identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039187D8FFC4BB13FF188075FF61FA6C.text	039187D8FFC4BB13FF188075FF61FA6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia armipes (Fairmaire 1896) Gompel 2020	<div><p>Zarcosia armipes (Fairmaire, 1896), comb. nov.</p><p>(Figures 1; 2l,m)</p><p>Xyophilus armipes Fairmaire, 1896: 44; Syntype: BMNH (female)</p><p>Zarcosia armipes (Fairmaire, 1896): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Karnataka, Belgaum .</p><p>Distribution. West India (Figure 1).</p><p>Material examined. India. Mysore State: T. V. C. (1 #m, BMNH); Karnataka: Belgaum (1 #f syntype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC4BB13FF188075FF61FA6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC4BB12FF1881AEFD9CFDE6.text	039187D8FFC4BB12FF1881AEFD9CFDE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia barclayi Gompel 2020	<div><p>Zarcosia barclayi n. sp.</p><p>(Figures 1; 3i,j; 4a; 6g)</p><p>Etymology. The species is named after Max Barclay in acknowledgment of his unflagging support over two decades to guide me through the insect research collections of the British Museum of Natural History.</p><p>Type locality. Malaysia: Sarawak, Gunung Mulu National Park, Deer Cave [04°05’’ N 114° 53’ E] .</p><p>Distribution. Malaysia (Figure 1).</p><p>Description. (Figure 3i,j). Body length. ♂: 1.74± 0.02 mm (n=2); ♀: 1.96± 0.13 mm (n=7). Dark brown, densely covered with silver applied pubescence. Appendages orange to brown.</p><p>Head. Wider than pronotum in males, as wide as pronotum in females. Palpi orange, last article securiform. Integument mat, finely rugose.</p><p>Antennae. A fourth to a third longer than elytra in males, shorter than elytra in females, orange, slightly and progressively thicker toward apex. Antennomere 4 and higher with a slight serration.</p><p>Pronotum (Figure 6g). Subcylindrical, convex, slightly longer than wide, marked by a strong lateral constriction on each side.</p><p>Elytra. Length: ♂: 1.20± 0.07 mm (n=2); ♀: 1.21± 0.08 mm (n=7). Finely and densely punctuated, covered with double pubescence. Moderately convex, depressed on disc.</p><p>Legs. No notable sexual characters. Male foretibia with a faint sinuation of their inner side.</p><p>Aedeagus (Figure 4a). Simple, narrow, progressively tapered, slightly and regularly bent in lateral view. Appendages visible near apex.</p><p>Type material. Holotype (♂, BMNH). Malaysia: Sarawak, Gunung Mulu National Park, Deer Cave, 04°07’’ N 114° 53’ E (360 m) (P. M. Hammond &amp; J. E. Marshall leg.) 6.V.1978 . Paratypes. Malaysia: Sarawak, Gunung Mulu National Park, Deer Cave, 04°07’’ N 114° 53’ E (360 m) (P. M. Hammond &amp; J. E. Marshall leg.) 6.V.1978 (3 ♀ paratypes, BMNH) . Sarawak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.916664&amp;materialsCitation.latitude=4.116667" title="Search Plazi for locations around (long 114.916664/lat 4.116667)">Gunung Mulu National Park</a>, Clearwater Cave, 04°07’ N 114°55’ E (360 m) (P. M. Hammond &amp; J. E. Marshall leg.) 13.V.1978 (1 ♀ paratype, BMNH) , 20.V.1978 (2 ♀ paratypes, BMNH; 1 ♀ paratype, 1 ♂ paratype, NGPC) .</p></div>	https://treatment.plazi.org/id/039187D8FFC4BB12FF1881AEFD9CFDE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC5BB12FF18821AFAA7F837.text	039187D8FFC5BB12FF18821AFAA7F837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia batuensis (Werner 1962) Gompel 2020	<div><p>Zarcosia batuensis (Werner, 1962), comb. nov.</p><p>(Figures 1; 2i)</p><p>Euglenes batuensis Werner, 1962: 125; Type: Bishop Museum (Hawaii, USA); 3 paratypes at the BMNH</p><p>Zarcosia batuensis (Werner, 1962): unpublished combination in Grzymala, 2016</p><p>Type locality. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves .</p><p>Distribution. Malaya, Batu caves (Figure 1).</p><p>Material examined. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves, Cavern C-Loc. 9 (86 m), Light trap, 10 hrs, (H. E. McClure leg.) 11.VIII.1959 (1 ♀ paratype, BMNH) ; Selangor Darul Ehsan, Gombak District, Batu caves, Cavern C-Loc. 3, Penny Room (H. E. McClure leg.) 8.XII.1959 (1 ♂ paratype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC5BB12FF18821AFAA7F837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC2BB15FF1884DDFAEFFC59.text	039187D8FFC2BB15FF1884DDFAEFFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia bedosae Gompel 2020	<div><p>Zarcosia bedosae n. sp.</p><p>(Figures 1; 3m; 4b; 6i)</p><p>Etymology. The species is named after Anne Bedos in recognition of her efforts to collect material for others.</p><p>Type locality. Laos: Khammouan province, Ban Thôngkha, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.51667&amp;materialsCitation.latitude=17.966667" title="Search Plazi for locations around (long 104.51667/lat 17.966667)">Tham Thon</a> [17°58’ N 104°31’ E] .</p><p>Distribution. Laos (Figure 1).</p><p>Description. (Figure 3m). Female unknown. Body length. ♂ holotype: 2.5 mm. Black, mat, covered with very short, pulverulent, silver pubescence. Appendages brown.</p><p>Head. Moderately transverse, wider than pronotum. Palpi yellowish, last article securiform. Integument mat, coarsely rugose, grainy. Occiput lined with a fringe of dense and short silver hairs posteriorly.</p><p>Antennae. A fifth longer than elytra, entirely dark brown, filiform.</p><p>Pronotum (Figure 6i). Convex, slightly longer than wide, sides parallel from the base to the posterior two thirds, then progressively convergent anteriorly. Sides marked with lateral impressions and callosities.</p><p>Elytra. Length: 1.75 mm. Punctation regular, as dense as on the pronotum, with larger points. Space between points mat, alutaceous. Disc subconvex, with two broad depression in the anterior third. Pubescence dense and short, comprised of very short hairs borne from the points and interspersed pulverulent hairs.</p><p>Legs. Particularly elongated. No other notable modifications.</p><p>Aedeagus (Figure 4b). Sturdy, sides weakly convergent in dorsal view and suddenly tapered apically. Strongly convex in lateral view, bent upward at apex.</p><p>Type material. Holotype (♂, MNHN). Laos: Khammouan province, Ban Thôngkha, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.51667&amp;materialsCitation.latitude=17.966667" title="Search Plazi for locations around (long 104.51667/lat 17.966667)">Tham Thon</a> (230 m), cave, by hand, 17°58’ N 104°31’ E (L. Deharveng &amp; A. Bedos) 12.II.1998.</p><p>Note. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.</p></div>	https://treatment.plazi.org/id/039187D8FFC2BB15FF1884DDFAEFFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC2BB15FF1887BDFE62FA96.text	039187D8FFC2BB15FF1887BDFE62FA96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia bipartita (Pic 1901) Gompel 2020	<div><p>Zarcosia bipartita (Pic, 1901), comb. nov.</p><p>(Figures 1; 2v; 4c)</p><p>Hylophilus bipartitus Pic, 1901: 96; Syntypes: MNHN, BMNH</p><p>Hylophilus bipartitus Pic, 1901: Pic, 1901</p><p>Zarcosia bipartitus [sic!] (Pic, 1901): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Malabar .</p><p>Distribution. India (Figure 1).</p><p>Material examined. India: Malabar District (1 ♂, MNHN), ex coll. Pic (1 ♂, 1 ♀, BMNH) ; Kerala, Malabar District, Mahé (3 ♂, MNCNM) .</p></div>	https://treatment.plazi.org/id/039187D8FFC2BB15FF1887BDFE62FA96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC2BB14FF1881F1FC9CFC7E.text	039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia brunnea Gompel 2020	<div><p>Zarcosia brunnea n. sp.</p><p>(Figures 1; 3t; 4w; 5s; 6o)</p><p>Etymology. The species is named in reference to the overall brown color of the specimens.</p><p>Type locality. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary [07°13’ N 80°57’ E] .</p><p>Distribution. Sri Lanka (Figure 1).</p><p>Description. (Figure 3t). Female unknown. Body length. ♂: 2.80± 0.13 mm (n=3). Robust, sturdy, uniformly reddish brown, covered with double pubescence.</p><p>Head. Transverse, wider than pronotum, covered with blond pubescence. Punctation fine and very dense. Palpi dark orange, last article securiform.</p><p>Antennae. Strong, shorter than elytra, antennomere 3–10 short, at most twice longer than wide, conical. Covered with hirsute blond pubescence.</p><p>Pronotum (Figure 6o). Globose, as long as wide, disc convex. Sides diverging outward from the base to the apical third, then converging markedly inward toward apical margin anteriorly. Densely punctuated, space between points smaller than the diameters of points. Covered with double pubescence, moderately long blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a dense fringe of golden pubescence anterior to the scutellum.</p><p>Elytra. Length: ♂: 1.92± 0.03 mm (n=3). Densely punctuated, space between points larger than the points themselves, covered with double pubescence. Moderately convex, depressed on disc. Apex with a few long hairs (similar to Z. srilankaensis n. sp., Figure 5h, but much fewer hairs).</p><p>Legs. Short, sturdy. Sexual characters: foretibia with a sharp tooth on their inner side (Figure 5s). Mesothoracic femora with an acute apical tooth, straight and pointing outward (similar to that of Z. dentatifemur (Pic, 1912), Figure 5w).</p><p>Aedeagus (Figure 4w). Sturdy, sides regularly parallel in dorsal view, strongly convergent in the apical fifth, ended with a small rounded lobe. Strongly and regularly bent in lateral view.</p><p>Type material. Sri Lanka: Holotype (♂, UCDC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 . Paratypes. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (1 ♂ paratype, NGPC) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2" title="Search Plazi for locations around (long 80.95/lat 7.2)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°12’ N 80°57’ E (135 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (1 ♂ paratype, UCDC) .