taxonID	type	description	language	source
039187D87C08FFC6FF11E0ABFBBEFDA7.taxon	diagnosis	Diagnosis. Surface strongly sculptured; pronotum with sublateral carina, three pairs of round depressions on anterior third and three large depressions on posterior third, forming a skull-like raised area medially; depressed area iridescent and raised area smooth. Elytral formula 12 d (I-II, III-IV, V-VI, VII-VIII, IX-X, XI-XII) + 1 a (aIX) + 1 sl / 5 d (III, V, VII, IX, XI) + 1 sl (XIII); striae formed by foveolate rows of punctures grouped in pairs, except for the accessory stria (aIX) and the lateral stria. Male genitalia with parameres slightly longer than phallobase process, and phallus about five times as long as paramere, with relatively wide digitiform apex.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C08FFC6FF11E0ABFBBEFDA7.taxon	description	Description: Holotype. Antennae black; head and body surface black and iridescent in dorsal view (Fig. 1 a); ventral surface dark brown (Fig. 1 b). Measurements: total length: 1.65 mm; maximum head width: 0.30 mm; pronotal length: 0.37 mm; maximum pronotal width: 0.58 mm; elytral length: 1.02 mm; maximum elytral width: 0.52 mm. Labrum subquadrate (Fig. 1 c); anterior margin straight, with some small teeth, long and fine setae on anterolateral angles; medial process projected on anterior margin; lateral margins parallel; surface with sparse punctures bearing setae. Anterior margin of clypeus (Fig. 1 d) slightly arcuate; anterolateral angles oblique; lateral margins subparallel; posterior margin arcuate with lateral angles feebly projected. Frons bearing a pair of lateral carinae from clypeal suture to the posterior margin of eyes, more conspicuous on anterior half; each carina with a depression on its medial side. Head surface rugose behind posterior margin of eyes. Antenna with nine antennomeres; last antennomere as long as antennomeres V – VII, bearing a pair of setae on ventral surface. Mandibles asymmetrical (Figs. 1 e – f); left mandible with two incisive teeth — proximal movable, distal immobile; distal tooth without denticles on inner margin; outer margin of left mandible projected on basal half; right mandible similar to left one, except for the lack of a proximal incisive tooth. Galea and lacinia fused into mala (Fig. 1 g); mala with apical cuticular tooth; maxillary palp with four articles; second article about two times as long as first one; third article 1.7 × as long as second one; apical one very small. Anterior margin of pronotum emarginate (Fig. 1 a), with anterolateral angles pronounced; lateral margin convex, sinuate and crenulate (coarsely on anterior half); greatest width on posterior third; posterior margin oblique with a medial incision; small fine sparse setae inserted on a fold of posterior margin of pronotum. Surface of pronotum strongly sculptured; anterior third with three pairs of round depressions: one near each anterolateral angle, fading posteriad; one on each side of medial line; and one between these two impressions; posterior half with three large depressions: one medial, divided by a raised medial line; and one on each side, divided by a sublateral carina, which extends from posterior to anterior margin; depressions with iridescent surface and raised parts smooth. Scutellar shield very small; rounded on anterior margin and obovate on posterior margin. Elytron (Fig. 1 a) with anterior margin sinuate with a small acute projection near humeral callus; lateral margin crenulate, converging to an acute apex; greatest width of elytra on the anterior third; elytral formula 12 d (I-II, III-IV, V-VI, VII- VIII, IX-X, XI-XII) + 1 a (aIX) + 1 sl / 5 d (III, V, VII, IX, XI) + 1 sl (XIII); complete stria formed by foveolate rows of punctures, grouped in pairs, except for the accessory stria (aIX) and the lateral stria; striation easier to observe on ventral side of elytron; sublateral carina beginning on the second fifth of the elytra, extending to the apex between striae XII and XIII; epipleuron smooth, broader on anterior third of elytron. Hind wings developed (Fig. 1 h); margin with fringe of long setae from radial hinge to the anterior part of the anal lobe; ScP and RA 1 + 2 extending to the radial hinge; RA 3 + 4 beginning near the radial hinge and extending toward the anterior margin, near a sinuosity on that margin, bordering a small pigmented area; RA 4 very weakly marked, not parallel to RA 3, fading before the margin; RP feebly marked, fading just after it begins; r 1 very short, not connected to any vein; r 4 obtuse, connecting RA 3 to RP 2 which is weakly marked and not reaching the margin; MP 1 + 2 strongly marked, angulated at medial hinge, crossing rp-mp 1 and rp-mp 2 (forming the oblong cell); medial spur slightly shorter than rp-mp 1, curved toward the posterior margin; Cu parallel to MP 1 + 2, turning toward posterior margin, ending on the AA 3; AA 3 + 4 weakly marked; AA 3 strongly marked, extending near to posterior margin; AP 1 + 2 absent; AP 3 + 4 strongly marked, bordering the posterior margin of the anal lobe which is relatively small and very narrow. Anterior margin of