taxonID	type	description	language	source
0390E827603CFF9D95D1F8A247F4F99C.taxon	type_taxon	Type species: A. omayan Huber, 2005.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	diagnosis	Diagnosis Even though the cladistic analysis identifies only two synapomorphies for Aetana, the genus is fairly easily distinguished from the putatively closest relatives by the following characters: retrolateral trichobothrium on leg 1 very proximal (at <5 % of tibia length) and presence of curved hairs on tibiae and / or metatarsi (both in contrast to Spermophora, Khorata Huber, 2005, Savarna, and an undescribed genus from Sarawak, below called ‘ Gen. n. Borneo’); sternum not dark, retrolateral trichobothrium on male palpal tibia in very distal position, and presence of epiandrous spigots (all in contrast to Khorata, Savarna, and ‘ Gen. n. Borneo’); male legs without spines and male palpal coxa unmodified (both in contrast to ‘ Gen. n. Borneo’); ALS with only two spigots, epigynal plate without external pair of pockets, and female genitalia without unpaired posterior pocket (all in contrast to Spermophora). Most characters previously thought to be diagnostic (Huber 2005 a) are rendered invalid due to the newly described species.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	description	Description Male MEASUREMENTS. Total body length ~ 2.5 – 4.5 (smallest species in A. kinabalu group; largest species in A. omayan group); carapace width 0.9 – 1.8; leg 1 length ~ 27 – 44; tibia 1 length ~ 6.0 – 11.0; tibia 2 / tibia 4 length 0.92 – 1.08; tibia 1 L / d ~ 55 – 95 (the largest species, A. omayan, has the relatively thickest legs; the smallest species, A. gaya, has the relatively thinnest legs). COLOR. In life mostly ochre-gray with brown and black marks (e. g., Figs 51 – 56, 105 – 110), only A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov. with light brown to orange prosoma and palps (Figs 8 – 12); sternum never dark; legs usually with indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); darker rings missing in A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov. BODY. Carapace either with shallow median furrow restricted to frontal part (Figs 63 – 64, 198) or without furrow (Figs 118, 130); ocular area raised, eye triads on short stalks directed toward lateral, in some species with median process (Figs 77, 96) or with pair of processes arising from near ALE (Figs 221, 226). AME absent. Clypeus high, either unmodified, with small paired processes (Fig. 191), with large median process (Figs 59, 77), or with indistinct lateral ridges (Figs 221, 226). Abdomen from slightly longer than high (Figs 51 – 56) to almost cylindrical (Figs 8, 102), pointed at spinnerets. Male gonopore with four epiandrous spigots in all species examined with SEM (e. g., Figs 37, 143, 162); each ALS with large widened spigot and pointed spigot, without further cylindrically shaped spigots (Figs 68, 210); PMS with two spigots each. CHELICERAE. Very variable, distal apophyses ranging from short processes in frontal position (Figs 15, 40) to long processes in lateral position (Figs 191, 226), absent in A. kiukoki group; proximal apophyses usually present, large in A. kiukoki group (Fig. 66), absent in A. ocampoi group (Figs 15, 40); chelicerae without modified hairs; without stridulatory ridges. PALPS. Coxa unmodified; trochanter usually with one retrolateral to ventral process, sometimes provided with scales or teeth (Fig. 158), in A. ocampoi group fused to femur (Figs 14, 27); trochanter in some species with additional prolateral apophysis (Fig. 189); femur rarely simple (Fig. 27), usually with one or more processes, in A. kinabalu group with complex set of up to four processes (e. g., Figs 150 – 151); patella either triangular in lateral view or ventral side longer than usual (e. g., Figs 14, 58); tibia either of usual shape (e. g., Figs 14, 127) or rather small and slender (A. kiukoki group; e. g., Fig. 58), with retrolateral trichobothrium in very distal position (close to tibia-tarsus joint; Figs 27, 114); palpal tarsus small, usually with capsular tarsal organ (Figs 134, 203), exposed in A. libjo Huber, sp. nov. (Fig. 35) (not clear in A. ocampoi Huber, sp. nov. and A. baganihan Huber, sp. nov.); procursus usually complex with proximal and distal parts connected by membranous hinge; procursus in A. kiukoki group unusually long (Figs 58, 76), in A. ocampoi group reduced to simple semi-transparent process (Figs 14, 27); bulb either with weakly sclerotized embolus as only process (Figs 114, 159), in A. kiukoki group with additional short membranous process (Figs 58, 75), in A. ocampoi group with additional novel processes (Figs 13 – 14, 26 – 27). LEGS. Without spines; with curved hairs on tibiae and / or metatarsi (Fig. 133); usually without or with few vertical hairs, only in A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov. with one dense row retrolatero-dorsally on each tibia; retrolateral trichobothrium on tibia 1 very proximal (at 2 – 4 % of tibia length), prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 usually with ~ 25 – 30 pseudosegments, fairly distinct distally; tarsus 4 with single row of ventral comb-hairs of a modified Belisana - type (cf. Huber & Fleckenstein 2008; Figs 32, 69, 82, 209). Female Similar to male but eye triads on lower humps (Figs 64, 80, 119) and closer together than in male (sexual dimorphism low in A. ocampoi group; in other groups, eye triads in females often less than half as wide apart as in males); clypeus and chelicerae unmodified; legs slightly shorter than in males (tibia 1 ~ 4.0 – 9.0). Females in A. omayan group with stridulatory apparatus between prosoma (elongate median plate on carapace posteriorly; Figs 186 – 188) against abdomen (indistinct hairless area frontally). Epigynum weakly to heavily sclerotized, sometimes with scape of variable length and shape (Figs 28, 60), never with external pair of pockets in anterior epigynal plate but in A. kinabalu and A. omayan groups sometimes with internal sclerotized pockets originating from ventral wall of uterus externus (Figs 111, 192 – 194), and in A. omayan group with pair of membranous external pockets in posterior epigynal area (Figs 192, 194, 222). Internal genitalia with pair of pore plates, in A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov. with unique median membranous structure (Figs 29, 44); in A. kiukoki group with distinctive serrated ridges (Figs 61, 73, 79).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	discussion	Monophyly and relationships Morphologically, the monophyly of Aetana appears weakly supported. Only two characters support this node (Fig. 1), both of them without homoplasy within the taxa included in the matrix but with considerable homoplasy among more distant Pholcinae relatives: (1) the very proximal position of the retrolateral trichobothrium on the leg tibiae (char. 30); and (2) the presence of curved hairs on leg tibiae and / or metatarsi (char. 31). However, preliminary analyses of molecular data (including many more potential close relatives) (A. Valdez-Mondragón, D. Dimitrov, B. A. Huber, unpubl. data) consistently support the monophyly of Aetana with high support values. The sister group of Aetana appears much better supported by morphology. Four characters suggest that three genera together (Khorata, Savarna, and ‘ Gen. n. Borneo’) are sister to Aetana. This, however, is in conflict with our preliminary molecular data that suggest a closer relationship between Aetana and East Asian Spermophora (incl. S. estebani) than between Aetana and Khorata, Savarna, and ‘ Gen. n. Borneo’. Such a close relationship with East Asian Spermophora was actually proposed in the original description (Huber 2005 a), at that time without a formal cladistic analysis. Within Aetana, four species groups receive strong support both from the morphological analysis herein (Fig. 1) and from our preliminary molecular data. Conflict exists regarding relationships among these groups. The present analysis supports a sister group relationship between the Aetana kinabalu and A. omayan groups, and at least one of the three characters supporting this node is a unique feature (the long retrolateral membranous process on the procursus; char. 21). Our preliminary molecular data do not strongly support any sister group relationships among the four species groups within Aetana.