</p><p>Note. This species was collected together with Z. dentatifemur (Pic, 1912), Z. spinifemur n. sp., Z. srilankaensis n. sp. and Z. intermedia n. sp. The sample contained a large series of females, which I could not assign to the respective species. In particular, Z. brunnea n. sp., Z. srilankaensis n. sp. and Z. intermedia n. sp. form a very homogeneous group. Their males are easily distinguished by stable sexual characters (aedeagus, teeth on the tibiae, hooks on the femora), but in spite of extensive comparisons of the females collected with them, I could not decide whether these females represented one or more taxa, nor which.</p></div>	https://treatment.plazi.org/id/039187D8FFC2BB14FF1881F1FC9CFC7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC3BB14FF188799FB8BFAFC.text	039187D8FFC3BB14FF188799FB8BFAFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia capitalis (Pic 1901) Gompel 2020	<div><p>Zarcosia capitalis (Pic, 1901), comb. nov.</p><p>(Figures 1; 2q)</p><p>Hylophilus capitalis Pic, 1901: 96; Syntype: MNHN</p><p>Zarcosia capitalis (Pic, 1901): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Malabar, Mahé.</p><p>Distribution. India: Kerala (Figure 1).</p><p>Material examined. India: Kerala, Malabar, Mahé (1 ♂ syntype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFC3BB14FF188799FB8BFAFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC3BB14FF18811EFE35F880.text	039187D8FFC3BB14FF18811EFE35F880.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia cephalica (Werner 1962) Gompel 2020	<div><p>Zarcosia cephalica (Werner, 1962), comb. nov.</p><p>(Figures 1; 2f; 4d; 5u)</p><p>Euglenes cephalicus Werner, 1962: 126; Holotype: Bishop Museum (Hawaii, USA); 1 paratype at the BMNH</p><p>Zarcosia cephalicus [sic!] (Werner, 1962): Grzymala, 2016</p><p>Type locality. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves .</p><p>Distribution. Malaysia: Batu caves; Sarawak, Gunung Mulu National Park (Figure 1).</p><p>Material examined. Malaysia: Sarawak, 4th Division, Gunung Mulu National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.916664&amp;materialsCitation.latitude=4.116667" title="Search Plazi for locations around (long 114.916664/lat 4.116667)">Deer Cave</a>, (360 m) 04°07’ N 114°55’ E (P. M. Hammond &amp; J. E. Marshall leg.) 6.V.1978 (1 ♂, 2 ♀, BMNH), 20.V.1978 (2 ♂, 1 ♀, BMNH; 1 ♂, 1 ♀, NGPC) ; Sarawak, 4th Division, Gunung Mulu National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.916664&amp;materialsCitation.latitude=4.116667" title="Search Plazi for locations around (long 114.916664/lat 4.116667)">Clearwater Cave</a>, (360 m) 04°07’ N 114°55’ E (P. M. Hammond &amp; J. E. Marshall leg.) 20.V.1978 (1 ♂, BMNH) . Republic of Indonesia: Aceh Selatan, Sawang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.12389&amp;materialsCitation.latitude=3.3930554" title="Search Plazi for locations around (long 97.12389/lat 3.3930554)">Tring Meuduro Tunong</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.12389&amp;materialsCitation.latitude=3.3930554" title="Search Plazi for locations around (long 97.12389/lat 3.3930554)">Gua Kiciarean</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.12389&amp;materialsCitation.latitude=3.3930554" title="Search Plazi for locations around (long 97.12389/lat 3.3930554)">Cave</a>, (30 m) 03°23’35” N 97°07’26” E (L. Deharveng &amp; A. Bedos leg.) 12.VII.1998 (1 ♂, 1 ♀, MNHN; 1 ♀, MZB), (J. Gibert leg.) 12.VII.1998 (1 ♀, MZB) ; Jawa Tengah, Cilacap, Nusakambangan Island, Goa Ratu, (70 m) cave, 07°43’ S 108°57’ (L. Deharveng &amp; A. Bedos leg.) 17.II.2011 (1♀, NGPC) .</p></div>	https://treatment.plazi.org/id/039187D8FFC3BB14FF18811EFE35F880	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC0BB17FF1884DDFAEFFBA5.text	039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia deharvengi Gompel 2020	<div><p>Zarcosia deharvengi n. sp.</p><p>(Figures 1; 3f; 4e; 5l; 6e)</p><p>Etymology. The species is named after Louis Deharveng in recognition of his efforts to collect material for others.</p><p>Type locality. Republic of Indonesia: Sumatera Barat, Baso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.46667&amp;materialsCitation.latitude=-0.23333333" title="Search Plazi for locations around (long 100.46667/lat -0.23333333)">Ngalau Simarasop</a> [00°14’ S 100°28’ E] .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Description. (Figure 3f). Female unknown. Body length. ♂: 2.43± 0.19 mm (n=3). Dark brown, darker anteriorly, gracile, covered with very short, pulverulent pubescence. Appendages orange-brown.</p><p>Head. Small, dark, barely wider than pronotum. Palpi yellow, last article securiform. Integument, finely and densely punctuated, grainy, looking like velvet. Occiput lined with a fringe of dense and short silver hairs posteriorly.</p><p>Antennae. 1.5 the length of elytra, orange-brown, regularly filiform with antenommeres 3–10 cylindrical.</p><p>Pronotum (Figure 6e). Round and convex, as long as wide, sides regularly curved. Base sinuated. Surface mat, rugose, covered with dense, velvety pubescence. Punctation undistinguishable.</p><p>Elytra. Length: 1.72± 0.13 mm (n=3). Punctation fine and shallow, dense (space between points as wide as the points themselves). Disc depressed. Pubescence dense and short, whitish, comprised of very short hairs borne from the points and interspersed pulverulent hairs.</p><p>Legs. Particularly elongated. Foretibia thickened in a smooth tooth on their inner side (Figure 5l).</p><p>Aedeagus (Figure 4e). Sides weakly and progressively convergent in dorsal view, apex rounded. Long, extremely bent in lateral view, apex with a brief downward deflection. Appendages visible near apex.</p><p>Type material. Holotype (♂, MNHN). Republic of Indonesia: Sumatra, West Sumatra, Baso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.46667&amp;materialsCitation.latitude=-0.23333333" title="Search Plazi for locations around (long 100.46667/lat -0.23333333)">Ngalau Simarasop</a>, (740 m) cave, 00°14’ S 100°28’ E (L. Deharveng &amp; A. Bedos leg.) 10.VII.2000 . Paratypes. Republic of Indonesia: Sumatra, Aceh, Aceh Besar Regency, Lhok’Nga, P. T. Semen Andalas, 05°27’ N 95°14’ E, Limestone Quarry, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.23333&amp;materialsCitation.latitude=5.45" title="Search Plazi for locations around (long 95.23333/lat 5.45)">Jungle Quarry Cave</a>, 2nd cavern, twilight zone, on sand near guano (I. Davis) 26.VII.1984 (1 ♂ paratype, NGPC) ; Sumatra, North Sumatra, Penen, Rampah Cave, 1 km E. of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.6325&amp;materialsCitation.latitude=3.3258333" title="Search Plazi for locations around (long 98.6325/lat 3.3258333)">Kota Tinggi</a>, 03°19’33” N 98°37’57” E (C. Covill) 26.VII.1984 (1 ♂ paratype, BMNH) .</p><p>Note. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.</p></div>	https://treatment.plazi.org/id/039187D8FFC0BB17FF1884DDFAEFFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC0BB17FF1880E1FEAAF9DC.text	039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia dentatifemur (Pic 1912) Gompel 2020	<div><p>Zarcosia dentatifemur (Pic, 1912), comb. nov.</p><p>(Figures 1; 2o; 3q; 4t; 5r, w; 6k)</p><p>Hylophilus dentatifemur Pic, 1912: 273; Holotype (male): BMNH</p><p>Zarcosia cephalicus [sic!] (Pic, 1912): unpublished combination in Grzymala, 2016</p><p>Type locality. Sri Lanka: Kandy .</p><p>Distribution. Sri Lanka: Kandy (Figure 1).</p><p>Material examined. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2" title="Search Plazi for locations around (long 80.95/lat 7.2)">Kandy</a> (G. E. Bryant). VIII.1908 (1 ♂ holotype, BMNH); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2" title="Search Plazi for locations around (long 80.95/lat 7.2)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°12‘ N 80°57‘ E (135 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (1 ♂, UCDC) .</p></div>	https://treatment.plazi.org/id/039187D8FFC0BB17FF1880E1FEAAF9DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC0BB17FF18823EFF63F837.text	039187D8FFC0BB17FF18823EFF63F837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia discoidalis (Pic 1902) Gompel 2020	<div><p>Zarcosia discoidalis (Pic, 1902), comb. nov.</p><p>(Figures 1; 2u)</p><p>Hylophilus discoidalis Pic, 1902: 34; Syntype (male): MNHN</p><p>Zarcosia discoidalis (Pic, 1902): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Malabar, Mahé.</p><p>Distribution. India: Kerala (Figure 1).</p><p>Material examined. India: Kerala, Malabar, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=75.53333&amp;materialsCitation.latitude=11.7" title="Search Plazi for locations around (long 75.53333/lat 11.7)">Mahé</a>, 11°42’ N 75°32’ E (P. Dorme) 1.XI.1888 (1 ♂ syntype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFC0BB17FF18823EFF63F837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC1BB16FF1884DDFF63FE14.text	039187D8FFC1BB16FF1884DDFF63FE14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia genjiensis (Pic 1917) Gompel 2020	<div><p>Zarcosia genjiensis (Pic, 1917), comb. nov.</p><p>(Figures 1; 2x)</p><p>Hylophilus genjiensis Pic, 1917: 260; Syntype (male): MNHN</p><p>Zarcosia genjiensis (Pic, 1917): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Rajasthan, Genji (« Inde: Coromandel, à Genji ») .</p><p>Distribution. India: Rajasthan (Figure 1).</p><p>Material examined. India: Rajasthan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.683334&amp;materialsCitation.latitude=23.7" title="Search Plazi for locations around (long 73.683334/lat 23.7)">Genji</a>, 23°42’ N 73°41’ E (M. Maindron) 15.IX.1901 (1 ♂ syntype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFC1BB16FF1884DDFF63FE14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC1BB16FF188676FBBFFA01.text	039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia gerstmeieri Gompel 2020	<div><p>Zarcosia gerstmeieri n. sp.</p><p>(Figures 1; 3d; 4f; 5j,v; 6c)</p><p>Etymology. The species is named after my colleague Roland Gerstmeier who collected this new species, in appreciation of his support of my work on Aderidae .</p><p>Type locality. Vietnam: Ninh Bình Province, Nho Quan District, Cúc Phýõng National Park .</p><p>Distribution. Vietnam: Ninh Bình Province (Figure 1).