prosternum feebly concave and crenulate; lateral margin converging posteriad on the anterior two thirds; posterior third (prosternal process) with lateral margins divergent posteriad and posterior margin slightly sinuate and wider than anterior margin of prosternum; with deep central depression on posterior half of prosternum. Mesoventrite with anterior margin slightly emarginate, with a medial gibbosity which fits into an internal concavity on the prothorax; lateral margins subparallel; posterior margin truncate and two thirds as wide as anterior margin. Metaventrite with discrimen marked from posterior margin, almost reaching anterior margin; posterior third of metaventrite slightly raised, especially near discrimen; metaventral carina complete. Profemur with a row of fine setae on posterior face; protibia with fine and very short pubescence dorsally and with anteroventral fringe of small setae; tarsus with four tarsomeres; third tarsomere twice as long as first and second together; last tarsomere as long as the third one, with a long seta on ventral face; mesofemur with a row of fine setae on anterior face and a shallow concavity on posterior face to fit the tibia; mesotibia and tarsus similar to the first pair of legs; metacoxa with shallow concavity on posterior face to fit the metafemur; metatrochanter with a small tooth-like projection; metafemur with similar concavity to fit metatibia; metatibia and tarsus similar to the other legs. First abdominal ventrite with a conspicuous medial carina fading near the posterior margin; ventrites 2 ¯ 5 with lateral margin raised with a fine pubescence. Male genitalia (Fig. 1 i – j) with short parameres, slightly longer than phallobase process; each paramere with pair of apical setae, as long as the paramere; phallus about five times as long as paramere; lateral margins parallel on basal half, converging on apical half; apex digitiform, relatively wide. Female: unknown. Variation. Total length: 1.57 ¯ 1.65 mm; maximum head width: 0.30 ¯ 0.31 mm; pronotal length: 0.32 ¯ 0.37 mm; maximum pronotal width: 0.55 ¯ 0.58 mm; elytral length: 0.93 ¯ 1.03 mm; maximum elytral width: 0.43 ¯ 0.52 mm. Comparative notes. Ytu thanatos sp. nov. is similar to species from the Ytu zeus group because of the presence of a carina on the first ventrite and its total length greater than 1.4 mm. It is especially similar to Y. cleideae Vanin because both have 13 striae on elytron. However, Ytu thanatos sp. nov. presents a different sculpture on pronotum and elytron, the pronotum having a group of depressions that form a skull-like raised area, not present on Y. cleideae, and the elytron of Ytu thanatos sp. nov. does not have even elytral intervals keeled like on Y cleideae (Vanin 1991, fig. 1). The male genitalia of these two species are completely different, being curved and abruptly bent downward at apex and forming an acute hook in Y. cleideae (Vanin 1991, figs. 2 and 4) and weakly curved but not forming a hook in Y. thanatos.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C08FFC6FF11E0ABFBBEFDA7.taxon	etymology	Etymology. This species is named after the entity Thanatos, the personification of death in reference to the skull-like surface of the pronotum.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C08FFC6FF11E0ABFBBEFDA7.taxon	materials_examined	Type material: Holotype. Male. Brasil, Espírito Santo, Castelo, Parque Estadual do Forno Grande, Afluente do Rio Caxixe, 20 ° 31 ’ 05.6 ” S; 41 ° 05 ’ 09.0 ” W, 30. III. 2011, Col.: Jardim, G. A.; Dumas, L. L.; Nessimian, J. L. [DZRJ 5638]; paratype: 1 male — Brasil, Espírito Santo, Domingos Martins, Parque Estadual da Pedra Azul, Córrego das Piscinas, 20 ° 23 ' 51.4 " S; 41 ° 01 ' 23.9 " W, 1236 m alt. 01. IV, 2011, Col.: Jardim, G. A. [DZRJ 5639]; 1 male — Brasil, Rio de Janeiro, Petrópolis, Fazenda Inglesa, BR- 040, Cachoeira do Brazão 22 ° 27 ’ 09.81 ” S; 43 ° 12 ’ 21.6 ” W, 822 m alt., 24. X. 2010, Col.: Moraes, C.; Ferreira-Jr, N.; Sampaio, B. H. L. [DZRJ 5640].	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0DFFC3FF11E4DBFE1BFB7A.taxon	diagnosis	Diagnosis. Pronotum with sublateral carinae complete; first third of pronotum with an impression near sublateral carinae and a pair of oval impressions medially; posterior two thirds of pronotum with a heart-shaped impression near each lateral impression; medial carina on posterior two thirds. Elytral formula 12 d (I-II, III-IV, V-VI, VII- VIII, XI-XII) + 2 a (IXa, aX) + 1 sl / 5 d (III, V, VII, IX, XI) + 1 sl (XIII); striae formed by a row of foveolate punctures and grouped in pairs, except for striae IX, X and their accessory striae. Male genitalia with parameres slightly longer than phallobase; phallus widened on apical half, in dorsal view; ventral face sinuous, apex curved toward ventral side.