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	biology_ecology	Natural history Most species were collected in forests, from well protected spaces close to the ground, under rocks and logs, in small holes and cavities. Few species occur higher in the vegetation, even in places directly reached by the sun, most notably the closely related A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov. and the putatively close relatives A. kinabalu and A. lambir Huber, sp. nov. An exception is the type species A. omayan, which was mainly collected in a cave but also among rocks in a treeless ravine. All species seem to build simple domed sheet webs (like most other pholcids studied), but the sheets are unusually strongly domed in A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov., and in A. kinabalu and A. lambir Huber, sp. nov. a second sheet occurs a few cm under the main sheet. In six cases, two species were found to share a locality, sometimes in different microhabitats (e. g., at Baganihan: A. kiukoki Huber, sp. nov. near the ground; A. baganihan Huber, sp. nov. among vegetation), sometimes in what seemed to be identical microhabitats (e. g., at Mt. Banahaw: A. manansalai Huber, sp. nov. and A. banahaw Huber, sp. nov., both close to the ground). In most species, males and females were often found together sharing a web. This was never observed in A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov., where males and females were sometimes observed very close to each other but in separate webs. When disturbed, Aetana spiders tend to run toward the periphery of the web rather than to vibrate in the ‘ typical’ pholcid way. Some do then vibrate vigorously for a very short time before becoming motionless and pressing their body against the substrate; others stop vibrating and start to gently move the abdomen in circles (A. libjo Huber, sp. nov. and A. baganihan Huber, sp. nov.). The closely related A. poring Huber, sp. nov. and A. indah Huber, sp. nov. barely reacted to disturbance.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	discussion	Composition The genus now includes 18 described species. The high species turnover in the Philippines and in northern Borneo, together with the wide distribution of one species group (A. omayan group; Philippines to Fiji Islands) and the huge sampling gaps in eastern Indonesia, New Guinea, and east to Fiji suggest that at least several dozen further species are likely to exist.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827603CFF9D95D1F8A247F4F99C.taxon	distribution	Distribution Ranging from northern Borneo and the Philippines to Fiji (Fig. 2). Most gaps and missing records are probably due to lack of adequate sampling, but the absence of records from two areas might reflect real absence. First, our intensive collecting in western Sarawak (at seven localities ranging from the Pueh foothills in the west to Niah in the east) did not result in a single specimen of Aetana, while many specimens were collected at all eight localities in eastern Sarawak (east of Niah) and Sabah. Second, northern Australia is relatively well sampled, but a revision of all the material available in collections (Huber 2001) did not reveal any Aetana.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	description	urn: lsid: zoobank. org: act: 85 B 2 FF 02 - FB 08 - 48 B 3 - 91 E 9 - 0 AB 54 D 7866 F 5 Figs 6 – 7, 13 – 25	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	diagnosis	Diagnosis Easily distinguished from closest known relatives (A. libjo Huber, sp. nov.; A. baganihan Huber, sp. nov.) by dark coloration (Figs 6 – 7) and by several details of the male palp (Figs 13 – 14; long cylindrical bulb; ventral process on femur), male chelicerae (Fig. 15; more proximal position of apophyses), and epigynum (Figs 16 – 19; short triangular scape).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	etymology	Etymology Named for the Filipino artist Hernando Ruiz Ocampo (1911 – 1978), famous for his work reflecting the harsh realities of his country after the Second World War, but also for his interest in depicting Philippine flora and fauna.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Luzon, Camarines Sur Prov., Mt. Isarog, W slope (13.664 ° N, 123.34 – 123.35 ° E), ~ 600 – 900 m a. s. l., forest, near ground, 23 Feb. 2014 (B. A. Huber), ZFMK (Ar 13927). Other material PHILIPPINES: Luzon, 1 ♂, 2 ♀♀ (morph A and morph B), same data as holotype, ZFMK (Ar 13928); 3 ♀♀ (2 ♀♀ morph A, 1 ♀ morph B), in pure ethanol, same data, ZFMK (Phi 222).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.7, carapace width 1.1. Leg 1: 28.3 (6.7 + 0.4 + 6.6 + 11.8 + 2.8), tibia 2: 3.9, tibia 3: 2.7, tibia 4: 4.0; tibia 1 L / d: 59. Distance PME-PME 185 µm, diameter PME 125 × 120 µm, distance PME-ALE 20 µm; AME absent. COLOR. Carapace pale ochre with wide median dark band including ocular area and narrower lateral black margins; clypeus pale ochre, distal half light brown; sternum pale ochre, medially light brown; legs light brown, with dark rings subdistally on femora and proximally on tibiae (incl. patellae); abdomen dorsally densely covered with dark marks and some white marks, ventrally gray, small dark mark behind gonopore, larger dark mark in front of spinnerets. BODY. Habitus as in Figs 6 – 7; ocular area slightly raised, each triad on low hump; carapace only anteriorly with very shallow and narrow median furrow (rather just a dark line); clypeus unmodified; sternum wider than long (0.60 / 0.55), unmodified. CHELICERAE. As in Fig. 15, with pair of frontal apophyses near median line, without proximal lateral apophyses; without modified hairs; without stridulatory ridges. PALPS. As in Figs 13 – 14, coxa unmodified, trochanter on retrolatero-ventral side with large apophysis fused to femur; femur with large retrolatero-ventral process; retrolateral trichobothrium on tibia very distal; tarsus with semitransparent simple procursus; genital bulb large, cylindrical, with retrolateral apophysis proximally and two distinctive processes distally: dorsal process with sclerotized black tip; ventral process with semitransparent flap. LEGS. Without spines, with curved hairs on tibiae 1 and metatarsi 1 – 4, few vertical hairs; retrolateral trichobothrium on tibia 1 at 3 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 25 pseudosegments, distally distinct. Male (variation) Tibia 1 in other male: 7.2; this male paler but otherwise identical. Female In general similar to male; eye triads closer together (distance PME-PME 140 µm); tibia 1 in 2 females: 5.5, 5.7. Epigynum apparently dimorphic: three females with small triangular sclerotized scape (Figs 16, 20; morph A), two females with much wider sclerotized area of epigynum (Figs 18, 23; morph B); internal genitalia also different, as in Figs 17, 19, 22, 25.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	biology_ecology	Natural history The spiders were found close to the ground, apparently more deeply hidden under rocks and logs than the sympatric A. lozadae Huber, sp. nov.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276037FF989598F9DE47FCFD7F.taxon	distribution	Distribution Known from type locality only (Fig. 3).