</p><p>Description. (Figure 3d). Female unknown. Body length. ♂ holotype: 3.25 mm. Robust, sturdy, entirely dark, covered with double golden pubescence.</p><p>Head. Weakly transverse, barely wider than pronotum, covered with white pubescence. Pubescence particularly dense anteriorly, between the antennal space. Punctation coarse and very dense. Palpi orange, last article securiform. Posterior margin lined with a fringe of white pubescence.</p><p>Antennae. Strong, serrated, as long as elytra, antennomeres 3–7 long, antennomeres 8–10 progressively shorter. Covered with hirsute blond pubescence.</p><p>Pronotum (Figure 6c). Convex, subrectangular, slightly longer than wide. Sides parallel from base to anterior fifth, then converging markedly inward toward apical margin anteriorly. Densely punctuated, points nearly adjacent. Covered with double pubescence, moderately long blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a dense fringe of golden pubescence anterior to the scutellum and at each posterior angle.</p><p>Elytra. Length: 1.15 mm. Punctation strong, points well delineated, deep, space between points larger than the points themselves. Densely covered with double pubescence (Figure 5j). Weakly convex.</p><p>Legs. Short, sturdy, dark. No visible sexual characters.</p><p>Aedeagus (Figure 4f). Very sturdy, strongly sclerotized, sides sinuated in dorsal view. Apex complex, lozenge shaped, terminated with a stinger. Strongly bent in lateral view, with an upward kink in apical fifth and a brief downward deflection at the tip.</p><p>Type material. Holotype (♂, NME). Vietnam: Ninh Bình Province, Nho Quan District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.59334&amp;materialsCitation.latitude=20.350279" title="Search Plazi for locations around (long 105.59334/lat 20.350279)">Cúc Phýõng National Park</a>, end of road, 20°21’01” N 105°35’36” E (375 m) (R. Gerstmeier) 6.V.2017.</p></div>	https://treatment.plazi.org/id/039187D8FFC1BB16FF188676FBBFFA01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC1BB16FF18821AFEF1F879.text	039187D8FFC1BB16FF18821AFEF1F879.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia glaucescens (Champion 1920) Gompel 2020	<div><p>Zarcosia glaucescens (Champion, 1920), comb. nov.</p><p>(Figures 1; 2s)</p><p>Xylophilus glaucescens Champion, 1920: 395; Holotype (male): BMNH</p><p>Type locality. Republic of the Philippines: Island of Basilan .</p><p>Distribution. Philippines: Island of Basilan (Figure 1).</p><p>Material examined. Republic of the Philippines: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=122.0&amp;materialsCitation.latitude=7.0" title="Search Plazi for locations around (long 122.0/lat 7.0)">Island</a> of Basilan, 07°00’ N 122°00’ E, 28.VIII.1921 (1 holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC1BB16FF18821AFEF1F879	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCCBB1BFF188709FAC9FB4F.text	039187D8FFCCBB1BFF188709FAC9FB4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia grandiceps Pic 1899	<div><p>Zarcosia grandiceps Pic, 1899, comb. nov.</p><p>(Figures 1; 2j)</p><p>Hylophilus grandiceps Pic, 1899: 756; Type: 1 holotype and 1 paratype, MNHN</p><p>Hylophilus grandiceps Pic, 1899: Pic, 1902</p><p>Hylophilus grandiceps Pic, 1899: Pic, 1905</p><p>Zarcosia grandiceps (Pic, 1899): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Sumatra, Palembang .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Material examined. Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.76667&amp;materialsCitation.latitude=-2.9666667" title="Search Plazi for locations around (long 104.76667/lat -2.9666667)">Palembang</a>, 02°58’ S 104°46’ E (2 sp., MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFCCBB1BFF188709FAC9FB4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCCBB1BFF18814BFE17F9AA.text	039187D8FFCCBB1BFF18814BFE17F9AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia holosericea (Champion 1916) Gompel 2020	<div><p>Zarcosia holosericea (Champion, 1916), comb. nov.</p><p>(Figures 1; 2n)</p><p>Xyophilus holosericeus Champion, 1916: 28; Holotype (male): BMNH</p><p>Zarcosia holosericeus [sic!] (Champion, 1916): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Himachal Pradesh, Kangra Valley .</p><p>Distribution. India: Himachal Pradesh (Figure 1).</p><p>Material examined. India: Himachal Pradesh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.5&amp;materialsCitation.latitude=32.15" title="Search Plazi for locations around (long 76.5/lat 32.15)">Kangra Valley</a>, 32°09’ N 76°30’ E (1400 m) (Dudgeon) VI.1899 (1 holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFCCBB1BFF18814BFE17F9AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCCBB1AFF1882EDFAEFFD56.text	039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia ilonae Gompel 2020	<div><p>Zarcosia ilonae n. sp.</p><p>(Figures 1; 3c; 4g; 5k; 6b)</p><p>Etymology. The species is named after my wife, Ilona Grunwald, to acknowledge her patience and support of my work with insects.</p><p>Type locality. Thailand: Surat Thani Province, Phanom District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.65&amp;materialsCitation.latitude=8.866667" title="Search Plazi for locations around (long 98.65/lat 8.866667)">Tham Wararam</a>, 08°52’ N 98°39’ E .</p><p>Distribution. Thailand (Figure 1).</p><p>Description. (Figure 3c). Female unknown. Body length. ♂ holotype: 2.55 mm. Slender, orange-brown with darker head, covered with very short, pulverulent, white pubescence. Appendages pale.</p><p>Head. Moderately transverse, barely wider than pronotum. Palpi yellow, last article securiform. Finely and densely punctuated. Occiput lined with a fringe of dense and short white hairs posteriorly.</p><p>Antennae. 1.5 times longer than elytra, entirely yellowish, filiform with antenommeres 3–10 elongated.</p><p>Pronotum (Figure 6b). Convex, slightly longer than wide, sides regularly curved, wider, almost gibbose in its anterior third.</p><p>Elytra. Length: 1.80 mm. Punctation strong and dense, with larger points. Space between points slightly wider than the points themselves. Disc subconvex. Pubescence dense and short, comprised of very short hairs borne from the points and interspersed pulverulent hairs.</p><p>Legs. Particularly elongated. Foretibia weakly broadened in the middle of their inner side (Figure 5k).</p><p>Aedeagus (Figure 4g). Sturdy, lightly pigmented, sides divergent from base in dorsal view, then largely rounded near apex and ending in a broad stinger. Nearly flat in lateral view, with no kink. Lateral appendages thick and as long as the apical stinger.</p><p>Type material. Holotype (♂, MNHN). Thailand: Surat Thani Province, Phanom District, Tham Wararam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.65&amp;materialsCitation.latitude=8.866667" title="Search Plazi for locations around (long 98.65/lat 8.866667)">Cave</a>, 08°52’ N 98°39’ E (250 m) (L. Deharveng and A. Bedos) 24.VII.1987.</p><p>Note. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.</p></div>	https://treatment.plazi.org/id/039187D8FFCCBB1AFF1882EDFAEFFD56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCDBB1DFF1886B1FC9CFF2A.text	039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia intermedia Gompel 2020	<div><p>Zarcosia intermedia n. sp.</p><p>(Figures 1; 3s; 4v; 5q; 6l)</p><p>Etymology. The species is named in reference of the intermediate morphology of its males, combining secondary sexual characters found in Z. srilankaensis and Z. dentatifemur .</p><p>Type locality. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary [07°13’ N 80°57’ E] .</p><p>Distribution. Sri Lanka (Figure 1).</p><p>Description. (Figure 3s). Female unknown. Body length. ♂: 2.83± 0.08 mm (n=3). Robust, sturdy, uniformly reddish brown, covered with double pubescence.</p><p>Head. Transverse, wider than pronotum, covered with blond pubescence. Punctation coarse and dense. Palpi orange, last article securiform.</p><p>Antennae. Strong, shorter than elytra, antennomere 3–10 short, 8–10 as long as wide, conical. Covered with hirsute blond pubescence.</p><p>Pronotum (Figure 6l). Globose, as long as wide, disc convex. Sides diverging outward from the base to the apical third, then converging markedly inward toward apical margin anteriorly. Densely punctuated, space between points smaller than the diameters of points. Covered with double pubescence, moderately long blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a dense fringe of golden pubescence anterior to the scutellum.</p><p>Elytra. Length: ♂: 1.86± 0.04 mm (n=3). Densely punctuated, space between points larger than the points themselves, covered with double pubescence. Moderately convex, depressed on disc. Apex with a a tuft of hairs along the suture (similar to Z. srilankaensis n. sp., Figure 5h).</p><p>Legs. Short, sturdy. Sexual characters: foretibia with a sharp tooth on their inner side (Figure 5q). Mesothoracic femora with an acute apical tooth, straight and pointing outward (similar to that of Z. dentatifemur (Pic, 1912), Figure 5w).</p><p>Aedeagus (Figure 4v). Sturdy, sides regularly parallel in dorsal view, strongly convergent in the apical fifth, shortly tapered. Moderately bent in lateral view with a slight apical sinuation (no deflection).</p><p>Type material. Holotype (♂, UCDC). Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 . Paratypes. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (1 ♂ paratype, UCDC; 1 ♂ paratype, NGPC) .</p><p>Note. This species was collected together with Z. dentatifemur (Pic, 1912), Z. spinifemur n. sp., Z. srilankaensis n. sp. and Z. brunnea n. sp. The sample contained a large series of females, which I could not assign to the respective species. In particular, Z. brunnea n. sp., Z. srilankaensis n. sp. and Z. intermedia n. sp. form a very homogeneous group. Their males are easily distinguished by stable sexual characters (aedeagus, teeth on the tibiae, hooks on the femora), but in spite of extensive comparisons of the females collected with them, I could not decide whether these females represented one or more taxa, nor which.