe with large brown area at middle, sometimes vaguely separated by narrow greyish line near middle; in paler colored specimens, dark markings on mesonotum weakened. Mediotergite weakly dark on posterior half in dorsal view. Pleuron grey, dorsopleural membrane brighter or yellowish. Wing (Fig. 4 C) tinged with yellow to brown, prearcular region indistinctly yellow in darker specimens. Stigma sometimes vaguely darker. Rs weakly curved near base; cell dm 2 – 2.5 times as long as wide. Legs covered with short black setae. Coxae and trochanters yellow to ocherous, coxae dusted with grey in various degrees, sometimes entirely grey. Remainders of legs yellow to dark brown; in darker specimens, femora to tarsi generally dark brown, basal 1 / 3 of femora gradually turning into yellow toward bases; in lighter colored specimens, femora to tarsi generally yellowish, tips of tibiae and distal parts of tarsi darkened. Halter yellow to dusky yellow, sometimes whitish. Abdomen. Dusky yellow to dark brown, often paler on sternites, sparsely covered with yellowish short setae; setae on hypopygium longer and darker. Hypopygium (Figs 4 D – J) with epandrium (Fig. 4 D) roughly trapezoid in dorsal view, wider distally, longer than wide. Hypandrium (Fig. 4 E) strongly and widely convex posteriorly at middle of posterior margin. Proctiger (Fig. 4 D) roughly squarish, lateral lobe (Fig. 4 F) slender tongue-shaped, narrower on distal 1 / 3, twice as long as wide. Gonocoxite (Fig. 4 G) 2.5 – 2.7 times as long as wide, and about 1.2 times longer than epandrium, slender and curved, basal part weakly wide. Outer gonostylus (Fig. 4 H) stout and bacilliform, 0.6 – 0.8 times as long as gonocoxite, slightly longer to shorter and much wider than inner gonostylus; inner gonostylus (Fig. 4 H) slender and strongly curved at middle, tip rounded. Interbase (Figs 4 I – J) with middle part fused and forming narrow band, bearing pair of claw-shaped ventral lobes, gradually narrow to tip and slightly arched; posterior margin of interbase without notch at middle; lateral arm wider on distal part, curved medially. Paramere (Figs 4 I – J) slender, weakly curved outward. Aedeagus (Figs 4 I – J) cylindrical, relatively slender, almost straight and bent posteroventrally on distal 1 / 3, tip ending at level of posterior end of interbase. Sperm pump (Figs 4 I – J) small, 0.4 times as long as and about three times as wide as aedeagus. Ejaculatory apodeme (Figs 4 I – J) with each arm narrow, directing anteriorly. Female. Body length 6.2 – 8.4 mm, wing length 7.2 – 8.3 mm. Generally resembling male. Female terminalia (Figs 5 A – F). Brown to yellow. Cercus (Fig. 5 B) yellow, slightly shorter than tergite 10, weakly constricted near middle of dorsal margin, basal part covered with long hair-like setae ventrally, distal part with very short setae; ventral margin at distal 1 / 6 strongly curved; distal 1 / 6 gradually narrowed to tip; posterodorsal edge produced posteriorly into small lobe. Sternite 8 (Fig. 5 D) squarish, slightly longer than wide in ventral view. Hypogynial valve wide blade-shaped, about twice as long as sternite 8, gradually narrowed to tip in ventral view; tip ending near level of middle of cercus. Sternite 9 (Fig. 5 C) narrow, nearly rectangular, about 1 / 4 times as long as wide, with posteriorly directed slender arm at posterolateral corner. Genital frame (Figs 5 E – F) with genital fork slender rod-shaped, curved in dorsoventral axis, about twice as long as remainder of genital frame, anterior part directed anterodorsally; lateral arm long oval, central part weakly produced; common spermathecal duct long, if bent anteriorly, furcation point situated at middle of genital fork.	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD4A24EE0C2FA230D2BFED7.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu, and Nansei Islands (Yakushima Island )) (Fig. 6 B) and? Russian Far East (Sakhalin) (Oosterbroek 2023).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
039E87DAFFD4A24EE0C2FA230D2BFED7.taxon	discussion	Remarks. The specimens collected in Yakushima Island were treated as the different subspecies, Dactylolabis longicauda megastylata Alexander, 1930, based on the larger outer gonostylus than that of D. l. longicauda Alexander, 1922. However, by examining additional specimens from various sites in Japan, including the site near the type locality of D. l. megastylata, the size of the outer gonostylus of the specimens in Yakushima Island has a variation and it falls within that of the specimens from the other localities in Japan. Therefore D. l. megastylata is synonymized with the nominotypical subspecies in this paper. This species resembles a Chinese species, D. (D.) gracilistylus Alexander, 1926, but is differentiated from it by the following characters: halter yellowish to whitish (dark brown in at least knob in D. gracilistylus); lateral arm of interbase slender, basal part distinctly narrower than distal part (Fig. 4 I) (stout, almost same width in whole length in D. gracilistylus); aedeagus ending at level of posterior margin of middle of interbase (Figs 4 I – J) (extending beyond posterior margin of middle of interbase in D. gracilistylus); ejaculatory apodeme almost straight and directed anteriorly to anterolaterally (Fig. 4 I); (curved and directed laterally in D. gracilistylus).	en	Kato, Daichi (2023): Taxonomic study of the genus Dactylolabis Osten Sacken, 1860 of Japan (Diptera: Limoniidae). Zootaxa 5375 (2): 227-240, DOI: 10.11646/zootaxa.5375.2.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.4/52324