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	description	urn: lsid: zoobank. org: act: 91 DA 8483 - 221 E- 4 FBB- 8 CD 8 - 453 CFD 23 B 90 F Figs 8 – 9, 26 – 40, 45 – 47	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	diagnosis	Diagnosis Distinguished from the very similar A. baganihan Huber, sp. nov. by shapes of bulbal processes (compare Figs 38 – 39 and 41 – 42), by shorter male palpal tibia (2.1 – 2.4 × longer than wide vs. 2.9 – 3.2 in A. baganihan Huber, sp. nov.), and by shorter epigynum and wider scape (compare Figs 28 and 43). Distinguished from next closest known relative (A. ocampoi Huber, sp. nov.) by light coloration and several details of male palp and epigynum (especially shapes of bulb and scape; Figs 26 – 28). From all other congeners by simple procursus and complex bulbal processes (Figs 26 – 27) and by narrow epigynal scape.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Dinagat Isl., near Libjo, Paragua Forest, ‘ site 1 ’ (10.222 ° N, 125.553 ° E), 130 m a. s. l., forest at brook, among low vegetation, 20 Feb. 2014 (B. A. Huber), ZFMK (Ar 13929). Other material PHILIPPINES: 6 ♂♂, 9 ♀♀, same data as holotype, ZFMK (5 ♂♂, 8 ♀♀) (Ar 13930 – 31) and MSU-IIT (1 ♂, 1 ♀); 3 juveniles in pure ethanol, same data, ZFMK (Phi 231).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.1, carapace width 1.1. Leg 1: 37.4 (9.0 + 0.5 + 8.8 + 16.2 + 2.9), tibia 2: 5.1, tibia 3: 3.3, tibia 4: 4.8; tibia 1 L / d: 74. Distance PME-PME 270 µm, diameter PME 125 × 95 µm, distance PME-ALE 20 µm; AME absent. COLOR. Carapace ochre-yellow with black lateral margins, dark mark behind ocular area and dark median mark on posterior rim; ocular area and clypeus pale ochre; sternum bright orange; leg coxae ochre-yellow, other leg segments greenish-ochre (especially femora) to light brown (distal segments); abdomen grey with distinct dorsal and lateral pattern of black marks, ventrally with brown band between gonopore and spinnerets (mid-section indistinct). BODY. Habitus as in Fig. 8; ocular area slightly raised, each triad on low hump directed toward lateral; carapace only anteriorly with very shallow and narrow median furrow; clypeus barely modified, indistinct median process near rim provided with some slightly stronger hairs; sternum wider than long (0.70 / 0.55), unmodified. Gonopore with four epiandrous spigots in two pairs (Fig. 37). ALS as in female (cf. Fig. 36). CHELICERAE. As in Fig. 40, with pair of ridge-shaped apophyses distally near median line, without proximal lateral apophyses; without modified hairs; without stridulatory ridges. PALPS. As in Figs 26 – 27, coxa unmodified, trochanter on retrolateral side with large apophysis fused to femur; femur with small retrolateral process distally; tibia length / width: 0.50 / 0.22; retrolateral trichobothrium on tibia very distal; tarsus with semitransparent simple procursus directed toward patella; tarsal organ exposed (Fig. 35); genital bulb large, with small retrolateral process proximally and three distinctive processes distally (Figs 30 – 31, 33, 38 – 39): dorsal process with distal hook; central hinged process with large prolateral flap and complex tip; ventral hinged process with simple pointed tip. Location of sperm duct opening unknown. LEGS. Without spines, with curved hairs on metatarsi, with vertical hairs in higher than usual density in one retrolatero-dorsal row on each tibia; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 30 pseudosegments, distally distinct. Tarsus 4 comb-hairs with very dense tines (Fig. 32). Male (variation) Tibia 1 in 6 other males: 8.3 – 9.0 (mean 8.6); ratio of palpal tibia length / width: 2.1 – 2.4. Some specimens with white marks on abdomen in addition to black marks. Female In general similar to male (Fig. 9); eye triads closer together (distance PME-PME 215 µm); clypeus unmodified; tibiae with short vertical hairs in low density; entire median area of carapace darker; clypeus also slightly darker; abdomen with or without white marks; tibia 1 in 9 females: 6.2 – 7.2 (mean 6.7). Epigynum with large, weakly sclerotized area, posteriorly protruding with short, narrow scape (Figs 28, 45 – 46); internal genitalia with large median membranous structure of unknown function (Fig. 29).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	biology_ecology	Natural history The spiders were found in strongly domed webs among the vegetation. When disturbed, they ran a short distance in the dome, vibrated vigorously for a short time, and then made slow circular movements with their abdomen. Males and females were often found close to each other, but always in separate webs. The locality is shared with the ground-dwelling A. paragua Huber, sp. nov.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276032FF879583FC8347FCFC69.taxon	distribution	Distribution Known from type locality only (Fig. 3).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	description	urn: lsid: zoobank. org: act: AA 06 D 026 - A 4 F 7 - 4 EA 9 - 9661 - 77707 B 2 CCFB 9 Figs 10 – 12, 41 – 44, 48 – 50	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	diagnosis	Diagnosis Distinguished from very similar A. libjo Huber, sp. nov. by shapes of bulbal processes (compare Figs 38 – 39 and 41 – 42), by longer male palpal tibia (2.9 – 3.2 × longer than wide vs. 2.1 – 2.4 in A. libjo Huber, sp. nov.), and by longer epigynum and narrower scape (compare Figs 28 and 43); from next closest known relative (A. ocampoi Huber, sp. nov.) by light coloration and several details of male palp and epigynum (especially shapes of bulb and scape). Distinguished from all other congeners by simple procursus and complex bulbal processes and by narrow epigynal scape.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Mindanao, Davao del Sur Prov., Marilog Distr., Baganihan (7.469 ° N, 125.250 ° E), 1210 m a. s. l., primary forest near road, among low vegetation, 15 Feb. 2014 (B. A. Huber), ZFMK (Ar 13932). Other material PHILIPPINES: Mindanao, 2 ♂♂, 3 ♀♀, same data as holotype, ZFMK (Ar 13933); 1 juvenile, in pure ethanol, same data as holotype, ZFMK (Phi 258).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.2, carapace width 1.1. Leg 1: 39.3 (9.0 + 0.5 + 9.1 + 17.4 + 3.3), tibia 2: 5.3, tibia 3: 3.6, tibia 4: 5.2; tibia 1 L / d: 77. Distance PME-PME 325 µm, diameter PME 135 × 110 µm, distance PME-ALE 20 µm; AME absent. COLOR. Carapace ochre-yellow with black lateral margins, pair of small dark marks behind ocular area, and dark median mark at posterior rim; ocular area and clypeus pale ochre; sternum bright orange; leg coxae ochre-yellow, other leg segments greenish-ochre (especially femora) to light brown (distal segments); abdomen grey, with distinct dorsal and lateral pattern of black marks and rows of white marks, ventrally with brown band between gonopore and spinnerets (mid-section indistinct). BODY. Habitus as in Figs 10 – 12; ocular area slightly raised, each triad on low hump directed toward lateral; carapace only anteriorly with very shallow and narrow median furrow; clypeus with indistinct pair of processes near rim, provided with some slightly stronger hairs; sternum wider than long (0.70 / 0.55), unmodified. CHELICERAE. As in A. libjo Huber, sp. nov. (cf. Fig. 40), with pair of ridge-shaped apophyses distally near median line, without proximal lateral apophyses; without modified hairs; without stridulatory ridges. PALPS. Proximal segments very similar to A. libjo (cf. Figs 26 – 27), trochanter apophysis and femur apophysis slightly larger, tibia longer (length / width: 0.70 / 0.24). Genital bulb longer than in A. libjo sp. nov., with all three processes different (Figs 41 – 42): dorsal process wider proximally; central hinged process with smaller prolateral flap and different tip; ventral hinged process relatively shorter. LEGS. Without spines, with curved hairs on metatarsi 1 – 3, with vertical hairs in higher than usual density in one retrolatero-dorsal row on each tibia; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 30 pseudosegments, distally distinct. Male (variation) Tibia 1 in 2 other males: 8.9, 9.3; ratio of palpal tibia length / width: 2.9, 3.2. Other males without or with indistinct white marks on abdomen. Female In general similar to male; eye triads closer together (distance PME-PME 230 µm); clypeus unmodified; tibiae with short vertical hairs in low density; entire median area on carapace darker; clypeus also slightly darker (one female with wide brown median band on carapace extending over ocular area and clypeus); abdomen with or without white marks; tibia 1 in 3 females: 6.7, 6.8, 7.0. Epigynum with large, weakly sclerotized area, posteriorly protruding with short, narrow scape (Figs 43, 48 – 49); internal genitalia with large median membranous structure of unknown function (Fig. 44).