</p></div>	https://treatment.plazi.org/id/039187D8FFCDBB1DFF1886B1FC9CFF2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCABB1DFF18856DFB11FD8A.text	039187D8FFCABB1DFF18856DFB11FD8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia javana (Pic 1894) Gompel 2020	<div><p>Zarcosia javana (Pic, 1894), comb. nov.</p><p>(Figures 1; 2z)</p><p>Hylophilus javanus Pic, 1894: 433; Type: MNHN</p><p>Hylophilus javanus Pic, 1894: Pic, 1899</p><p>Zarcosia javanus [sic!] (Pic, 1894): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Java .</p><p>Distribution. Republic of Indonesia: Java (Figure 1).</p><p>Material examined. Republic of Indonesia: Java (Staudinger) (2 syntypes, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFCABB1DFF18856DFB11FD8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCABB1DFF18868DFCF6FBE4.text	039187D8FFCABB1DFF18868DFCF6FBE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia kempi (Blair 1924) Gompel 2020	<div><p>Zarcosia kempi (Blair, 1924), comb. nov.</p><p>(Figures 1; 2c; 4h)</p><p>Hylophilus kempi Blair, 1924: 121; Type: BMNH</p><p>Zarcosia kempi (Blair, 1924): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Meghalaya, Assam, Siju cave, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=90.683334&amp;materialsCitation.latitude=25.35" title="Search Plazi for locations around (long 90.683334/lat 25.35)">Garo Hills</a> [25°21’ N 90°41’ E] .</p><p>Distribution. India: Meghalaya, Assam (Figure 1).</p><p>Material examined. India: Meghalaya, Assam, Siju cave, 400 ft from entrance, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=90.683334&amp;materialsCitation.latitude=25.35" title="Search Plazi for locations around (long 90.683334/lat 25.35)">Garo Hills</a>, 25°21’ N 90°41’ E (60 m)(S. K. &amp; B. N. C.) II.1922 (1 ♂ holotype &amp; 1 ♀, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFCABB1DFF18868DFCF6FBE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCABB1CFF188026FF63FBCA.text	039187D8FFCABB1CFF188026FF63FBCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia lemairei Gompel 2020	<div><p>Zarcosia lemairei n. sp.</p><p>(Figures 1; 3a,b; 4i; 5 a–d; 6a)</p><p>Etymology. The species is named after Jean-Michel Lemaire, who sent me the first specimens of this species, thereby seeding the present synopsis.</p><p>Type locality. Laos: Khammouan province, Boualapha, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.833336&amp;materialsCitation.latitude=17.583334" title="Search Plazi for locations around (long 105.833336/lat 17.583334)">Tham Nam Ock</a> [17°35’ N 105°50’ E] .</p><p>Distribution. Laos, Myanmar, Vietnam (Figure 1).</p><p>Description. (Figure 3 a–b). Body length. ♂: 3.70± 0.28 mm (n=6); ♀: 3.98± 0.29 mm (n=12). Entirely black, slender, with long appendages. Body shape reminiscent of strict cave beetles ( Carabidae: Trechini or Leiodidae: Leptodirini). Pubescence dense, very short, almost pulverulent, composed of stout bristles arising from the punctation, interspersed with very short whitish hairs, arising from the entire body surface, and giving a mat appearance to the insect.</p><p>Head (Figure 5 a–c). As wide as pronotum in dorsal view. Eyes bigger in males than in females. Palpi black, last article securiform, as broad as the length of the first two antennomeres together. Frons deeply carved in males, flat in females. The deep pit on the male forehead differs in shape from that of Zarcosia palliditarsis (Pic, 1912) (Figure 5 e–g) and possesses a long and wavy tuft of silky yellow bristles, borne from the bottom of the head pit.</p><p>Antennae. Filiform, 20% longer than elytra in males, hardly longer than elytra in females.</p><p>Pronotum. Slightly longer than wide. Sides convergent in their anterior third, otherwise roughly parallel to each other, with a slight sinuosity or constriction. The dorsal surface is regularly rounded, with an anterior gibbosity. Combined, the anterior gibbosity and the slight posterior constriction place the largest width of the pronotum in its anterior third.</p><p>Base of the pronotum with two sinuosities: one at the level of the midline and two flanking this first one, on each side. Sexual dimorphism: in males, the dorsal surface is posteriorly impressed by ill-defined foveola that singles out a carena-like bulge along the midline.</p><p>Elytra. ♂: 2.61± 0.18 mm (n=6); ♀: 2.71± 0.21 mm (n=12). 2.3 longer than wide in both sexes, with sides slightly but regularly curved in females, straight, convergent in their posterior half in males. Dorsal surface moderately convex.</p><p>Legs. Black, except the second half of the penultimate and the last tarsal segment that are orange in both sexes. Surface of femora micro-reticulated. Sexual dimorphism: (i) Tibias of forelegs and hindlegs slightly thickened and sinuated in males, flattened on their inner face; (ii) first tarsal segments of mesothoracic legs thickened in males (Figure 5d, left); (iii) hindtarsae extraordinarily elongated in males, in particular the last segment, which is 50–70% the length of the first tarsal segment (Figure 5d, right). By contrast, in females, the last segment of the hindtarsae is 40% the length of the first tarsal segment.</p><p>Aedeagus. Figure 4i. Slender, narrow, sides weakly convergent in dorsal view. Slight curvature in lateral view, with apex sharply bent upward.</p><p>Type material. Holotype (♂, MNHN). Laos: Khammouan province, Boualapha, Tham Nam Ock, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.833336&amp;materialsCitation.latitude=17.583334" title="Search Plazi for locations around (long 105.833336/lat 17.583334)">Cave</a>, Hin Nam <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.833336&amp;materialsCitation.latitude=17.583334" title="Search Plazi for locations around (long 105.833336/lat 17.583334)">No Project</a>, 17°35’ N 105°50’ E (270 m) (H. Steiner) 3.III.2016 . Paratypes. Laos: Ban Du, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.816666&amp;materialsCitation.latitude=17.55" title="Search Plazi for locations around (long 105.816666/lat 17.55)">Tham Pha Houm</a>, cave, 17°33’ N 105°49’ E (170 m) (J. Lips) 1.III.2016 (2 ♂ paratypes, NGPC; 3 ♀ paratypes, JMLC) ; Nong Ping, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.9&amp;materialsCitation.latitude=17.35" title="Search Plazi for locations around (long 105.9/lat 17.35)">Tham Pha Pong</a>, cave, 17°21’ N 105°54’ E (226 m) (J. Lips) 25.II.2016 (1 ♀ paratype, JMLC) ; Vientiane Province, Vang Vieng, Tham Ling Kham, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.433334&amp;materialsCitation.latitude=18.933332" title="Search Plazi for locations around (long 102.433334/lat 18.933332)">Cave</a>, 18°56’ N 102°26’ E, by hand (L. Johansson) 18.II.2004 (1 ♀ paratype, MNHN) ; Vientiane, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.88333&amp;materialsCitation.latitude=19.016666" title="Search Plazi for locations around (long 101.88333/lat 19.016666)">Tham Paka</a>, 19°01’ N 101°53’ E, cave (M. Ferrand) 5.III.2016 (1 ♂ and 1 ♀ paratypes, JMLC) ; Khammouan province, Boualapha, Tham Nam Ock, Cave, Hin Nam <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.833336&amp;materialsCitation.latitude=17.583334" title="Search Plazi for locations around (long 105.833336/lat 17.583334)">No Project</a>, 17°35’ N 105°50’ E (270 m) (H. Steiner) 3.III.2016 (1 ♀ paratype, NGPC) ; Khammouan province, Boualapha, Tham Xe Bangfai, cave, Hin Nam <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.833336&amp;materialsCitation.latitude=17.366667" title="Search Plazi for locations around (long 105.833336/lat 17.366667)">No Project</a>, 17°22’ N 105°50’ E (270 m) (H. Steiner) 22.II.2016 (1 ♀ paratype, NGPC) ; Myanmar: Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.86667&amp;materialsCitation.latitude=22.2" title="Search Plazi for locations around (long 95.86667/lat 22.2)">Mondawa Gu</a>, cave, 22°12’ N 95°52’ E, by hand (F. Brehier) 1.XII.2015 (1 ♀ paratype, MNHN) ; Vietnam: Hà Nam Ninh Province, Cúc Phuong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.6&amp;materialsCitation.latitude=20.3" title="Search Plazi for locations around (long 105.6/lat 20.3)">Water Fairy cave</a>, 20°18’ N 105°36’ E, by hand (L. Deharveng) 11.X.1998 (1 ♀ paratype, NGPC) ; Bắc Kạn Province, Ba B ể District, Hoàng Trĩ, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=22.316668" title="Search Plazi for locations around (long 105.63333/lat 22.316668)">Nà</a> sai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=22.316668" title="Search Plazi for locations around (long 105.63333/lat 22.316668)">Tham Phay Cave</a>, 22°19’ N 105°38’ E (490 m) (A. Faille) 22.X.2018 (1 ♂ paratype, SMNS; 1 ♂ paratype, NGPC) ; Quang Binh, Cha Noi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.1&amp;materialsCitation.latitude=17.633333" title="Search Plazi for locations around (long 106.1/lat 17.633333)">Hang Cha Noi</a>, 17°38’ N 106°06’ E (260 m), by hand, (L. Deharveng and A. Bedos) 8.I.1995 (1 ♀ paratype, MNHN) ; Quang Binh, Dong Hoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.23333&amp;materialsCitation.latitude=17.65" title="Search Plazi for locations around (long 106.23333/lat 17.65)">Grotte de Troc</a>, 17°39’ N 106°14’ E (70 m), by hand (L. Deharveng and A. Bedos) 15.III.1997 (1 ♀ paratype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFCABB1CFF188026FF63FBCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCBBB1CFF1880CDFE7FFA2A.text	039187D8FFCBBB1CFF1880CDFE7FFA2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia longithorax (Pic 1899) Gompel 2020	<div><p>Zarcosia longithorax (Pic, 1899), comb. nov.</p><p>(Figures 1; 2g)</p><p>Hylophilus longithorax Pic, 1899: 755; Type: MNHN</p><p>Zarcosia longithorax (Pic, 1899): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Sumatra .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Material examined. Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.666664&amp;materialsCitation.latitude=2.3333333" title="Search Plazi for locations around (long 98.666664/lat 2.3333333)">Lau Rakit</a>, 02°20’ N 98°40’ E (J. B. Corporaal) 28.VIII.1921 (1 sp., BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFCBBB1CFF1880CDFE7FFA2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFCBBB1CFF18826DFDFBF866.text	039187D8FFCBBB1CFF18826DFDFBF866.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia lucifuga (Heberdey 1931) Gompel 2020	<div><p>Zarcosia lucifuga (Heberdey, 1931), comb. nov.</p><p>(Figures 1; 2 aa)</p><p>Hylophilus lucifugus Heberdey, 1931: 678; Type: The original description refers to 8 males; 1 syntype at MNHN</p><p>Hylophilus lucifugus Heberdey, 1931: Werner, 1962</p><p>Zarcosia lucifugus [sic!] (Heberdey, 1931): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Sumatra, Fort de Kock .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Material examined. Republic of Indonesia: Sumatra, Fort de Kock, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.36667&amp;materialsCitation.latitude=-0.3" title="Search Plazi for locations around (long 100.36667/lat -0.3)">Pauh Cave</a>, 00°18’ S 100°22’ E (Feuerborn) 13.III.1929 (1 ♂ syntype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFCBBB1CFF18826DFDFBF866	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC8BB1FFF1884DDFD07FE14.text	039187D8FFC8BB1FFF1884DDFD07FE14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia luteitarsis (Champion 1924) Gompel 2020	<div><p>Zarcosia luteitarsis (Champion, 1924), comb. nov.</p><p>(Figures 1; 2e)</p><p>Xylophilus luteitarsis Champion, 1924: 484; Holotype (male): BMNH</p><p>Zarcosia luteitarsis (Champion, 1924): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Uttarakhand, Ranikhet, Kumaon .</p><p>Distribution. Northern India (Figure 1).</p><p>Material examined. India: Uttarakhand, Ranikhet, Kumaon (1 ♂ holotype, BMNH) ; Sikkim, Gopaldhara, Rungbong Valley (H. Stevens) (2 ♂, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC8BB1FFF1884DDFD07FE14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC8BB1FFF188676FBA8FC6F.text	039187D8FFC8BB1FFF188676FBA8FC6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia malayana (Werner 1962) Gompel 2020	<div><p>Zarcosia malayana (Werner, 1962), comb. nov.</p><p>(Figures 1; 2h)</p><p>Euglenes malayanus Werner, 1962: 125; Holotype: Bishop Museum (Hawaii, USA); 4 paratypes at the BMNH</p><p>Zarcosia malayanus [sic!] (Werner, 1962): unpublished combination in Grzymala, 2016</p><p>Type locality. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves .</p><p>Distribution. Malaysia: Selangor Darul Ehsan, Gombak District (Figure 1).</p><p>Material examined. Malaysia: Selangor Darul Ehsan, Gombak District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.683334&amp;materialsCitation.latitude=3.2333333" title="Search Plazi for locations around (long 101.683334/lat 3.2333333)">Batu</a> caves, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.683334&amp;materialsCitation.latitude=3.2333333" title="Search Plazi for locations around (long 101.683334/lat 3.2333333)">Cavern</a> C-Loc. 9, 03°14’ N 101°41’ E (86 m), light trap (H. E. McClure) 5.IX.1959 (1 ♂ paratype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC8BB1FFF188676FBA8FC6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC8BB1FFF1887A8FF63FAC9.text	039187D8FFC8BB1FFF1887A8FF63FAC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia nathani (Pic 1943) Gompel 2020	<div><p>Zarcosia nathani (Pic, 1943), comb. nov.</p><p>(Figures 1; 2 ai)</p><p>Hylophilus nathani Pic, 1943: 5; Holotype: MNHN</p><p>Zarcosia nathani (Pic, 1943): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Tamil Nadu, Coimbatore .</p><p>Distribution. India: Tamil Nadu (Figure 1).</p><p>Material examined. India: Tamil Nadu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.96667&amp;materialsCitation.latitude=11.066667" title="Search Plazi for locations around (long 76.96667/lat 11.066667)">Coimbatore</a>, 11°04’ N 76°58’ E, 9.VIII.1932 (1 ♂ holotype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFC8BB1FFF1887A8FF63FAC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC8BB1FFF188129FF68F924.text	039187D8FFC8BB1FFF188129FF68F924.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia nigrofasciata (Pic 1899) Gompel 2020	<div><p>Zarcosia nigrofasciata (Pic, 1899), comb. nov.</p><p>(Figures 1; 2 ad)</p><p>Hylophilus nigrofasciatus Pic, 1899: 756; Holotype: MNHN</p><p>Zarcosia nigrofasciatus [sic!] (Pic, 1899): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Sumatra, Palembang .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Material examined. Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.76667&amp;materialsCitation.latitude=-2.9666667" title="Search Plazi for locations around (long 104.76667/lat -2.9666667)">Palembang</a>, 02°58’ S 104°46’ E (1 ♂ holotype, BMNH)</p></div>	https://treatment.plazi.org/id/039187D8FFC8BB1FFF188129FF68F924	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC8BB1EFF188366FBD5FBF5.text	039187D8FFC8BB1EFF188366FBD5FBF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia obscuricornis (Pic 1927) Gompel 2020	<div><p>Zarcosia obscuricornis (Pic, 1927), comb. nov.</p><p>(Figures 1; 2 ag; 4j)</p><p>Hylophilus grandicornis v. obscuricornis Pic, 1927: 30; Type: MNHN</p><p>Hylophilus grandicornis Pic: Pic, 1952</p><p>Zarcosia grandicornis (Pic): unpublished combination in Grzymala, 2016</p><p>Zarcosia grandicornis v. obscuricornis (Pic, 1927): unpublished combination in Grzymala, 2016</p><p>Hylophilus grandicornis was never described. Pic (1952) refers to this taxon, and the Zoological record (1923, vol. LX) indicates a reference for this description (Mélanges Exotico-Entomologiques, Vol. 38, p. 4), but after a careful check, I found no such description in the volume cited, nor in the entire publications by Pic in Mélanges Exotico-Entomologiques. I also surveyed a large number of Pic’s papers where Aderidae were discussed and could not find the trace of a description of this taxon. Winkler’s catalogue, 1927 does not list it either. Leaning on the ZCN article 11.1 (ICZN, 1999), I therefore consider that the name Hylophilus grandicornis Pic is not an available name, although, there is a single specimen from Pic’s original collection at the MNHN labelled “ Hylophilus grandicornis n. sp. ”. I suppose that the description has remained in litteris.</p><p>In 1927, Pic described a variety of Hylophilus grandicornis under the name Hylophilus grandicornis v. obscuricornis . The description is limited to 3 words (Antennis nigro-piceis) and was made in reference to grandicornis s. str. After examining the type specimen of this taxon in Pic’s collection (MNHN), it appears to be a bona fide species. I therefore consider that the name Hylophilus obscuricornis Pic, 1927 is available, and that this taxon should be placed in the genus Zarcosia, under the name Zarcosia obscuricornis (Pic, 1927), comb. nov.</p><p>Finally, the type in litteris of Hylophilus grandicornis is similar to that of Hylophilus obscuricornis Pic, 1927 and belongs to this taxon.</p><p>Type locality. Republic of Indonesia: Sumatra, Palembang .</p><p>Distribution. East China, Vietnam (Figure 1).</p><p>Material examined. China: no locality (1 sp., MNHN); Foochow (M. S. Yang) 1938 (1 ♂, NGPC, from collection Pic) ; Yunnan, Xishuangbanna, Guo Men Shan (NNNR), 37 km NW Jinghong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.63333&amp;materialsCitation.latitude=22.283333" title="Search Plazi for locations around (long 100.63333/lat 22.283333)">Transition</a> zone MF, 22°17’ N 100°38’ E (1080 m), (A. Weigel) 26.V.2008 (1 ♂, NME) ; Vietnam: Ho Chi Minh City (J. Barbier) 24.V.1950 (1 sp., MNHN); Hòa Bình (1 ♂ holotype and 17 sp., MNHN); Hòa Bình (J. De Cooman) 1923 (1 ♂, type in litteris of Hylophilus grandicornis Pic, MNHN); Lạc Thô, 20°24’ N 105°36’ E (1 ♂, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFC8BB1EFF188366FBD5FBF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC9BB1EFF188011FE08F8ED.text	039187D8FFC9BB1EFF188011FE08F8ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia palliditarsis (Pic 1912) Gompel 2020	<div><p>Zarcosia palliditarsis (Pic, 1912), comb. nov.</p><p>(Figures 1; 2t; 4k; 5 e–g)</p><p>Hylophilus palliditarsis Pic, 1912: 274; Type: MNHN (1 male “ syntype ”); BMNH (2 male “co-types”)</p><p>Xylophilus palliditarsis Pic, 1912: Champion, 1915</p><p>Zarcosia palliditarsis (Pic, 1912): unpublished combination in Grzymala, 2016</p><p>This species and Zarcosia lemairei n. sp. are remarkable among all Zarcosia species discussed in this work by the morphology of the male head (Fig. 5 a–c, e–g). In both cases, the head capsule is carved dorsally and possesses a wavy brush of long yellow bristles. This sexually dimorphic trait is reminiscent of cephalic modification in some Scydmaenus species (Scydmanenidae), while these bristles are similar to the trichomes present in Paussinae Carabidae and other inquilines.</p><p>Zarcosia palliditarsis (Pic, 1912) may very well be a junior synonym of Euglenus planipennis Motschulsky, 1863, also from Sri Lanka. The type material of this species is presumably kept in St. Petersburg or in Moscow and I did not manage to locate it. It is not possible on the sole basis of Motschulsky’s description, 1863 to clarify the taxonomic status of this species.</p><p>Type locality. Sri Lanka: Kandy .</p><p>Distribution. Sri Lanka (Figure 1).</p><p>Material examined. Sri Lanka: Kandy (G. E. Bryant) VI.1908 (1 ♂ and 1 ♀ syntype, MNHN; 2 ♂ co-types, 1 ♂ and 2 ♀, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFC9BB1EFF188011FE08F8ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFC9BB01FF188329FF68FE9C.text	039187D8FFC9BB01FF188329FF68FE9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia patucki (Champion 1924) Gompel 2020	<div><p>Zarcosia patucki (Champion, 1924), comb. nov.</p><p>(Figures 1; 2 ae)</p><p>Xylophilus patucki Champion, 1924: 483; Holotype: BMNH</p><p>Zarcosia patucki (Champion, 1924): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Andhra Pradesh, Chipurupalle .