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0DFFC3FF11E4DBFE1BFB7A.taxon	description	Description: Holotype. Antennae black; head dark and shiny dorsally; gena light brown; pronotum dark and shiny brown; hypomeron laterally light brown (Fig. 2 a); elytron dark brown medially; carinae shiny; ventrally light brown; legs light brown, except for the tibiofemoral joint and tarsi, which are dark brown (Fig. 2 b). Measurements: total length: 1.85 mm; maximum head width: 0.41 mm; pronotal length: 0.35 mm; maximum pronotal width: 0.65 mm; elytral length: 1.12 mm; maximum elytral width: 0.41 mm. Labrum subtrapezoidal (Fig. 2 c); anterior margin bearing denticles and filiform setae; medial region of anterior margin slightly emarginate; lateral margin oblique; surface with sparse punctures on anterior half, bearing setae. Anterior margin of clypeus slightly arcuate (Fig. 2 d); anterolateral angles and lateral margin oblique; posterior margin sinuate, feebly emarginate on lateral parts, medially projected. Head with conspicuous supra-orbital carinae, from clypeus to near the posterior margin of eyes; medial carina extending from the posterior margin of eyes, fading before the margin of clypeus; surface with a microrugose area behind the eyes. Antenna with nine antennomeres; last antennomere with a pair of apical setae. Mandibles asymmetrical (Figs. 2 e – f); left mandible with two incisive teeth — one immobile (distal), one movable (proximal); distal incisor without denticles on inner margin; outer margin of mandible with rounded projection; in dorsal view, depressed from distal incisor to basal third, where it presents a raised V-shaped area from the projection on outer margin to the region where the movable tooth is inserted; right mandible similar to left one, but bearing only an immobile incisor. Maxilla (Fig. 2 g) with galea and lacinia fused; with apical cuticular teeth; maxillary palp with four articles; second article three times as long as first one; third article slightly longer than second one; fourth article bearing an apical tuft of setae. Anterior margin of pronotum sinuate (Fig. 2 a); lateral margin oblique on first third and slightly convex on posterior two thirds; posterior margin sinuate, with a median incision; sublateral carinae extending from anterior to posterior margin; first third of pronotum with an impression near each sublateral carinae and a pair of oval impressions medially; posterior two thirds of pronotum with a heart-shaped impression near each lateral impression; medial carina more conspicuous on posterior two thirds; region between sublateral carinae and lateral margin depressed. Scutellar shield very small and diamond-shaped. Elytron with anterior margin bisinuate (Fig. 2 a), slightly pronounced on humeral callus; lateral margin arcuate, converging to an acute apex; elytral formula 12 d (I-II, III-IV, V-VI, VII-VIII, XI-XII) + 2 a (IXa, aX) + 1 sl / 5 d (III, V, VII, IX, XI) + 1 sl (XIII); striae formed by a row of foveolate punctures; complete striae are grouped in pairs except for striae IX and X, which are separated by the two accessory striae (IXa and aX); odd intervals carinate, except for first interval; carinae less conspicuous on second fourth of elytron, especially the carina on the 9 th and 11 th intervals, the last between accessory striae; sublateral carina strongly raised on 13 th interval, beginning on second fourth of elytron, extending to the apex; epipleuron broader on anterior third, with a constriction near the hind coxae. Hind wings developed (Fig. 2 h); margin fringed from radial hinge to anterior region of anal lobe; ScP e RA 1 + 2 extending to radial hinge; RA 3 + 4 beginning near radial hinge, extending toward anterior margin, just after a feeble sinuosity on anterior margin; RA 3 + 4 delimits a pigmented area; RA 4 feebly marked, fading without reaching the margin; RP feebly marked, extending shortly; r 1 short, not connecting to any vein; r 4 oblique; MP 1 + 2 well marked, angulated on medial hinge, crossing rp-mp 1 and rp-mp 2, forming oblong cell; Cu vein parallel to MP 1 + 2 toward posterior margin, reaching AA 3, which extends to posterior margin; AA 3 + 4 restricted to base; AP 3 + 4 well marked, following the margin of anal lobe. Anterior margin of prosternum feebly emarginate (Fig. 2 b); lateral margin subparallel, diverging on prosternal process; posterior margin of prosternal process slightly arcuate, with acute lateral angles; anterior margin of mesoventrite sinuate, with a medial gibbosity, which fits into an internal concavity of prothorax; lateral margin oblique; posterior margin truncate. Metaventrite with shallow depression at the discrimen, extending from anterior to posterior margin; metaventral carina complete, extending from anterior to posterior margin; metaventral tubercle absent. Procoxae with a pronounced trochantin; profemur with a shallow concavity on anterior face to fit the tibia, and a row of fine setae on posterior face; mesofemur with a row of fine setae on anterior face and a shallow concavity on posterior face to fit tibia; metacoxa with a shallow concavity on posterior face to fit femur; metafemur with shallow concavity on posterior face to fit metatibia; metatarsus 4 - segmented; first and second tarsomeres with some setae on ventral side; third tarsomere longer than twice the first and second together; third tarsomere with three apical setae on ventral side; the middle setae much longer than the other two; last