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	biology_ecology	Natural history The spiders were found in strongly domed sheet-webs among high grasses in well-preserved forest. As in A. libjo Huber, sp. nov., males and females were sometimes found close to each other but in separate webs. The locality is shared with the ground-dwelling A. kiukoki Huber, sp. nov.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602DFF86956AFC7547FCF878.taxon	distribution	Distribution Known from type locality only (Fig. 3).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	description	urn: lsid: zoobank. org: act: 23 ED 16 FE-E 6 AA- 40 B 4 - B 55 E-A 2 C 31 CB 86 A 22 Figs 51 – 52, 57 – 69, 87 – 89	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. paragua Huber, sp. nov., A. loboc Huber, sp. nov., A. pasambai Huber, sp. nov.) by distinctive modification of male clypeus (Figs 59, 62, 65; similar only in A. paragua Huber, sp. nov., see Fig. 72), and long tongue-shaped posterior projection of epigynum (Figs 60, 87; very similar in A. paragua Huber, sp. nov., see Fig. 73; much shorter in A. loboc Huber, sp. nov., see Fig. 78; female of A. pasambai Huber, sp. nov. unknown). Distinguished from A. paragua Huber, sp. nov. also by longer male eye stalks (Fig. 59), more strongly curved apophysis on male palpal femur (Fig. 58), different shape of distal procursus elements (Fig. 58), and pore plates closer together (Fig. 61). Distinguished from A. loboc Huber, sp. nov. and A. pasambai Huber, sp. nov. also by modification of male palpal femur (only one large retrolateral process; Fig. 58) and absence of median process on male ocular area.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	etymology	Etymology Named for Filipino painter Ang Kiukok (1931 – 2005).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Mindanao, Davao del Sur Prov., Marilog Distr., Baganihan (7.469 ° N, 125.250 ° E), 1210 m a. s. l., primary forest near road, near ground, 15 Feb. 2014 (B. A. Huber), ZFMK (Ar 13934). Other material PHILIPPINES, Mindanao Isl.: 7 ♂♂, 7 ♀♀, 9 juvs, same data as holotype, ZFMK (Ar 13935 - 36); 1 ♀, 6 juvs, in pure ethanol, same data, ZFMK (Phi 256). – 1 ♀, 11 juvs from Barangay Baganihan, ‘ site 1 ’ (7.438 ° N, 125.226 ° E), 1000 m a. s. l., 5 Dec. 2014 (M. A. Responte), MSU-IIT; 4 ♀♀, 17 juvs, from same locality, ‘ site 2 ’ (7.456 ° N, 125.239 ° E), 6 Dec. 2014 (M. A. Responte), MSU-IIT; 9 juvs, from same locality, ‘ site 3 ’ (7.470 ° N, 125.245 ° E), 7 Dec 2014 (M. A. Responte), MSU-IIT. – 4 ♀♀, 1 juv., Marilog Distr., Epol Spring Resort (7.456 ° N, 125.237 ° E), ~ 1100 m a. s. l., degraded forest, near ground, 15 Feb. 2014 (B. A. Huber), ZFMK (Ar 13937); 3 ♀♀, in pure ethanol, same data, ZFMK (Phi 253). – 1 ♂, 1 ♀, 10 juvs, from Epol Falls, ‘ site 2 ’ (7.454 ° N, 125.239 ° E), 1150 m a. s. l., 2 Dec. 2014 (M. A. Responte), MSU-IIT; 1 ♀, 4 juvs, from same locality, ‘ site 3 ’ (7.451 ° N, 125.240 ° E), 1200 m a. s. l., 3 Dec. 2014 (M. A. Responte), MSU-IIT; 1 juv., from same locality, ‘ site 1 ’ (7.455 ° N, 125.237 ° E), 1100 m a. s. l., 1 Dec. 2014 (M. A. Responte), MSU-IIT. – 6 ♂♂, 2 ♀♀, 2 juvs, Mt. Matutum, Kawit Forest, ‘ site 1 ’ (6.338 ° N, 125.104 ° E), 950 m a. s. l., along brook, near ground, 13 Feb. 2014 (B. A. Huber), ZFMK (Ar 13938); 4 juvs, in pure ethanol, same data, ZFMK (Phi 268). – 1 ♂, 2 ♀♀, Bukidnon Prov., Barangay San Jose, Blue Water Cave (7.706 ° N, 125.032 ° E), 200 m a. s. l., near ground at cave entrance, 16 Feb. 2014 (B. A. Huber), ZFMK (Ar 13939); 4 ♀♀, 1 juv., in pure ethanol, same data, ZFMK (Phi 251). – 2 ♂♂, 2 ♀♀, 1 juv., Barangay San Jose, Kabyaw Cave (~ 7.704 ° N, 125.038 ° E), ~ 200 m a. s. l., near ground near cave entrance, 16 Feb. 2014 (B. A. Huber), ZFMK (Ar 13940). – 4 ♂♂, 8 ♀♀, Barangay San Jose, doline near Kabyaw Cave (7.703 ° N, 125.038 ° E), 220 m a. s. l., near ground, 16 Feb. 2014 (B. A. Huber), ZFMK (Ar 13941). – 3 ♀♀, Bukidnon Prov., CEDAR (Center for Ecological Development and Recreation) (8.251 ° N, 125.034 ° E), 760 m a. s. l., forest along river, near ground, 16 Feb. 2014 (B. A. Huber), ZFMK (Ar 13942); 1 ♀, in pure ethanol, same data, ZFMK (Phi 246). – 1 ♂, 4 ♀♀, 3 juvs, from CEDAR (8.251 ° N, 125.027 ° E), 15 Nov. 2014 (E. P. Mondejar), MSU-IIT. – 3 ♂♂, 7 ♀♀, Bukidnon Prov., Santo Domingo (7.782 ° N, 125.397 ° E), 560 m a. s. l., forest remnant along brook, near ground, 8 – 9 Feb. 2014 (B. A. Huber), ZFMK (Ar 13943); 1 ♀, 6 juvs, in pure ethanol, same data, ZFMK (Phi 285). – 2 ♂♂, Bukidnon Prov., near Santo Domingo, Penolohan (7.769 ° N, 125.420 ° E), 640 m a. s. l., forest above Salug River, near ground, 8 Feb. 2014 (B. A. Huber) ZFMK (Ar 13944); 2 ♀♀ in pure ethanol, same data, ZFMK (Phi 277). – 5 ♂♂, 15 ♀♀, 3 juvs, Misamis Occidental Prov., Iligan, NPC Nature’s Park near Cristina Falls (8.186 ° N, 124.192 ° E), 90 m a. s. l., near ground, 17 Feb. 2014 (B. A. Huber), ZFMK (Ar 13945 - 46); 3 ♀♀, 2 juvs, in pure ethanol, same data, ZFMK (Phi 244). PHILIPPINES, Camiguin Isl.: 13 ♂♂, 11 ♀♀, 1 juv., Katibawasan Falls (9.215 ° N, 124.720 ° E), 300 m a. s. l., near ground, 19 Feb. 2014 (B. A. Huber, P. N. Banaag), ZFMK (Ar 13947 - 48); 1 ♀, in pure ethanol, same data, ZFMK (Phi 236). – 9 ♂♂, 12 ♀♀, 20 juvs, from Katibawasan (9.213 ° N, 124.718 ° E), 5 – 6 May 2014 (E. P. Mondejar), MSU-IIT. – 8 ♂♂, 6 ♀♀, 1 juv., Mt. Hibok Hibok (9.196 ° N, 124.692 ° E), 600 m a. s. l., near ground, 18 Feb. 2014 (B. A. Huber, P. N. Banaag), ZFMK (Ar 13949 - 50); 2 ♀♀, 1 juv., in pure ethanol, same data, ZFMK (Phi 240). Assigned tentatively (no male available) PHILIPPINES, Mindanao Isl., Davao Oriental: 1 ♀, from Mount Hamiguitan WS (access San Isidro), ‘ site 1 ’ (6.720 ° N, 126.172 ° E), 490 m a. s. l., 9 Feb. 2015 (M. A. Responte), ZFMK (Ar 13951); 7 juvs, from same locality, ‘ site 3 ’ (6.732 ° N, 126.179 ° E), 1250 m a. s. l., 11 Feb. 2015 (M. A. Responte), MSU-IIT.