</p><p>Distribution. India: Andhra Pradesh (Figure 1).</p><p>Material examined. India: Andhra Pradesh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=83.566666&amp;materialsCitation.latitude=18.3" title="Search Plazi for locations around (long 83.566666/lat 18.3)">Chipurupalle</a>, 18°18’ N 83°34’ E (R. S. Patuck) (1 ♂ holotype, BMNH)</p></div>	https://treatment.plazi.org/id/039187D8FFC9BB01FF188329FF68FE9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD6BB01FF1885FFFC9BFBE6.text	039187D8FFD6BB01FF1885FFFC9BFBE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia picina (Fairmaire 1893) Gompel 2020	<div><p>Zarcosia picina (Fairmaire, 1893), comb. nov.</p><p>(Figures 1; 2p; 4l)</p><p>Xylophilus picinus, Fairmaire, 1893: 36; Type: MNHN (1 specimen labelled “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.333336&amp;materialsCitation.latitude=10.366667" title="Search Plazi for locations around (long 106.333336/lat 10.366667)">Mt</a> [interpreted as Vietnam, Mỹ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.333336&amp;materialsCitation.latitude=10.366667" title="Search Plazi for locations around (long 106.333336/lat 10.366667)">Tho</a>, 10°22’ N 106°20’ E, a locality referred too multiple times in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.333336&amp;materialsCitation.latitude=10.366667" title="Search Plazi for locations around (long 106.333336/lat 10.366667)">Fairmaire’s</a> original paper]; Museum Paris, Collection Léon Fairmaire 1906 ” read as lectotype, pres. design.)</p><p>Zarcosia picinus [sic!] (Fairmaire, 1893): unpublished combination in Grzymala, 2016</p><p>Hylophilus picinus var. rufescens Pic, 1905, syn. nov.; Type: MNHN</p><p>Type locality. Vietnam: Mỹ Tho .</p><p>Distribution. Singapore, Vietnam (Figure 1).</p><p>This species presents striking morphological similarities with Z. dentatifemur (Pic, 1912), including a longitudinal stripe of pubescence on the pronotum midline and the overall shape of the male genitalia. The shape of the tibiae, however, notably on the forelegs is unambiguously different.</p><p>Material examined. Singapore: (C. J. Saunders) V.1922 (1 ♂, BMNH); (C. J. Saunders) 1.I.1923 (1 ♀, BMNH); Vietnam: Ho Chi Minh City (1 ♂ and 2 sp., MNHN); Ho Chi Minh City (J. Barbier) (5 sp., MNHN; 1 ♂, NGPC, from Pic collection); Mt [interpreted as Vietnam, M ỹ Tho] (1 ♂ lectotype, label added « Hylophilus picinus Fairmaire, 1893, Lectotype ♂, N. Gompel des. 2018 ») .</p></div>	https://treatment.plazi.org/id/039187D8FFD6BB01FF1885FFFC9BFBE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD6BB01FF188020FF63F908.text	039187D8FFD6BB01FF188020FF63F908.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia plumbea (Champion 1916) Gompel 2020	<div><p>Zarcosia plumbea (Champion, 1916), comb. nov.</p><p>(Figures 1; 2d)</p><p>Xylophilus plumbeus Champion, 1916: 31; Holotype: BMNH</p><p>Zarcosia plumbeus [sic!] (Champion, 1916): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Karnataka, Kanara .</p><p>Distribution. India: Karnataka (Figure 1).</p><p>Material examined. India: Karnataka, Kanara (H. L. Andrewes) (1 ♂ holotype, BMNH) .</p><p>Zarcosia proprior (Pic, 1920), comb. nov.</p><p>(Figures 1; 2 ab)</p><p>Hylophilus proprior Pic, 1920: 15; Type: MNHN</p><p>Zarcosia proprior (Pic, 1920): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia: Java, Tji Solak, Winjnkoopsbai .</p><p>Distribution. Republic of Indonesia: Java (Figure 1).</p><p>Material examined. Republic of Indonesia: Java, Tji Solak, Winjnkoopsbai (1 ♀ holotype and 1 ♂ syntype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFD6BB01FF188020FF63F908	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD6BB00FF188303FEA9FE9A.text	039187D8FFD6BB00FF188303FEA9FE9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia rubrobasalis (Pic 1921) Gompel 2020	<div><p>Zarcosia rubrobasalis (Pic, 1921), comb. nov.</p><p>(Figures 1; 2w; 4m)</p><p>Hylophilus rubrobasalis Pic, 1921: 6; Type: MNHN</p><p>Zarcosia rubrobasalis (Pic, 1921): unpublished combination in Grzymala, 2016</p><p>Type locality. Republic of Indonesia, Sumatra .</p><p>Distribution. Republic of Indonesia, Sumatra; Malaysia; Singapore (Figure 1).</p><p>Material examined. Republic of Indonesia: Sumatra (1 type sp., MNHN); Malaysia: Negeri Sembilan Darul Khusus, Port Dickson (H. M. Pendlebury) 4.I.1935 (1 sp., BMNH) ; Singapore: (C. J. Saunders) (1 sp. and 1 ♂, BMNH); (C. J. Saunders) III.1922 (2 sp., BMNH); (C. J. Saunders) VI.1922 (1 ♀, NGPC); (C. J. Saunders) IV.1923 (1 ♂, NGPC) .</p></div>	https://treatment.plazi.org/id/039187D8FFD6BB00FF188303FEA9FE9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD7BB00FF1885FDFBBDFD18.text	039187D8FFD7BB00FF1885FDFBBDFD18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia rufotestacea (Pic 1901) Gompel 2020	<div><p>Zarcosia rufotestacea (Pic, 1901), comb. nov.</p><p>(Figures 1; 2r)</p><p>Hylophilus rufotestaceus Pic, 1901: 96; Type: MNHN</p><p>Zarcosia rufotestaceus [sic!] (Pic, 1901): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Kerala, Malabar, Mahé.</p><p>Distribution. India: Kerala (Figure 1).</p><p>Material examined. India: Kerala, Malabar, Mahé (1 syntype sp., MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFD7BB00FF1885FDFBBDFD18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD7BB00FF188772FDC8F94C.text	039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia schawalleri Gompel 2020	<div><p>Zarcosia schawalleri n. sp.</p><p>(Figures 1; 3k,l; 4n; 5m; 6h)</p><p>Etymology. The species is named after Wolfgang Schawaller, who collected the only known specimens.</p><p>Type locality. Malaysia, Sabah, Sandakan, Sepilok .</p><p>Distribution. Malayan Borneo (Figure 1).</p><p>Description. (Figure 3k,l). Body length. ♂ holotype: 1.63 mm; ♀ paratype: 1.68 mm. Bicolor, head black, pronotum dark-brown and elytra orange with a broad dark transverse area, covered with short double pubescence.</p><p>Head. Transverse, wider than pronotum in both sexes, covered with short white pubescence. Integument black, punctation fine and very dense, rugose. Palpi orange, last article securiform. Eyes much bigger in ♂.</p><p>Antennae. Orange, slender, slightly longer than elytra in ♂, shorter than elytra in ♀, antennomere 3–7 at most twice longer than wide, 8–10 shorter. Covered with hirsute blond pubescence.</p><p>Pronotum (Figure 6h). Narrow, subcylindrical in ♂, trapezoidal in ♀, as long as wide. Lateral constrictions in the posterior half, on each side. Finely and densely punctuated, space between points much smaller than the diameters of points. Integument mat, rugose. Covered with very short, white double pubescence.</p><p>Elytra. Length: ♂ holotype: 1.10 mm; ♀ paratype: 1.15 mm. Orange, with a broad dark area with no defined boundaries, spanning half of the elytral length, from anterior fifth to posterior third. Punctation dense but shallow, covered with double pubescence. Moderately convex, depressed on disc.</p><p>Legs. Gracile. Sexual characters: forelegs with first tarsomere enlarged in male, tibia bent inward (Figure 5m); mesothoracic femora with a sharp apical thorn on their inner side, curved and pointing outward.</p><p>Aedeagus (Figure 4n). Sides regularly convergent in dorsal view, with an apical button. Strongly bent in lateral view with a sudden apical deflection upward.</p><p>Type material. Holotype (♂, NME). Malaysia: Sabah, Sandakan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.933334&amp;materialsCitation.latitude=5.866667" title="Search Plazi for locations around (long 117.933334/lat 5.866667)">Sepilok</a>, 05°52’ N 117°56’ E (50 m) (W. Schawaller) 13.III.2007 . Paratype. Malaysia: Sabah, Sandakan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.933334&amp;materialsCitation.latitude=5.866667" title="Search Plazi for locations around (long 117.933334/lat 5.866667)">Sepilok</a>, 05°52’ N 117°56’ E (50 m) (W. Schawaller) 13.III.2007 (♀ paratype, NME) .</p></div>	https://treatment.plazi.org/id/039187D8FFD7BB00FF188772FDC8F94C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD7BB03FF18834EFA9CFF2B.text	039187D8FFD7BB03FF18834EFA9CFF2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia sellata (Champion 1916) Gompel 2020	<div><p>Zarcosia sellata (Champion, 1916), comb. nov.</p><p>(Figures 1; 2 af)</p><p>Xyophilus sellatus Champion, 1916: 29; Holotype: BMNH</p><p>Zarcosia sellatus [sic!] (Champion, 1916): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Tamil Nadu .</p><p>Distribution. India: Tamil Nadu (Figure 1).</p><p>Material examined. India: Tamil Nadu, Nilgiri Hills (H. L. Andrewes) (1 ♂ holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFD7BB03FF18834EFA9CFF2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD4BB03FF18856BFA8EFDAE.text	039187D8FFD4BB03FF18856BFA8EFDAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia sexdentata (Champion 1924) Gompel 2020	<div><p>Zarcosia sexdentata (Champion, 1924), comb. nov.</p><p>(Figures 1; 2k)</p><p>Xylophilus sexdentatus Champion, 1924: 482; Holotype (male): BMNH</p><p>Zarcosia sexdentatus [sic!] (Champion, 1924): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Andhra Pradesh, Chipurupalle .</p><p>Distribution. India: Andhra Pradesh (Figure 1).</p><p>Material examined. India: Andhra Pradesh, Chipurupalle (R. S. Patuck) (1 ♂ holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFD4BB03FF18856BFA8EFDAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD4BB03FF1886D5FB7EFA29.text	039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia sinuata Gompel 2020	<div><p>Zarcosia sinuata n. sp.</p><p>(Figures 1; 3n,o; 4o; 5n; 6j)</p><p>Etymology. The species’ name refers to the marked bilateral sinuation of the base of its pronotum.</p><p>Type locality. Thailand: Mae Hong Son Province, Pai District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.416664&amp;materialsCitation.latitude=19.116667" title="Search Plazi for locations around (long 98.416664/lat 19.116667)">Mae Sae</a> [19°07’ N 98°25’ E] .