tarsomere longer than the third (less than 1.5 × length of third tarsomere). First abdominal ventrite with a conspicuous medial carina fading near the posterior margin; ventrites 2 – 5 with lateral margin raised with a fine pubescence. Male genitalia (Figs. 2 i – j) with short parameres, slightly longer than phallobase; each paramere with a pair of apical setae, shorter than parameres; phallus widened in apical half, in dorsal view; ventral face sinuous, curved apex pointing toward ventral side. Female: unknown. Variation. Total length: 1.63 ¯ 1.85 mm; maximum head width: 0.32 ¯ 0.41 mm; pronotal length: 0.33 ¯ 0.37 mm; maximum pronotal width: 0.57 ¯ 0.65 mm; elytral length: 1.00 ¯ 1.12 mm; maximum elytral width: 0.37 ¯ 0.41 mm. Comparative notes. Ytu hypnos sp. nov. is very similar to Ytu thanatos sp. nov., differing slightly in the pronotum sculpture and conspicuously in male genitalia. Ytu hypnos sp. nov. presents the skull-like raised area on pronotum, as in Ytu thanatos sp. nov., but the impressions near sublateral carinae are more conspicuously marked in Ytu thanatos sp. nov.. The phallus of Ytu hypnos sp. nov. is sinuous in lateral view and widened on distal half, dorsal view, while on Ytu thanatos sp. nov. it is arcuate in lateral view and does not present a widened area in dorsal view.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0DFFC3FF11E4DBFE1BFB7A.taxon	etymology	Etymology. This species is named after the Greek mythological entity Hypnos, personification of sleep and twin brother of Thanatos, in reference to a similar surface of its pronotum.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0DFFC3FF11E4DBFE1BFB7A.taxon	materials_examined	Type material: Holotype. Male. Brasil, Rio de Janeiro, São Fidelis, Serra Bela Joana, Filme d’água em pedra, 22 ° 05 ’ 03.8 ” S; 41 ° 26 ’ 05.3 ” W, 25. X. 2009, Col.: Clarkson, B.; Cardoso-Costa, G.; Ferreira-Jr, N. [DZRJ 5641]; paratype: 1 male — Brasil, Rio de Janeiro, Guapimirim, Parque Nacional da Serra dos Órgãos, BR- 116, Km 92.5 (aprox.), parede com água escorrendo, 22 ° 28 ’ 16.8 ” S; 43 ° 00 ’ 04.4 ” W, 15. V. 2011, Col.: Ferreira-Jr, N.; Sampaio, B. H. L. [DZRJ 5642]; 1 male — Brasil, São Paulo, Ubatuba, BR- 101, Km 1, 300 m antes da cachoeira da escada (sentido RJ-SP), Riacho de segunda ordem, 23 ° 21 ’ 14.5 ” S; 44 ° 46 ’ 12.1 ” W, 221 m alt., 09. IX. 2011 Col.: Sampaio, B. H. L. [DZRJ 5643].	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0FFFCCFF11E15BFACDFF47.taxon	diagnosis	Diagnosis. Dorsal surface coarsely granulated and iridescent, except for an area on each elytron which combined takes a form similar to an " X ". Elytra together much broader than pronotum; elytral formula 10 d + 2 a (IVa, VIa) + 1 sl / 0 d + 0 sl; with ten complete striae and two accessory striae each: one between striae IV and V, another between striae VI and VII; intervals raised, with iridescent microscales, and row of granules. Male genitalia with parameres conspicuously longer than phallobase’s process and long phallus, with a slight constriction on basal half; digitiform apex and some regions of phallus less chitinous.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0FFFCCFF11E15BFACDFF47.taxon	description	Description: Holotype. Antennae black; dorsal surface black and shiny (Fig. 3 a); ventrally dark brown (Fig. 3 b), except for epipleuron which is light brown; legs brown, darker on distal parts. Measurements: total length: 2.22 mm; maximum head width: 0.37 mm; pronotal length: 0.52 mm; maximum pronotal width: 0.67 mm; elytral length: 1.37 mm; maximum elytral width: 0.62 mm. Labrum (Fig. 3 c) with anterior margin arcuate with denticules; anterolateral angles bearing claviform setae; medial region of anterior margin with a small process; lateral margin oblique; surface smooth and glabrous, except for the region near anterolateral angles where there are punctures bearing setae. Clypeus (Fig. 3 d) with anterior margin slightly arcuate; anterolateral angles oblique; lateral margin subparallel; posterior margin arcuate, especially on medial region; punctures visible near lateral and anterior margin. Head surface with iridescent microscales from clypeus to posterior margin; surface on a band posterior to eyes slightly less iridescent. Supraorbital carina extending from clypeal suture to posterior margin of eye; middle line of head feebly raised because of a depressed area beside each supra-orbital carina. Antenna with nine antennomeres; last antennomere as long as antennomeres IV – VIII together. Mandibles asymmetrical (Figs. 3 e – f); left mandible with two incisive teeth — one immobile (distal), one movable (proximal); distal incisor bearing two denticules on inner margin; outer margin of mandibles arcuate, without projection; right mandible similar to the left one, but with only an immobile incisor; incisor of right mandible with outer margin slightly more arcuate than on left mandible. Maxilla (Fig. 3 g) with lacinia and galea fused into mala; mala bearing apical cuticular teeth; maxillary palp with four articles; second article 1.5 × as long as first one; third article 1.5 × as long as second