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.0, carapace width 1.2. Leg 1: 34.2 (7.9 + 0.5 + 8.0 + 14.8 + 3.0), tibia 2: 5.0, tibia 3: 3.5, tibia 4: 5.1; tibia 1 L / d: 72. Distance PME-PME 355 µm, diameter PME 150 × 120 µm, distance PME-ALE ~ 35 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including ocular area and clypeus; sternum with pair of wide brown bands converging posteriorly; legs greenish ochre with slightly darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black and white marks, ventrally with dark band behind gonopore and very indistinct mark in front of spinnerets. BODY. Habitus as in Figs 51 – 52; ocular area raised, each triad on additional short stalk directed toward lateral (Figs 59, 62 – 63), without median process; carapace with very shallow median furrow in anterior part only (Fig. 63); clypeus with large distinctive process, strongly protruding in upper part, with two pairs of rounded apophyses in distal part, densely covered with small scales (Figs 65, 66); sternum wider than long (0.75 / 0.60), unmodified. ALS and PMS as in Fig. 68. CHELICERAE. As in Fig. 59, with pair of dark lateral apophyses proximally and pair of weakly sclerotized humps laterally, without modified hairs; without stridulatory ridges. PALPS. As in Figs 57 – 58; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with strong retrolateral apophysis distally curved toward ventral; patella large; tibia small, dorsal trichobothrium in very proximal position, retrolateral trichobothrium in very distal position; tarsus with long procursus, distally complex, apparently with two hinged structures; bulb with large embolus and smaller, semitransparent, pointed process. LEGS. Without spines; with curved hairs on metatarsi 1 – 3 (few curved hairs also on tibiae 1 – 2); few vertical hairs; retrolateral trichobothrium on tibia 1 at 3 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 30 pseudosegments, only distally distinct. Male (variation) Tibia 1 in 48 other males: 7.0 – 9.8 (mean: 8.2). Ventral mark behind gonopore in most males slightly asymmetrical. Female In general similar to male (Fig. 53) but clypeus unmodified; eye triads much closer together (distance PME-PME 155 µm), not on stalks (Fig. 64); without stridulatory apparatus between carapace and abdomen. Tibia 1 in 56 females: 4.9 – 7.5 (mean: 6.1); dark and light rings on legs often more distinct than in males. Tarsus 4 comb-hairs with very dense tines (Fig. 69). Epigynum in anterior part weakly sclerotized, internal structures visible through cuticle, with flat, tongue-shaped scape (Figs 60, 87), without membranous pockets behind epigynum. Internal genitalia as in Figs 61 and 89; pore plates close together; without sclerotized internal pockets; with distinct transversal sclerotized ridges provided with many small teeth. The single female from Mt. Hamiguitan cannot be unambiguously assigned to this species rather than to A. paragua and is therefore assigned tentatively.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	biology_ecology	Natural history The spiders were found in domed sheet webs close to the ground, usually in well protected dark spaces among and under large rocks and logs in forests. Males and females were sometimes found together in one web. At Baganihan, some specimens (especially juveniles) were found among mosses in deep furrows of trees up to 1.5 m above the ground. At both localities on Camiguin Island, the spiders were observed to be much more agile and quick when disturbed than at other localities.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827602BFF8C959EFEB041B3FC92.taxon	distribution	Distribution Known from numerous localities on Mindanao Island and from Camiguin Island (Fig. 4).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	description	urn: lsid: zoobank. org: act: 9 E 9 FEDE 8 - BEB 5 - 46 D 4 - 929 C- 6 C 2259 AA 7 FDB Figs 54, 70 – 74, 90 – 92	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. kiukoki Huber, sp. nov., A. loboc Huber, sp. nov., A. pasambai Huber, sp. nov.) by distinctive modification of male clypeus (Fig. 72; similar only in A. kiukoki Huber, sp. nov.), and tongue-shaped posterior projection of epigynum (Fig. 73; very similar to A. kiukoki Huber, sp. nov.; much shorter in A. loboc Huber, sp. nov.; female of A. pasambai Huber, sp. nov. unknown). Distinguished from A. kiukoki Huber, sp. nov. also by shorter male eye stalks (Fig. 72), weakly curved apophysis on male palpal femur (Fig. 71), different shape of distal procursus elements (Fig. 71), and pore plates wider apart (Fig. 74). Distinguished from A. loboc Huber, sp. nov. and A. pasambai Huber, sp. nov. also by modification of male palpal femur (only one large retrolateral process) and absence of median process in male ocular area.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Dinagat Isl., near Libjo, Paragua Forest, ‘ site 1 ’ (10.222 ° N, 125.553 ° E), 130 m a. s. l., forest at brook, near ground, 20 Feb. 2014 (B. A. Huber), ZFMK (Ar 13952). Other material PHILIPPINES: Dinagat Isl., 2 ♂♂, 4 ♀♀, same data as holotype, ZFMK (Ar 13953); 1 ♂, in pure ethanol, same data, ZFMK (Phi 230). – 1 ♀, 1 juv., Paragua Forest, ‘ site 2 ’ (10.241 ° N, 125.545 ° E), 240 m a. s. l., forest along brook, near ground, 20 Feb. 2014 (B. A. Huber), ZFMK (Ar 13954).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.0, carapace width 1.2. Leg 1: 35.2 (8.0 + 0.5 + 8.2 + 15.5 + 3.0), tibia 2: 4.9, tibia 3: 3.4, tibia 4: 4.9; tibia 1 L / d: 67. Distance PME-PME 230 µm, diameter PME 135 × 110 µm, distance PME-ALE ~ 35 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including ocular area and clypeus; sternum with pair of wide brown bands converging posteriorly, labium proximally dark brown; legs greenish ochre with barely visible darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, with few indistinct white marks, ventrally with dark band behind gonopore and very indistinct mark in front of spinnerets. BODY. Habitus as in A. kiukoki Huber, sp. nov. (cf. Figs 51 – 52); ocular area raised, each triad on additional hump directed toward lateral, without median process; carapace with very shallow median furrow in anterior part only; clypeus with large distinctive process (Fig. 72), strongly protruding in upper part, with two pairs of rounded apophyses set with small scales; sternum wider than long (0.80 / 0.55), unmodified. CHELICERAE. As in Fig. 72, with pair of dark lateral apophyses proximally, barely visible lateral humps more distally; without modified hairs; without stridulatory ridges. PALPS. As in Figs 70 – 71; very similar to A. kiukoki Huber, sp. nov., but retrolateral apophysis of femur less strongly curved, distal elements of procursus different, and semitransparent pointed process on bulb smaller. LEGS. Without spines; with curved hairs on metatarsi 1 – 3 (few curved hairs also on tibiae 1 – 2); few vertical hairs; retrolateral trichobothrium on tibia 1 at 3 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 30 pseudosegments, only distally distinct. Male (variation) Tibia 1 in 2 other males: 8.2, 8.3. Female In general similar to male but clypeus unmodified; eye triads much closer together (distance PME-PME 125 µm); without stridulatory apparatus between carapace and abdomen. Tibia 1 in 4 females: 5.5, 5.6, 5.6, 5.8; dark and light rings on legs often more distinct than in males. Epigynum and internal genitalia very similar to those of A. kiukoki Huber, sp. nov., but tongue-shaped posterior projection slightly shorter and wider (Figs 73, 90), and pore plates wider apart (Fig. 74).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	biology_ecology	Natural history The spiders were found in domed webs very close to the ground. At the same locality, A. libjo Huber, sp. nov. occurred higher among the vegetation.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276026FF8E959BFCD44164FAD3.taxon	distribution	Distribution Known from two neighboring localities on Dinagat Island only (Fig. 4).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	description	urn: lsid: zoobank. org: act: CAD 63 F 67 - D 442 - 43 BF-ACBC- 062 F 4 B 3 DCA 0 A Figs 55 – 56, 75 – 86, 93 – 95	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. pasambai Huber, sp. nov.) by distinctive modification of male clypeus (Fig. 77; longer and narrower), by median process between eye stalks larger and in more dorsal position (Fig. 77), by longer male palpal tibia and procursus (Fig. 76), and by stronger ventral apophysis on male palpal femur (female of A. pasambai Huber, sp. nov. unknown); from other close relatives (A. paragua Huber, sp. nov.; A. kiukoki Huber, sp. nov.) by modifications of male palpal femur (two processes instead of one), presence of median process between eye stalks, and very short tongueshaped posterior projection of epigynum (Fig. 78).