</p><p>Distribution. Thailand: Mae Hong Son Province (Figure 1).</p><p>Description. (Figure 3n,o). Body length. ♂ holotype: 2.63 mm; ♀: 2.99± 0.19 mm (n=10). Black, elongated, covered with white double pubescence.</p><p>Head. Moderately transverse, slightly wider than pronotum in both sexes. Punctation coarse and dense. Palpi dark orange, last article securiform.</p><p>Antennae. Moderately long, regularly filiform, somewhat shorter than elytra in ♂, a fourth shorter in ♀. Antennomere 3–8 more than twice longer than wide, the last three much shorter. Covered with long white hairs.</p><p>Pronotum (Figure 6j). Convex, slightly longer than wide in both sexes. Sides parallel from the base to the apical third, then converging markedly inward toward apical margin anteriorly. Densely punctuated, space between points smaller than the diameters of points. Points large and deep. Covered with double pubescence, moderately long white hairs borne from the punctation and pulverulent interspersed silver hairs. Sinuation at base particularly marked.</p><p>Elytra. Length: ♂ holotype: 1.88 mm; ♀: 1.89± 0.09 mm (n=10). Punctation similar to that of the pronotum, with slightly larger points. Covered with double pubescence, as on pronotum. Moderately convex, depressed on disc.</p><p>Legs. Rather long and thin. No sexual characters.</p><p>Aedeagus (Figure 4o). Sides regularly convergent in dorsal view, rounded at apex, with an apical button. Moderately bent in lateral view with a weak apical deflection upward.</p><p>Type material. Holotype (♂, SMNS). Thailand: Mae Hong Son Province, Pai District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.416664&amp;materialsCitation.latitude=19.116667" title="Search Plazi for locations around (long 98.416664/lat 19.116667)">Mae Sae</a>, 19°07’ N 98°25’ E (750 m), (L. Bocák) 15.V.1993 . Paratypes. Thailand: Mae Hong Son Province, Pai District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.416664&amp;materialsCitation.latitude=19.116667" title="Search Plazi for locations around (long 98.416664/lat 19.116667)">Mae Sae</a>, 19°07’ N 98°25’ E (750 m), (L. Bocák) 15.V.1993 (9 ♀ paratypes, SMNS; 2 ♀ paratypes, NGPC) .</p></div>	https://treatment.plazi.org/id/039187D8FFD4BB03FF1886D5FB7EFA29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD4BB05FF18826EFC5AFBCA.text	039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia spinifemur Gompel 2020	<div><p>Zarcosia spinifemur n. sp.</p><p>(Figures 1; 3p; 4s; 5x; 6n)</p><p>Etymology. The species’ name refers to the acute teeth that ornate the inner side of male femora.</p><p>Type locality. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary [07°13’ N 80°57’ E] .</p><p>Distribution. Sri Lanka: Central, Kandy district (Figure 1).</p><p>Description. (Figure 3p). Female unknown. Body length. ♂ holotype: 2.23 mm. Bicolor with dark head, red pronotum and red elytra marked with a broad dark area. Covered with short and dense double pubescence.</p><p>Head. Transverse, wider than pronotum, covered with white pubescence. Finely and densely punctuated. Palpi orange, last article securiform.</p><p>Antennae. Long and strong, a third longer than elytra, with antennomere 4–8 at least twice longer than wide, conical. Covered with hirsute white pubescence.</p><p>Pronotum (Figure 6n). Dark red. Trapezoidal with divergent sides from base to apex, strongly convex. Sides straight. Densely punctuated, space between points smaller than the diameters of points. Covered with short double pubescence. Base sinuated with a dense fringe of golden pubescence anterior to the scutellum, half the length of the base and centered on the dorsal midline.</p><p>Elytra. Length: ♂ holotype: 1.50 mm. Dark red with a broad dark transversal area from excluding the basal third and the apical fifth of the elytra. Densely punctuated, but not as densely as on the pronotum. Space between points as large as the points themselves. Covered with double pubescence. Regularly convex.</p><p>Legs. Long and thin. Prothoracic and mesothoracic legs entirely orange, metathoracic legs dark, except base of femora dark-orange. Sexual characters: mesothoracic femora with an acute apical thorn, curved and pointing outward (Figure 5x).</p><p>Aedeagus (Figure 4s). Sturdy, sides regularly convex in dorsal view, ending in an apical triangle. Moderately bent near its base in lateral view, with a weak apical upward deflection.</p><p>Type material. Holotype (♂, UCDC). Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999.</p></div>	https://treatment.plazi.org/id/039187D8FFD4BB05FF18826EFC5AFBCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD2BB07FF1880CDFC9CFB18.text	039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia srilankaensis Gompel 2020	<div><p>Zarcosia srilankaensis n. sp.</p><p>(Figures 1; 3r; 4u; 5h,p; 6m)</p><p>Etymology. The species’ name refers to the geographical origin of the holotype.</p><p>Type locality. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary [07°13’ N 80°57’ E] .</p><p>Distribution. Sri Lanka: Central, Kandy district (Figure 1).</p><p>Description. (Figure 3t). Female unknown. Body length. ♂: 2.92± 0.16 mm (n=12). Robust, sturdy, uniformly reddish brown, covered with double pubescence.</p><p>Head. Transverse, barely wider than pronotum, covered with blond pubescence, pubescence denser between antenna insertion points. Punctation coarse and very dense. Palpi dark orange, last article securiform.</p><p>Antennae. Strong, shorter than elytra, antennomere 3–10 short, at most twice longer than wide, conical. Covered with hirsute blond pubescence.</p><p>Pronotum (Figure 6m). Longer than wide, disc convex. Sides subparallel from base to the apical third, then converging markedly inward toward apical margin anteriorly. Densely punctuated, space between points smaller than the diameters of points. Covered with double pubescence, short blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a light fringe of golden pubescence anterior to the scutellum.</p><p>Elytra. Length: ♂: 2.03± 0.12 mm (n=12). Densely punctuated, space between points larger than the points themselves. Points larger and shallower than those of the pronotum. Covered with double pubescence. Moderately convex, depressed on disc. Apex with a tuft of long hairs along the suture (Figure 5h).</p><p>Legs. Short, sturdy. Sexual characters: foretibia with 2 large sharp teeth, one half the inner side and one apically, together carving a wrench in the tibia (Figure 5p). Mesothoracic femora with an minute apical tooth, not visible in dorsal view.</p><p>Aedeagus (Figure 4u). Sturdy, sides subparallel in dorsal view, progressively convergent in the apical fourth, truncated apically. Moderately bent in lateral view, apex not deflected upward.</p><p>Type material. Holotype (♂, UCDC). Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 . Paratypes. Sri Lanka: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2166667" title="Search Plazi for locations around (long 80.95/lat 7.2166667)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°13’ N 80°57’ E (445 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (6 ♂ paratypes, UCDC; 4 ♂ paratypes, NGPC), 30.VIII.1999 (1 ♂ paratype, NGPC) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.95&amp;materialsCitation.latitude=7.2" title="Search Plazi for locations around (long 80.95/lat 7.2)">Central</a>, Kandy district, Victoria Rendenigala Ramtembe Sanctuary, 07°12‘ N 80°57‘ E (135 m) (M. &amp; J. Wasbauer) 23.VIII.1999 (1 ♂ paratype, UCDC) .</p><p>Note. This species was collected together with Z. dentatifemur (Pic, 1912), Z. spinifemur n. sp., Z. brunnea n. sp. and Z. intermedia n. sp. The sample contained a large series of females, which I could not assign to the respective species. In particular, Z. brunnea n. sp., Z. srilankaensis n. sp. and Z. intermedia n. sp. form a very homogeneous group. Their males are easily distinguished by stable sexual characters (aedeagus, teeth on the tibiae, hooks on the femora), but in spite of extensive comparisons of the females collected with them, I could not decide whether these females represented one or more taxa, nor which.</p></div>	https://treatment.plazi.org/id/039187D8FFD2BB07FF1880CDFC9CFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD0BB06FF188173FA8EFE9A.text	039187D8FFD0BB06FF188173FA8EFE9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia subglaber , Pic 1952	<div><p>Zarcosia subglaber Pic, 1952</p><p>(Figures 1; 2y)</p><p>Zarcosia subglaber Pic, 1952: 5; Lectotype, pres. design.: one female specimen labelled « Zarcosia inglaber [sic!] n. sp. », MNHN</p><p>Zarcosia subglaber Pic, 1952: Grzymala, 2016</p><p>Type locality. Vietnam: Ho Chi Minh City .</p><p>Distribution. Vietnam: Ho Chi Minh City (Figure 1).</p><p>Note on type material: There is no specimen of Zarcosia labelled subglaber from Pic’s collection at the MNHN. There is, however, a single specimen of Zarcosia with multiple labels « Zarcosia inglaber n. sp. » from Pic’s handwritting. This specimen also carries a locality label « Saigon [= Ho Chi Minh city], 5 VI 1950, J. Barbier ». Given that no other Aderidae to date has a species name with the radical *glaber*, it is likely that the name inglaber is a lapsus calami, and that Pic meant subglaber . This hypothesis is reinforced by the original description of subglaber, for 3 reasons. First, the only Zarcosia that Pic described under this genus (rather than under Hylophilus or Xylophilus) is Zarcosia subglaber . Among all the Zarcosia in Pic’s collection, a single specimen is labelled « Zarcosia » with Pic’s handwriting, the very specimen mentioned above and labeled « Zarcosia inglaber n. sp. ». Second, in his description of Zarcosia subglaber, Pic explicitly indicates that the material was given to him by Jean Barbier, who collected it in Saigon. Finally. Pic writes in his paper « Voisin de lateniger Pic », referring to the similarity between Zarcosia subglaber and « lateniger ». Although I could not find out what « lateniger Pic» is, the specimen labeled « Zarcosia inglaber n. sp. » also carries a label that reads « sp près lateniger » from Pic’s handwriting. Together, these elements unambiguously support that the specimen labeled « Zarcosia inglaber n. sp. » at MNHN was used by Pic for the description of Zarcosia subglaber Pic, 1952 . I design it here as a lectotype and have added a label « Zarcosia subglaber Pic, 1952, Lectotype ♀, N. Gompel des. 2020 ».