one; apical one very small. Anterior margin of pronotum sinuate (Fig. 3 a), slightly projected medially; anterolateral angles pronounced; lateral margin arcuate; posterior margin bisinuate, pronounced medially as much as on posterolateral angles; posterior margin with a single medial incision; pronotum broader on posterior half; surface with microscales and sparse granules larger than an ommatidium; small oval impression on each side of median line on the anterior third; medial inconspicuous depression on second third; last third of pronotum impressed near posterolateral angles. Scutellar shield black and equilateral triangle shaped. Elytra together (Fig. 3 b) much broader than pronotum; anterior margin of each elytron bisinuate; humeral callus feebly pronounced; lateral margin crenulate, subparallel on anterior half and convergent on posterior half; elytral formula 10 d + 2 a (IVa, + VIa) + 1 sl / 0 d + 0 sl; elytral intervals raised, with iridescent microscales, and with row of granules larger than those of pronotum; humeral callus with confused distribution of granules, reflecting poorly marked striae; epipleuron smooth, wide on anterior half of elytron. Hind wings developed (Fig. 3 h); margin fringed with long setae from radial hinge to anterior region anal lobe; ScP and RA 1 + 2 extending toward radial hinge; RA 3 + 4 beginning on radial hinge, extending toward anterior margin just after a feeble sinuosity on margin, limiting a pigmented area; RA 4 and RP very weakly marked; r 1 very short, not crossing any other vein; r 4 oblique; MP 1 + 2 strongly marked, angulated on medial hinge, crossing rp-mp 1 and rp-mp 2, forming oblong cell, its apex pointing toward posterior margin; Cu vein parallel to MP 1 + 2, extending toward posterior margin, reaching AA 3, which extends to posterior margin; AA 3 + 4 limited to basal area; AP 3 + 4 following posterior margin of anal lobe, which is relatively narrow. Anterior margin of prosternum (Fig. 3 b) crenulate and feebly emarginate; prosternal carina slightly oblique, constricted between coxae, diverging posteriad; posterior margin of prosternal process arcuate. Mesoventrite with a gibbosity on anterior margin, which fits an internal concavity on prothorax; lateral margin oblique; posterior margin truncate. Metaventrite with discrimen marked from anterior to posterior margin; metaventral carinae complete, extending from anterior to posterior margin; metaventral tubercles absent. Legs with fine sparse setae on femora, tibiae and tarsi; profemur robust, with a shallow impression on anterior face to fit tibia; protibia with a row of fine setae on anterior face; meso- and metafemur with shallow impression on posterior face to fit tibiae; metacoxa with similar impression to fit hind femur; tarsi four-segmented; third tarsomere twice as long as first and second together; last tarsomere twice as long as third one. Abdomen with visible plastron except for a narrow strip near ventrite margins; first ventrite with medial carina almost reaching posterior margin; posterior margin of ventrites 1 ¯ 4 microserrate. Male genitalia (Figs. 3 i – h) with parameres conspicuously longer than phallobase process; each paramere with a pair of apical setae as long as half of paramere; phallus long, with a slight constriction on basal half and digitiform apex; some regions of phallus less chitinous. Female: unknown. Variation. Total length: 2.22 ¯ 2.31 mm; maximum head width: 0.37 ¯ 0.41 mm; pronotal length: 0.40 ¯ 0.52 mm; maximum pronotal width: 0.67 ¯ 0.75 mm; elytral length: 1.37 ¯ 1.51 mm; maximum elytral width: 0.56 ¯ 0.62 mm. Comparative notes. Ytu hermes sp. nov. is very similar to Y. hephaestus Reichardt in its external morphology. They differ in the phallus which is conspicuously sinuate on the apical portion in Y. hephaestus, forming a small apical hook (Reichardt & Vanin 1977, fig. 8), while in Ytu hermes sp. nov. the phallus presents a feeble sinuosity on different portions and does not form an apical hook. Ytu hermes sp. nov. also presents an area not sclerotized which is lacking in Y. hephaestus.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0FFFCCFF11E15BFACDFF47.taxon	etymology	Etymology. This species is named after Hermes, God of commerce and the messenger of gods in Greek mythology, in reference to its similarities to Ytu zeus group (Hermes is son of Zeus).	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C0FFFCCFF11E15BFACDFF47.taxon	materials_examined	Type material: Holotype. Male. Brasil, Espírito Santo, Dores do Rio Preto, Pedra Menina, Parque Nacional do Caparaó, Cachoeira Sete Pilões, 20 ° 28 ’ 56.8 ” S; 41 ° 49 ’ 50.3 ” W, 1869 m alt., 06. I. 2013, Col.: Sampaio, B. H. L. [DZRJ 5644]; paratype: 1 male — Brasil, Minas Gerais, Alto Caparaó, Parque Nacional do Caparaó, trilha para a base Terreirão Rio José Pedro, 20 ° 24 ’ 51 ” S; 41 ° 49 ’ 34.4 ” W, 2111 m alt., 19. I. 2014, Col.: Sampaio, B. H. L. [DZRJ 5645]; 1 male — Brasil, Espírito Santo, Dores do Rio Preto, Pedra Menina, Parque Nacional do Caparaó, Afluente do Rio Preto, Laje, 20 ° 29 ’ 26.2 ” S; 41 ° 49 ’ 17.1 ” W, 1507 m alt., 24. III. 2012, Col.,: Jardim, G. A. [DZRJ 5646].	