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Bohol Isl., near Loboc, above Loboc River (9.651 ° N, 124.022 ° E), ~ 50 m a. s. l., ravine in degraded forest, 5 Mar. 2014 (B. A. Huber), ZFMK (Ar 13955). Other material PHILIPPINES: Bohol Isl., 4 ♂♂, 13 ♀♀, 1 juv., same data as holotype, ZFMK (Ar 13956 - 57); 3 ♀♀, 3 juvs, in pure ethanol, same data as holotype, ZFMK (Phi 201). – 3 ♂♂, 3 ♀♀, at Loboc River (9.651 ° N, 124.022 ° E), near ground, 20 m a. s. l., 4 Mar. 2014 (B. A. Huber), ZFMK (Ar 13958). – 1 ♂, 1 ♀, Rajah Sikatuna (Magsaysay Park) (9.705 ° N, 124.123 ° E), 430 m a. s. l., forest, 6 Mar. 2014 (B. A. Huber), ZFMK (Ar 13959).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.1, carapace width 1.2. Leg 1: 32.4 (7.5 + 0.5 + 7.6 + 14.1 + 2.7), tibia 2: 4.5, tibia 3: 3.2, tibia 4: 4.9; tibia 1 L / d: 73. Distance PME-PME 340 µm, diameter PME 135 × 120 µm, distance PME-ALE ~ 35 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including ocular area and clypeus; sternum with pair of wide brown bands converging posteriorly, labium proximally not dark brown; legs ochre to light brown, with indistinct darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, with few indistinct white marks, ventrally with small spot behind gonopore and very indistinct mark in front of spinnerets. BODY. Habitus as in Fig. 55; ocular area raised, each triad on short stalk directed toward lateral, with distinctive median process (Fig. 77); carapace with very shallow median furrow in anterior part only; clypeus with large distinctive process (Fig. 77), strongly protruding in upper part, with distinctive pair of distal apophyses; sternum wider than long (0.75 / 0.60), unmodified. CHELICERAE. As in Fig. 77, with pair of dark lateral apophyses proximally, without lateral humps more distally; without modified hairs; without stridulatory ridges. PALPS. As in Figs 75 – 76; coxa unmodified; trochanter with wide ventral apophysis; femur large, with strong ventral and slender retrolateral apophyses; patella very large; tibia small but long, dorsal trichobothrium in very proximal position, retrolateral trichobothrium in very distal position; tarsus with very long procursus, distally complex, apparently with two hinged structures; bulb with large embolus and small semitransparent pointed process. LEGS. Without spines; with curved hairs on metatarsi 1 – 3 (few curved hairs also on tibiae 1 – 2); few vertical hairs; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, only distally distinct. Male (variation) Tibia 1 in 7 other males: 6.5 – 7.7 (mean 7.1). Female In general similar to male but clypeus unmodified; eye triads much closer together (distance PME-PME 135 µm); without stridulatory apparatus between carapace and abdomen; without ventral dark band on abdomen. Tibia 1 in 17 females: 5.0 – 5.7 (mean: 5.3); dark and light rings on legs often more distinct than in males. Epigynum in anterior part weakly sclerotized, internal structures partly visible through cuticle, with short posterior projection with submarginal transversal groove (Figs 78, 85 – 86, 93), without membranous pockets behind epigynum; internal genitalia as in Figs 79 and 95; without sclerotized internal pockets; with distinct transversal sclerotized ridges provided with many small teeth.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	biology_ecology	Natural history The spiders were found in domed webs close to the ground, hidden under rocks or logs.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276024FFB5958DFA1F4151FE4D.taxon	distribution	Distribution Known from two neighboring localities on Bohol Island only (Fig. 4).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	description	urn: lsid: zoobank. org: act: 041 DF 955 - 4589 - 4503 - AF 5 D- 1 EC 15 C 107 E 8 E Figs 96 – 98	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. loboc Huber, sp. nov.) by distinctive modification of male clypeus (Fig. 96; shorter and wider), by median process between eye stalks smaller and in more frontal position (Fig. 96), by shorter male palpal tibia and procursus (Fig. 98), and by smaller ventral apophysis on male palpal femur (female of A. pasambai Huber, sp. nov. unknown); from other close relatives (A. paragua sp. nov., A. kiukoki Huber, sp. nov.) by modifications of male palpal femur (two processes instead of one), and by presence of median process between eye stalks.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	etymology	Etymology Named for Philippine-born cellist Wilfredo Pasamba.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Negros Isl., Negros Oriental Prov., Casaroro Falls (9.281 ° N, 123.208 ° E), 550 m a. s. l., forest along river below waterfall, 10 Mar. 2014 (B. A. Huber), ZFMK (Ar 13960). Other material None.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.3, carapace width 1.1. Leg 1: 32.7 (7.7 + 0.5 + 7.7 + 14.1 + 2.7), tibia 2: 4.7, tibia 3: 3.7, tibia 4: 5.0; tibia 1 L / d: 69. Distance PME-PME 385 µm, diameter PME 135 × 120 µm, distance PME-ALE ~ 35 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including ocular area and clypeus; sternum with pair of wide brown bands converging posteriorly, with darker triangular mark posteriorly, labium proximally not dark brown; legs greenish ochre, with indistinct darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, with some white marks, ventrally with dark band behind gonopore and very indistinct mark in front of spinnerets. BODY. Habitus as in A. loboc Huber, sp. nov. (cf. Fig. 55); ocular area raised, each triad on short stalk directed toward lateral, with small median process in frontal position (Fig. 96); carapace with very shallow median furrow in anterior part only; clypeus with large distinctive process (Fig. 96), protruding in upper part, with distinctive pair of distal apophyses; sternum wider than long (0.80 / 0.55), unmodified. CHELICERAE. As in Fig. 96, with pair of dark lateral apophyses proximally, without lateral humps more distally; without modified hairs; without stridulatory ridges. PALPS. As in Figs 97 – 98; very similar to A. loboc Huber, sp. nov., but femur apophyses of different sizes and positions, tibia and procursus shorter. LEGS. Without spines; with curved hairs on tibiae and metatarsi 1 – 2; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, only distally distinct. Female Unknown	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	biology_ecology	Natural history The specimen was found in a small hole in the ground.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601FFFB79562FD9046CDFD72.taxon	distribution	Distribution Known from type locality on Negros Island only (Fig. 4).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	description	Figs 99 – 103, 111 – 112, 163 – 165	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	discussion	Note The original description was based on two males. Here we present data on new material from the type locality, a description of the female, and an emended diagnosis to account for the newly described congeners.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. lambir Huber, sp. nov.) by shape of prolatero-ventral apophysis of male palpal femur (pointed tip and subdistal branch of approximately same length; cf. Huber 2005 a: fig. 121); also by presence of prolateral apophysis on femur, by details of procursus (shapes of sclerites on complex distal part; cf. Huber 2005 a: figs 121 – 122), and by female genitalia (pair of internal pockets; position of pore plates; Figs 111 – 112).