taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
039187D8FFC4BB13FF188075FF61FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC4BB13FF188075FF61FA6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC4BB12FF1881AEFD9CFDE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC4BB12FF1881AEFD9CFDE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC4BB12FF1881AEFD9CFDE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC4BB12FF1881AEFD9CFDE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC5BB12FF18821AFAA7F837.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC5BB12FF18821AFAA7F837.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1884DDFAEFFC59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1884DDFAEFFC59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1884DDFAEFFC59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1884DDFAEFFC59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1887BDFE62FA96.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1887BDFE62FA96.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB15FF1887BDFE62FA96.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC2BB14FF1881F1FC9CFC7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF188799FB8BFAFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF188799FB8BFAFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF18811EFE35F880.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF18811EFE35F880.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF18811EFE35F880.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC3BB14FF18811EFE35F880.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1884DDFAEFFBA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF1880E1FEAAF9DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF18823EFF63F837.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC0BB17FF18823EFF63F837.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF1884DDFF63FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF1884DDFF63FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF188676FBBFFA01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF18821AFEF1F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC1BB16FF18821AFEF1F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1BFF188709FAC9FB4F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1BFF188709FAC9FB4F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1BFF18814BFE17F9AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1BFF18814BFE17F9AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCCBB1AFF1882EDFAEFFD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCDBB1DFF1886B1FC9CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1DFF18856DFB11FD8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1DFF18856DFB11FD8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1DFF18868DFCF6FBE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1DFF18868DFCF6FBE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1DFF18868DFCF6FBE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1CFF188026FF63FBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1CFF188026FF63FBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1CFF188026FF63FBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCABB1CFF188026FF63FBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCBBB1CFF1880CDFE7FFA2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCBBB1CFF1880CDFE7FFA2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCBBB1CFF18826DFDFBF866.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFCBBB1CFF18826DFDFBF866.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF1884DDFD07FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF1884DDFD07FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF188676FBA8FC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF188676FBA8FC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF1887A8FF63FAC9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF1887A8FF63FAC9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF188129FF68F924.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1FFF188129FF68F924.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1EFF188366FBD5FBF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC8BB1EFF188366FBD5FBF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB1EFF188011FE08F8ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB1EFF188011FE08F8ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB1EFF188011FE08F8ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB1EFF188011FE08F8ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB01FF188329FF68FE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFC9BB01FF188329FF68FE9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB01FF1885FFFC9BFBE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB01FF1885FFFC9BFBE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB01FF1885FFFC9BFBE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB01FF188020FF63F908.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB01FF188020FF63F908.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB00FF188303FEA9FE9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB00FF188303FEA9FE9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD6BB00FF188303FEA9FE9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF1885FDFBBDFD18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF1885FDFBBDFD18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB00FF188772FDC8F94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB03FF18834EFA9CFF2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD7BB03FF18834EFA9CFF2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF18856BFA8EFDAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF18856BFA8EFDAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB03FF1886D5FB7EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD4BB05FF18826EFC5AFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD2BB07FF1880CDFC9CFB18.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD0BB06FF188173FA8EFE9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD0BB06FF188173FA8EFE9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB06FF1885FDFF63FCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB06FF1885FDFF63FCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB06FF18872DFA8EFB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB06FF18872DFA8EFB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFD1BB09FF1880B9FAEFFE76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188591FBFEFD0E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188591FBFEFD0E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188709FF61FB16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188709FF61FB16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188709FF61FB16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188171FB88F98A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDEBB09FF188171FB88F98A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403789/files/figure.png	https://doi.org/10.5281/zenodo.4403789	FIGURE 2. continued next page.	FIGURE 2. continued next page.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403787/files/figure.png	https://doi.org/10.5281/zenodo.4403787	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453388/files/figure.png	https://doi.org/10.5281/zenodo.4453388	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4453392/files/figure.png	https://doi.org/10.5281/zenodo.4453392	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403791/files/figure.png	https://doi.org/10.5281/zenodo.4403791	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
039187D8FFDCBB0AFF1884DDFD43FD56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4403795/files/figure.png	https://doi.org/10.5281/zenodo.4403795	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm.	2020-08-26	Gompel, Nicolas		Zenodo	biologists	Gompel, Nicolas			