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C00FFCEFF11E507FAECFBE7.taxon	diagnosis	Diagnosis. Middle line of pronotum with a narrow longitudinal impression from posterior margin to first third; a small impression on each side of medial impression on first third, and a sublateral impression on first third. Elytron with edges flattened; elytral formula 8 d + 0 a + 1 sl / 0 d + 0 sl; nine complete stria formed by a rows of punctures; elytral intervals with iridescent microscales and feebly raised; striae forming sulci. Male genitalia with long parameres (half length of phallus); phallus narrow in lateral view; dorsal surface convex up to apical fourth, where it is slightly convex and presents small setae; feebly widened digitiform apex (in relation to phallus width), bearing long setae on dorsal face.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C00FFCEFF11E507FAECFBE7.taxon	description	Description: Holotype. Antennae black; dorsal surface shining black (Fig. 4 a); ventral surface dark brown (Fig. 4 b); legs dark brown, darker distally. Measurements: total length: 1.70 mm; maximum head width: 0.30 mm; pronotal length: 0.32 mm; maximum pronotal width: 0.48 mm; elytral length: 1.07 mm; maximum elytral width: 0.40 mm. Labrum (Fig. 4 c) slightly longer than wide; anterior margin slightly convex with denticles; anterolateral angles arcuate with claviform setae; medial region with a process; lateral margin slightly arcuate; surface smooth and glabrous on posterior third and medial region; other regions of labrum with sparse punctures bearing setae. Clypeus (Fig. 4 d) with anterior margin arcuate; anterolateral angles arcuate; lateral margin oblique; posterior margin arcuate with posterolateral angles projected. Head surface with iridescent microscales, except for a narrow band near the pronotum; supra-orbital carina extending from clypeus to posterior margin of eyes; region beside each carina depressed; head medially feebly raised. Antenna with nine antennomeres; last antennomere as long as VI – VIII combined. Mandibles asymmetrical (Figs. 4 e – f); left mandible with two incisors — one immobile (distal), one movable (proximal); distal incisor with three denticles on inner margin; outer margin strongly arcuate on medial third; right mandible similar to left one, bearing only an immobile incisor. Maxilla with galea and lacinia fused into a mala; mala with apical cuticle teeth; claviform setae on inner margin; maxillary palp four-segmented, second article twice as long as first one; third article 1.5 × longer than second one; apical one very small. Anterior margin of pronotum (Fig. 4 a) sinuate, medially protuberant; anterolateral angles strongly projected; lateral margin arcuate; greatest width at middle third; posterior margin bisinuate, medially more pronounced than at the posterolateral angles; posterior margin with single small medial incision; middle line of pronotum with a narrow shallow longitudinal impression extending from posterior margin to first third; a small round impression on each side of medial longitudinal impression on first third; sublateral round impression, smaller than the medial, on first third; posterolateral angles with shallow impression on the lateral thirds of pronotum. Anterior margin of elytron (Fig. 4 a) arcuate; lateral margin arcuate, converging to acute apex; edges of elytron explanate; elytral formula 8 d + 0 a + 1 sl / 0 d + 0 sl; striae formed by row of punctures, sublateral stria beginning in middle fifth; elytral intervals with iridescent microscales and feebly raised, so that the striae form sulci. Epipleuron broad and smooth on anterior two fifths. Hind wings reduced (Fig. 4 h); marginal fringe with very short setae, beginning just distal the radial hinge (which is weakly marked); RA 3 + 4 beginning near radial hinge, extending toward the anterior margin, just distal to a convexity on anterior margin of wing; RA 3 + 4 borders a pigmented area; RA 4 not visible; RP weakly marked, visible briefly, near basis; r 1 short, oblique, not crossing any vein; r 4 oblique; MP 1 + 2 strongly marked, angulate on medial hinge, crossing rp-mp 1 and rp-mp 2 (which is weakly marked), forming oblong cell; Cu vein almost parallel to MP 1 + 2, extending toward posterior margin, ending at AA 3, which extends to posterior margin; AA 3 + 4 present only near base; anal lobe very small and narrow. Anterior margin of prosternum arcuate and crenulate (Fig. 4 b); prosternal carinae slightly oblique; prosternal process narrower between procoxae, widening toward the posterior margin; posterior margin of prosternal process bisinuate, oblique laterally, and slightly concave medially. Anterior margin of mesoventrite feebly projected, with a medial gibbosity which fits into an internal concavity on prothorax; posterior margin truncate between the middle coxae. Metaventrite depressed medially, but without a conspicuous discrimen; metaventral carinae and tubercles absent. Metacoxae with posterior margin slightly projected over trochanter and femur and feebly serrated; femora with a fringe of long setae on anterior face; tibiae