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	materials_examined	New material examined MALAYSIA-BORNEO, Sabah: 1 ♂, 1 ♀, 1 juv., Mt. Kinabalu, forest along Silau Silau Trail (6.010 – 6.017 ° N, 116.537 – 116.543 ° E), 1550 – 1650 m a. s. l., domed webs among vegetation, 6 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13961); 2 ♀♀, 1 juv., in pure ethanol, same data, ZFMK (Bor 211). – 1 ♀, Kinabalu N. P., 1550 m a. s. l., 2 – 8 Apr. 1998 (C. L. Deeleman-Reinhold, P. Zborowski), RMNH. – 2 ♂♂, 5 ♀♀, 3 juvs, Crocker Range between Kota Kinabalu and Tambuan, S-slope, forest along river (5.783 ° N, 116.338 – 116.340 ° E), 1430 – 1480 m a. s. l., domed webs among vegetation, 3 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13962); 4 juvs, in pure ethanol, same data, ZFMK (Bor 171). – 2 ♂♂, 3 ♀♀, in very poor condition, Tawau (4.406 ° N, 117.892 ° E), 6 Sep. 2009 (A. Floren), RMNH. MALAYSIA-BORNEO, Sarawak: 2 ♂♂, 6 ♀♀, 3 juvs, Gunung Mulu N. P., forest near Deer Cave (4.027 ° N, 114.818 ° E), 60 m a. s. l., 23 – 24 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13963 - 64); same data, 3 ♀♀, 5 juvs, in pure ethanol, ZFMK (Bor 182). – 5 ♂♂, 2 ♀♀, 4 juvs, Bario, forest along river W of town (3.736 ° N, 115.437 – 115.443 ° E), 1150 – 1250 m a. s. l., domed webs among vegetation, 30 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13965). – 1 ♀, 1 juv., Bario, forest along river N of town (3.765 – 3.771 ° N, 115.444 – 115.448 ° E), 1170 – 1250 m a. s. l., domed webs among vegetation, 29 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13966); 9 juvs, in pure ethanol, same data, ZFMK (Bor 233).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	description	Description (amendments to Huber 2005 a) Male Thoracic furrow absent (contra Huber 2005 a); clypeus slightly protruding (more than in female). Tibia 2 slightly longer than tibia 4 (e. g., 5.1 / 4.9; 4.9 / 4.8); curved hairs on metatarsi 2; tibia 1 in 7 males: 7.6 – 8.8 (mean: 8.3). Female Eye triads much closer together than in male (distance PME-PME ~ 235 µm vs. 375 µm), not on stalks. Tibia 1 in 15 females: 5.9 – 7.3 (mean 6.4). Epigynum simple plate, darker laterally anteriorly; anterior internal arc and internal sclerotized pockets visible through cuticle (Figs 111, 163 – 164). Internal genitalia as in Figs 112 and 165, with pair of sclerotized pockets and more lateral pair of membranous pockets. Variation In males from Gunung Mulu, the tiny cone-shaped process on the genital bulb (cf. Huber 2005 a: figs 121 and 122) is absent. Males from Bario with small additional process at basis of prolatero-ventral apophysis of femur.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	biology_ecology	Natural history Webs were in most cases made of two sheets: an upper sheet in which the spider hung, and a lower sheet that had to be removed in order to catch the spider. At most localities, webs were found high among the vegetation, even in the sunlight. At Gunung Mulu, webs were found among mosses and low vegetation on perpendicular rock surfaces about 2 m above the ground. At several localities, A. kinabalu was sympatric with a ground-dwelling congener: with A. poring Huber, sp. nov. at Mt. Kinabalu and Poring; with A. indah Huber, sp. nov. at Crocker Range.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601DFFB09597FD7F46C0FD81.taxon	distribution	Distribution Widely distributed in Sabah and eastern Sarawak (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	description	urn: lsid: zoobank. org: act: EFFAC 871 - 4657 - 461 F- 9875 - 807 E 767 DB 6 B 8 Figs 105 – 106, 113 – 125, 166 – 168	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. kinabalu) by shape of prolatero-ventral apophysis of male palpal femur (Fig. 113; pointed tip much longer than subdistal branch); also by details of procursus (shapes of sclerites on complex distal part), by absence of prolateral apophysis on femur, and by female genitalia (no internal sclerotized pockets; pore plates wider apart; Figs 116 – 117).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	materials_examined	Material examined Holotype MALAYSIA-BORNEO: ♂, Sarawak, Lambir Hills N. P. (4.198 – 4.207 ° N, 114.034 – 114.045 ° E), 60 – 150 m a. s. l., in domed webs among low vegetation, 22 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13967). Other material MALAYSIA-BORNEO, Sarawak: 9 ♂♂, 11 ♀♀, same data as holotype, ZFMK (8 ♂♂, 10 ♀♀; Ar 13968 - 69) and SMK (1 ♂, 1 ♀); 1 ♂, 3 ♀♀, 4 juvs, in pure ethanol, same data, ZFMK (Bor 201). – 3 ♂♂, 4 ♀♀, Niah Cave N. P., forest near headquarters (3.820 ° N, 113.763 ° E), 40 m a. s. l., night collecting, domed webs among vegetation, 28 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13970). – 6 ♂♂, 3 ♀♀, Niah Cave N. P., forest near cave (3.814 ° N, 113.771 ° E), 40 m a. s. l., among low vegetation, 28 July 2014 (B. A. Huber), ZFMK (Ar 13971). – 3 ♂♂, 11 ♀♀, 1 juv., Niah Cave N. P., forest along main trail (3.814 – 3.821 ° N, 113.763 – 113.771 ° E), 20 – 40 m a. s. l., domed webs among vegetation, 27 July 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13972 - 73); 1 ♂, 1 ♀, 1 juv., same data, in pure ethanol, ZFMK (Bor 178).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.5, carapace width 0.95. Leg 1: 33.1 (7.9 + 0.4 + 7.6 + 14.3 + 2.9), tibia 2: 4.4, tibia 3: 2.8, tibia 4: 4.2; tibia 1 L / d: 87. Distance PME-PME 340 µm, diameter PME 105 µm, distance PME-ALE 25 µm; AME absent. COLOR. Carapace pale ochre with black lateral margins and brown median band including ocular area. Clypeus mostly pale ochre, at rim brown. Sternum pale ochre, laterally slightly darker. Legs ochre to light brown, slightly darker rings on femora (subdistally) and tibiae (proximally and subdistally); tips of femora and tibiae whitish. Abdomen with dorsal and lateral pattern of black and white marks; ventrally with brown mark near spinnerets. BODY. Habitus as in Fig. 105; ocular area slightly raised, each triad on short stalk directed toward lateral; carapace without thoracic furrow; clypeus slightly more protruding than usual; sternum wider than long (0.60 / 0.45), unmodified. CHELICERAE. As in Fig. 115, with pair of proximal lateral apophyses and pair of simple distal apophyses in very lateral position; without modified hairs; without stridulatory ridges. PALPS. As in Figs 113 – 114, coxa unmodified; trochanter with short, rounded ventral apophysis; femur with distinct retrolatero-ventral apophysis, long prolatero-ventral apophysis with side branch, no prolateral process. Procursus complex, ventral transparent lamina proximally wide and slightly sclerotized; retrolatero-ventral process indistinct, apparently fused to procursus. Bulb simple, with short and wide embolus. LEGS. Without spines, with curved hairs on metatarsi 2 only (single dorsal row; proximal half), with few vertical hairs; retrolateral trichobothrium on tibia 1 at 2.5 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with> 20 pseudosegments, only distally fairly distinct. Male (variation) Tibia 1 in 19 other males: 6.1 – 8.5 (mean 7.4); most males with small dark mark at gonopore. Female In general similar to male; eye triads closer together (distance PME-PME 220 µm), not on stalks; clypeus less protruding; no curved hairs on metatarsi. Tibia 1 in 26 females: 5.6 – 6.6 (mean 6.2). Epigynum very simple plate, wider than long (Fig. 125), anterior internal arc visible through cuticle (Figs 116, 166). Internal genitalia very simple, as in Figs 117 and 168, without internal sclerotized pockets, with pair of lateral membranous pockets. ALS as in Fig. 124.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	biology_ecology	Natural history At the type locality (Lambir), the spiders were found among vegetation close to the ground, while at Niah they occurred higher among the vegetation, very similar to A. kinabalu, in the same type of twolayered web (see above).