</p><p>Material examined. Vietnam: Ho Chi Minh City (J. Barbier) 5.VI.1950 (1 ♀ lectotype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFD0BB06FF188173FA8EFE9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD1BB06FF1885FDFF63FCEA.text	039187D8FFD1BB06FF1885FDFF63FCEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia subrobusta (Pic 1943) Gompel 2020	<div><p>Zarcosia subrobusta (Pic, 1943), comb. nov.</p><p>(Figures 1; 2 ac)</p><p>Hylophilus subrobustus Pic, 1943: 5; Type: MNHN</p><p>Zarcosia subrobustus [sic!] (Pic, 1943): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Tamil Nadu, Coimbatore .</p><p>Distribution. India: Tamil Nadu (Figure 1).</p><p>Material examined. India: Tamil Nadu, Coimbatore, On edge of field (P. S. Nathan) VIII.1932 (2 syntypes, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFD1BB06FF1885FDFF63FCEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD1BB06FF18872DFA8EFB5E.text	039187D8FFD1BB06FF18872DFA8EFB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia subrufa (Champion 1924) Gompel 2020	<div><p>Zarcosia subrufa (Champion, 1924), comb. nov.</p><p>(Figures 1; 2 aj)</p><p>Xylophilus subrufus Champion, 1924: 490; Type: BMNH</p><p>Zarcosia subrufus [sic!] (Champion, 1924): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Andhra Pradesh, Chipurupalle .</p><p>Distribution. India: Andhra Pradesh (Figure 1).</p><p>Material examined. India: Andhra Pradesh, Chipurupalle (R. S. Patuck) (1 ♀ holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFD1BB06FF18872DFA8EFB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFD1BB09FF1880B9FAEFFE76.text	039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia sumatrensis Gompel 2020	<div><p>Zarcosia sumatrensis n. sp.</p><p>(Figures 1; 3g,h; 4p; 5o; 6f)</p><p>Etymology. The species’ name refers to its geographical origin.</p><p>Type locality. Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.166664&amp;materialsCitation.latitude=4.233333" title="Search Plazi for locations around (long 98.166664/lat 4.233333)">Goea Limau Moangkaer</a> [04°14’ N 98°10’ E] .</p><p>Distribution. Republic of Indonesia: Sumatra (Figure 1).</p><p>Description. (Figure 3g,h). Body length. ♂ holotype: 3.00 mm; ♀: 2.65± 0.19 mm (n=7). Dark brown, darker anteriorly, gracile, covered with very short, pulverulent pubescence. Appendages orange-brown.</p><p>Head. Small, dark, barely wider than pronotum. Palpi orange, last article securiform. Integument, finely and densely punctuated, lightly rugose. Occiput lined with a very fine fringe of dense and short silver hairs posteriorly.</p><p>Antennae. A third longer than elytra in ♂, as long as elytra in ♀, orange-brown, regularly filiform with antenommeres 3–10 cylindrical.</p><p>Pronotum (Figure 6f). Round and convex, slightly elongated in ♂, as long as wide in ♀. Sides regularly curved. Surface mat, finely and densely punctuated, covered with dense, pruinose double pubescence.</p><p>Elytra. Length: ♂ holotype: 2.00 mm; ♀: 1.95± 0.10 mm (n=7). Punctation fine and shallow, dense (space between points as wide as the points themselves). Disc depressed. Pubescence dense and short, whitish, comprised of short hairs borne from the points and interspersed pulverulent hairs.</p><p>Legs. Particularly elongated. Foretibia thickened in a smooth tooth on their inner side (Figure 5o). Mesothoracic legs not observed (a single male devoid of mesothoracic legs was present in the type series).</p><p>Aedeagus (Figure 4p). Sides subparallel in dorsal view, apex tapered in apical fifth, ending with a button. Strongly bent in lateral view, apex with a weak upward deflection. Appendages visible near apex.</p><p>Type material. Holotype (♂, BMNH). Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.166664&amp;materialsCitation.latitude=4.233333" title="Search Plazi for locations around (long 98.166664/lat 4.233333)">Goea Limau Moangkaer</a>, 04°14’ N 98°10’ E (J. C. van der Meer Mohr) 20.III.1932 . Paratypes. Republic of Indonesia: Sumatra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.166664&amp;materialsCitation.latitude=4.233333" title="Search Plazi for locations around (long 98.166664/lat 4.233333)">Goea Limau Moangkaer</a>, 04°14’ N 98°10’ E (J. C. van der Meer Mohr) 20.III.1932 (2 ♀ paratypes, BMNH) ; Sumatra, Goea Lau Rakit I, 02°20’ N 98°40’ E (J. C. van der Meer Mohr) 5.X.1934 (6 ♀ paratypes, BMNH; 2 ♀ paratypes, NGPC) .</p><p>Note. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.</p></div>	https://treatment.plazi.org/id/039187D8FFD1BB09FF1880B9FAEFFE76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFDEBB09FF188591FBFEFD0E.text	039187D8FFDEBB09FF188591FBFEFD0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia testaceitarsis (Pic 1926) Gompel 2020	<div><p>Zarcosia testaceitarsis (Pic, 1926), comb. nov.</p><p>(Figures 1; 2 ah)</p><p>Hylophilus testaceitarsis Pic, 1926: 13; Holotype: MNHN</p><p>Type locality. Vietnam: Lạc Thô (Tonkin, Lactho) .</p><p>Distribution. Vietnam (Figure 1).</p><p>Material examined. Vietnam: Lạc Thô (A. De Cooman) (1 ♂ holotype, MNHN) .</p></div>	https://treatment.plazi.org/id/039187D8FFDEBB09FF188591FBFEFD0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFDEBB09FF188709FF61FB16.text	039187D8FFDEBB09FF188709FF61FB16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia troglodytes (Champion 1916) Gompel 2020	<div><p>Zarcosia troglodytes (Champion, 1916), comb. nov.</p><p>(Figures 1; 2a,b; 4q)</p><p>Xylophilus troglodytes Champion, 1916: 30; Holotype: BMNH</p><p>Euglenes troglodytes (Champion, 1916): Werner, 1962</p><p>Zarcosia troglodytes (Champion, 1916): unpublished combination in Grzymala, 2016</p><p>Type locality. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves .</p><p>Distribution. Malaya, Batu caves (Figure 1).</p><p>Material examined. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves (H. N. Ridley) (1 ♂ holotype and 1 ♀, BMNH) ; Selangor Darul Ehsan, Gombak District, Batu caves (C. Dover) VII.1926 (2 ♂ and 1 ♀, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFDEBB09FF188709FF61FB16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFDEBB09FF188171FB88F98A.text	039187D8FFDEBB09FF188171FB88F98A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia uncifer (Champion 1916) Gompel 2020	<div><p>Zarcosia uncifer (Champion, 1916), comb. nov.</p><p>(Figures 1; 2 ak)</p><p>Xylophilus uncifer Champion, 1916: 32; Holotype (male): BMNH</p><p>Zarcosia uncifer (Champion, 1916): unpublished combination in Grzymala, 2016</p><p>Type locality. India: Maharashtra, Mumbai .</p><p>Distribution. India: Maharashtra (Figure 1).</p><p>Material examined. India: Maharashtra, Mumbai (1 ♂ holotype, BMNH) .</p></div>	https://treatment.plazi.org/id/039187D8FFDEBB09FF188171FB88F98A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
039187D8FFDCBB0AFF1884DDFD43FD56.text	039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zarcosia weigeli Gompel 2020	<div><p>Zarcosia weigeli n. sp.</p><p>(Figures 1; 3e; 4r; 5i,t; 6d)</p><p>Etymology. The species is named after Andreas Weigel in recognition of her efforts to collect material for others.</p><p>Type locality. China: Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.666664&amp;materialsCitation.latitude=22.15" title="Search Plazi for locations around (long 100.666664/lat 22.15)">Na Ban</a> [22°09’ N 100°40’ E] .</p><p>Distribution. China: Yunnan.</p><p>Description. (Figure 3i,j). Female unknown. Body length. ♂: 2.53± 0.04 mm (n=2). Entirely black, elongated, covered with thin blond pubescence.</p><p>Head. Small, weakly transverse, slightly wider than pronotum. Palpi dark brown, last article securiform. Punctation coarse, dense. Posterior margin lined with a fringe of dense white pubescence.</p><p>Antennae. Subfiliform, slightly longer than elytra, black. Antennomere 3–8 more than twice as long as wide. Covered with hirsute white pubescence.</p><p>Pronotum (Figure 6d). Subcylindrical, convex, longer than wide, sides weakly but regularly convex. Strongly and densely punctuated, points deep.</p><p>Elytra. Length: ♂: 1.75±0.00 mm (n=2). Punctation stronger than on pronotum, dense, points deep, well defined. Covered with double pubescence: moderately long hair borne from punctation and interspersed whitish pulverulent hairs (Figure 5i). Moderately convex, depressed on disc.</p><p>Legs. Long and thin, black, except last two tarsomeres, yellow. Sexual characters. Male foretibia weakly bent inward, with a small acute tooth on their inner side (Figure 5t).</p><p>Aedeagus (Figure 4r). Simple, sturdy, sides parallel in dorsal view, progressively tapered in a regular triangle in apical fourth. Hardly bent in lateral view, with a weak apical upward deflection.</p><p>Type material. Holotype (♂, NME). China: Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. Na Ban (NNNR), 22°09’ N 100°40’ E (730 m), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.666664&amp;materialsCitation.latitude=22.15" title="Search Plazi for locations around (long 100.666664/lat 22.15)">Transition</a> zone MF (A. Weigel) 23.V.2008 . Paratype. China: Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.666664&amp;materialsCitation.latitude=22.15" title="Search Plazi for locations around (long 100.666664/lat 22.15)">Na Ban</a> (NNNR), 22°09’ N 100°40’ E (730 m) , secondary forest, Rubber plant (A. Weigel) 20.V.2008 (1 ♂ paratype, NGPC) .</p></div>	https://treatment.plazi.org/id/039187D8FFDCBB0AFF1884DDFD43FD56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Gompel, Nicolas	Gompel, Nicolas (2020): Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species. Zootaxa 4838 (2): 151-178, DOI: 10.11646/zootaxa.4838.2.1