with a fringe of small setae on anterior and posterior face; tarsus with four articles; third tarsomere 1.5 × longer than first and second together; fourth tarsomere as long as twice the third one. Abdomen with visible plastron, covering the whole surface; ventrites 1 ¯ 4 with posterior margin microserrate; posterolateral angles of ventrites acutely projected, except for the last; first ventrite without medial carina. Male genitalia (Figs. 4 i – j) with long parameres, reaching half length of phallus; pair of apical setae on each paramere, one-third length of paramere. Phallus narrow in lateral view; dorsal surface convex up to apical fourth, where it presents a feeble concavity; concave area bears small setae; phallus with digitiform apex, slightly wide, bearing long setae on dorsal face. Female: unknown. Variation. Total length: 1.70 ¯ 1.91 mm; maximum head width: 0.30 ¯ 0.33 mm; pronotal length: 0.32 ¯ 0.40 mm; maximum pronotal width: 0.48 ¯ 0.58 mm; elytral length: 1.05 ¯ 1.12 mm; maximum elytral width: 0.40 ¯ 0.43 mm. Comparative notes. Ytu nyx sp. nov. is similar to species of the Ytu demeter group, because the lack of carina on ventrite 1 and the total length greater than 1.4 mm. Ytu nyx sp. nov. is especially similar to Ytu godoyi Reichardt & Vanin because it presents nine striae on elytron. These two species differ from each other by the following characteristics: Ytu nyx sp. nov. has supra-orbital carinae complete (interrupted abruptly on Y. godoyi); this new species lacks metaventral tubercles and carinae (Y. godoyi has metaventral tubercles and presents faded carinae); the male genitalia of Ytu nyx sp. nov. resembles that of the Ytu phebo group (Reichardt & Vanin 1977, figs. 13 ̄ 17).	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C00FFCEFF11E507FAECFBE7.taxon	etymology	Etymology. This species is named after the Greek mythological entity Nyx, personification of the night, in reference to its dark color and its resemblance to Y. thanatos sp. nov. and Y. hypnos sp. nov. (in Greek mythology Nyx is mother of Thanatos and Hypnos).	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C00FFCEFF11E507FAECFBE7.taxon	materials_examined	Type material: Holotype. Male. Brasil, Espírito Santo, Dores do Rio Preto, Pedra Menina, Parque Nacional do Caparaó, Rio São Domingos, Cachoeira do Aurélio, 20 ° 28 ’ 56.5 ” S; 41 ° 50 ’ 14.6 ” W, 1814 m alt., 6. I. 2013, Col.: Sampaio, B. H. L. [DZRJ 5647]; paratype: 1 male — Brasil, Espírito Santo, Dores do Rio Preto, Pedra Menina, Parque Nacional do Caparaó, Cachoeira Sete Pilões, 20 ° 28 ’ 56.8 ” S; 41 ° 49 ’ 50.3 ” W, 1869 m alt., 06. I. 2013, Col.: Sampaio, B. H. L. [DZRJ 5648]; 1 male — Brasil, São Paulo, Atibaia, 11. III. 2006, Col.: Roque, F. [DZRJ 5649].	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C02FFC8FF11E0E7FC1CF886.taxon	diagnosis	Diagnosis. Lateral margins of pronotum convex, abruptly curving inward on anterior fifth; the anterior third of pronotum with two pairs of shallow impressions, one medial and other sublateral; posterior half of pronotum with two pairs of shallow impressions, one near posterolateral angles and the other sublateral. Elytral formula 8 d + 2 a (IVa, VIa) + 1 sl / 0 d + 0 sl; elytral intervals feebly raised; surface of striae and intervals with iridescent scales. Male genitalia with long parameres and phallus curved from basis to apex in laterally; apex digitiform bearing a pair of setae on dorsally.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C02FFC8FF11E0E7FC1CF886.taxon	description	Description: Holotype. Antennae black; dorsal surface black and shiny (Fig. 5 a); ventrally dark brown (Fig. 5 b); legs darker than ventral surface. Measurements: total length: 1.76 mm; maximum head width: 0.32 mm; pronotal length: 0.33 mm; maximum pronotal width: 0.55 mm; elytral length: 1.12 mm; maximum elytral width: 0.41 mm. Labrum (Fig. 5 c) with arcuate anterior margin bearing some denticles; medial region of anterior margin with a small process; anterolateral angles bearing claviform setae; lateral margin slightly arcuate; labrum surface pubescent on anterior two thirds. Anterior margin of clypeus (Fig. 5 d) feebly arcuate; anterolateral angles oblique with a pair of setae each. Head surface with iridescent microscales; supra-orbital carina extending from clypeus to eyes, fading after reaching eyes; medial side of carinae depressed; medial part of head not raised. Antenna with nine antennomeres; last antennomere as long as V – VIII together. Mandibles asymmetrical (Figs. 5 e – f); left mandible with an immobile incisive tooth and a pair of movable incisive teeth; immobile incisor with two denticles on inner margin; outer margin of mandibles arcuate from base to apex; dorsal face with longitudinal carina, beginning on base of mandible extending toward its outer margin; right mandible similar to the left one, but bearing only an immobile incisor. Maxilla (Fig. 5 g) with galea and lacinia fused into mala; apex of mala with cuticular teeth; maxillary palp four-segmented; second article twice as long as first one; third article twice as long as second one; apical one very small. Anterior margin of pronotum arcuate (Fig. 5 a), almost truncate medially; anterolateral angle pronounced; lateral margin arcuate, oblique on anterior fifth; posterior margin bisinuate, with arcuate part medially; anterior third of pronotum with two pairs of shallow impressions, one medial and other sublateral; posterior half with two pairs of shallow impressions, one near posterolateral angles other sublateral. Scutellar shield very small, subtriangular, slightly longer than wide; anterior margin arcuate, lateral margin converging to acute apex. Anterior margin of elytron arcuate and flattened (Fig. 5 a); lateral margin subparallel on anterior two thirds, converging on posterior third; elytral formula 8 d + 2 a (IVa, VIa) + 1 sl / 0 d + 0 sl; striae formed by a row of punctures; accessory striae long; sublateral carina on posterior three-fifths; elytral intervals feebly raised; surface of striae and intervals with iridescent microscales. Epipleuron smooth and wide to level of hind coxa. Hind wing developed (Fig. 5 h); anterior margin fringed with long setae from radial hinge to anterior region of anal lobe; ScP and RA 1 + 2 extending to radial hinge; RA 3 + 4 beginning on radial hinge, extending toward anterior margin on a feeble sinuosity; RA 3 + 4 borders a pigmented area; RA 4 weakly marked, fading before reaching wing margin; RP weakly marked, visible only near the basis; r 1 short, not crossing any vein; r 4 oblique; MP 1 + 2 well marked, angulate on medial hinge, crossing rp-mp 1 and rp-mp 2, forming oblong cell; cubital vein parallel to MP 1 + 2, extending toward posterior margin of wing, reaching AA 3, which extends to posterior margin; AP 3 + 4 well marked following the margin of anal lobe, which is small and narrow. Anterior margin of prosternum arcuate and crenulate (Fig. 5 b); prosternal carinae parallel from anterior margin to the procoxae; posterior margin of prosternal process wide and arcuate. Anterior margin of mesoventrite with three gibbosities; medial gibbosity larger than the others, fitting in an internal concavity on prosternum; anterior half of mesoventrite with a transverse fusiform depression which fits the posterior margin of prosternal process. Metaventral carinae complete; metaventral tubercles absent. Anterior face of profemur with a shallow depression on distal two thirds, to fit the fore tibia; protibia with a row of setae on anterior face; meso- and metafemora with shallow depression to fit meso- and metatibiae, respectively; tarsi foursegmented; third tarsomere twice as long as tarsomeres I – II together; fourth tarsomere twice as long as the third one. Abdomen with ventrites iridescent; carina present on first ventrite, extending from anterior to posterior margin; last ventrite with a pair of weak longitudinal carinae. Male genitalia (Figs. 5 i – j) with long parameres, not reaching half of phallus length; each paramere with a pair of apical setae, as long as half of paramere length; phallus curved from base to apex in lateral view; apex of phallus digitiform with a pair of setae on dorsal region. Female: unknown. Variation. Total length: 1.76 ¯ 1.83 mm; maximum head width: 0.32 ¯ 0.33 mm; pronotal length: 0.31 ¯ 0.33 mm; maximum pronotal width: 0.55 ¯ 0.60 mm; elytral length: 1.12 ¯ 1.25 mm; maximum elytral width: 0.40 ¯ 0.41 mm. Comparative notes. Ytu coeus sp. nov. is similar to species of the Ytu zeus group, because of the presence of a carina on ventrite 1 and its length greater than 1.4 mm. It is especially similar to Ytu zeus Reichardt because both species present nine elytral striae (8 p + 1 sl). Ytu coeus sp. nov. lacks prosternal and metaventral setae (“ metasternal hairs ” in Reichardt & Vanin 1977), as in Ytu zeus. The male genitalia of Ytu coeus sp. nov. resemble those of the Ytu phebo group (Reichardt & Vanin 1977, figs. 13 ̄ 17).	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C02FFC8FF11E0E7FC1CF886.taxon	etymology	Etymology. This species is named after Coeus, the Greek Titan God of intellect, in reference to its similarity to Y. phebo group, especially on genitalia characteristics; Coeus was a brother and consort of Phoebe.	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
039187D87C02FFC8FF11E0E7FC1CF886.taxon	materials_examined	Type material: Holotype. Male. Brasil, Espírito Santo, Dores do Rio Preto, Pedra Menina, Parque Nacional do Caparaó, abrigo Casa Queimada, riacho de laje, nascente do Rio São Domingos, 20 ° 27 ’ 21.6 ” S; 41 ° 48 ’ 30.8 ” W, 2211 m alt., 05. I. 2013, Col.: Sampaio, B. H. L. [DZRJ 5650]; paratype: 1 male — same data as holotype, Cachoeira Sete Pilões, 20 ° 28 ’ 56.8 ” S; 41 ° 49 ’ 50.3 ” W, 1869 m alt., 06. I. 2013, Col.: Sampaio, B. H. L. [DZRJ 5661]; 1 male — Brasil, Ceará, Ubajara, Parque Nacional de Ubajara, trilha Araticum, Rio da Minas, 3 ° 50 ’ 3 ” S; 40 ° 54 ’ 18 ” W, 524 m alt., 19. III. 2013, Col.: Santos, A. P. M.; Takiya, D. M. [DZRJ 5651].	en	Sampaio, Brunno Henrique Lanzellotti, Ferreira-Jr, Nelson (2018): Five new species of Ytu Reichardt (Coleoptera: Myxophaga: Torridincolidae) and new records from Brazil. Zootaxa 4402 (3): 508-524, DOI: 10.11646/zootaxa.4402.3.5