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827601AFFBC9594FDCD4623FEDE.taxon	distribution	Distribution Known from two localities in eastern Sarawak (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	description	urn: lsid: zoobank. org: act: 3 FF 192 B 2 - 3 F 88 - 4 BBF-A 675 - CA 77 FF 9 A 72 F 1 Figs 104, 126 – 144, 169 – 171	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. kinabalu, A. lambir Huber, sp. nov., A. indah Huber, sp. nov., A. poring Huber, sp. nov.) by shape of prolatero-ventral apophysis of male palpal femur (Fig. 126; single pointed process without side branch); also by apophysis on male palpal trochanter (longer and more sclerotized than in A. kinabalu and A. lambir Huber, sp. nov.; shorter than in A. indah Huber, sp. nov. and A. poring Huber, sp. nov.), by shapes of sclerites on procursus (Figs 126 – 127), and by female genitalia (Figs 128 – 129, 169 – 171; short and wide epigynum with straight posterior margin; distinctive internal structures).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	materials_examined	Material examined Holotype MALAYSIA-BORNEO: ♂, Sabah, Gaya Island, forest along small stream (6.014 – 6.018 ° N, 116.020 ° E), 30 – 80 m a. s. l., among rocks and tree buttresses, 4 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13974). Other material MALAYSIA-BORNEO: Sabah, 12 ♂♂, 13 ♀♀, 3 juvs, same data as holotype, ZFMK (11 ♂♂, 12 ♀♀; Ar 13975 - 76) and SMK (1 ♂, 1 ♀); same data, 7 ♀♀, 2 juvs, in pure ethanol, ZFMK (Bor 165).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.5, carapace width 0.9. Leg 1: 32.6 (7.6 + 0.4 + 7.6 + 14.2 + 2.8), tibia 2: 4.4, tibia 3: 2.8, tibia 4: 4.2; tibia 1 L / d: 96. Distance PME-PME 370 µm, diameter PME 105 µm, distance PME-ALE 25 µm; AME absent. COLOR. Carapace pale ochre with black lateral bands and wide brown median band including ocular area. Clypeus pale ochre with pair of brown marks at rim. Sternum medially ochre, laterally slightly darker. Legs ochre to light brown, indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); tips of femora and tibiae whitish. Abdomen grey with dorsal and lateral pattern of black and white marks; ventrally with small brown marks near spinnerets and in genital area. BODY. Habitus as in Fig. 104; ocular area slightly raised, each triad on short stalk directed toward lateral (Figs 130 – 131); carapace without thoracic furrow (Fig. 135; only dark line in anterior part); clypeus slightly more protruding than usual; sternum wider than long (0.75 / 0.55), unmodified. Gonopore with four epiandrous spigots (Fig. 143). Spinnerets as in Figs 141 – 142. CHELICERAE. As in A. lambir Huber, sp. nov. (cf. Fig. 115), with pair of proximal lateral apophyses and pair of simple distal apophyses in very lateral position; without modified hairs; without stridulatory ridges. PALPS. As in Figs 126 – 127, coxa unmodified; trochanter with short ventral apophysis with small teeth prolaterally and small prolateral branch; femur with rounded retrolatero-ventral apophysis, long pointed prolatero-ventral apophysis without side branch, with small ventral process. Tarsal organ capsulate (Fig. 134). Procursus complex (Figs 137 – 140); retrolatero-ventral process with simple tip (Fig. 136). Bulb simple, with short and wide embolus. LEGS. Without spines, with curved hairs dorsally on metatarsi 1 and 2 only (mostly on proximal half), with few vertical hairs; retrolateral trichobothrium on tibia 1 at 2.5 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 30 pseudosegments, distally fairly distinct. Male (variation) Tibia 1 in 10 other males: 7.2 – 8.1 (mean 7.6). Female In general similar to male; eye triads closer together (distance PME-PME 185 µm), not on stalks (Fig. 132); clypeus less protruding; no curved hairs on metatarsi. Tibia 1 in 11 females: 4.9 – 5.8 (mean 5.3). Epigynum short and wide plate (Fig. 144), slightly protruding, with pair of distinctive curved darker marks (Figs 128, 169). Internal genitalia as in Figs 129 and 171, with sclerotized structures but apparently without sclerotized pockets, with pair of lateral membranous pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	biology_ecology	Natural history Most specimens were found close to the ground among logs, but some (mostly juveniles) built their webs in less protected places higher on trees.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276016FFB89581FE2047F9FC48.taxon	distribution	Distribution Known from Gaya Island only (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	description	urn: lsid: zoobank. org: act: 2 D 2 D 173 E- 9 AFC- 4 CDF- 877 F- 40 C 44043 E 1 C 5 Figs 107 – 108, 145 – 149, 172 – 174	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. indah Huber, sp. nov.) by shape of prolatero-ventral apophysis of male palpal femur (Fig. 145; pointed tip much shorter), by distal cheliceral apophyses (Fig. 147; more pointed and gradually narrowing), and by female genitalia (Figs 148, 172; pair of dark lines; posterior rim curved toward posterior); from all other relatives also by strong apophysis prolateroproximally on male palpal femur (Fig. 145; present but smaller in A. indah Huber, sp. nov.).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	materials_examined	Material examined Holotype MALAYSIA-BORNEO: ♂, Sabah, Mt. Kinabalu, Poring Hot Springs, forest along Kipungit River (6.049 ° N, 116.712 ° E), 450 m a. s. l., near ground, 7 Aug. 2014 (B. A. Huber), ZFMK (Ar 13977). Other material MALAYSIA-BORNEO, Sabah: 2 ♂♂, 4 ♀♀, same data as holotype, ZFMK (Ar 13978 - 79); 1 ♀, 2 juvs, in pure ethanol, same data, ZFMK (Bor 206). – 4 ♀♀, Mt. Kinabalu, forest along Silau Silau Trail (6.010 – 6.017 ° N, 116.537 – 116.543 ° E), 1550 – 1650 m a. s. l., near ground, 6 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13980); 2 juvs, in pure ethanol, in ZFMK (Bor 210), same data. – 2 ♂♂, 1 ♀, Mt. Kinabalu, forest above Kinabalu Mountain Lodge (6.012 – 6.014 ° N, 116.534 ° E), 1570 – 1650 m a. s. l., near ground, 5 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13981); 1 ♀, 1 juv., in pure ethanol, same data, ZFMK (Bor 215). – 5 ♀♀, 2 juvs, Kinabalu N. P., 1550 m a. s. l., 3 June 1979 and 26 July 1980 (C. L. & P. R Deeleman), RMNH (2 vials).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.6, carapace width 1.1. Leg 1: 26.9 (6.4 + 0.4 + 6.4 + 11.2 + 2.5), tibia 2: 3.8, tibia 3: 2.7, tibia 4: 4.0; tibia 1 L / d: 63. Distance PME-PME 455 µm, diameter PME 105 µm, distance PME-ALE 35 µm; AME absent. COLOR. Carapace pale ochre with black lateral bands and wide brown median band including ocular area. Clypeus pale ochre with pair of brown marks at rim. Sternum medially ochre, laterally slightly darker. Legs ochre to light brown, indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); tips of femora and tibiae whitish. Abdomen grey with dorsal and lateral pattern of black and indistinct white marks; ventrally with small brown mark near spinnerets and in genital area. BODY. Habitus as in Figs 107 – 108; ocular area slightly raised, each triad on short stalk directed toward lateral; carapace without thoracic furrow (only dark line in anterior part); clypeus slightly more protruding than usual; sternum wider than long (0.75 / 0.55), unmodified. CHELICERAE. As in Fig. 147, with pair of proximal lateral apophyses and pair of simple distal apophyses in very lateral position; without modified hairs; without stridulatory ridges. PALPS. As in Figs 145 – 146; coxa unmodified; trochanter with slender ventral apophysis with very small teeth prolaterally; femur with rounded retrolatero-ventral apophysis, long prolatero-ventral apophysis with side branch, with two prolateral processes, one of them very close to trochanter. Procursus complex; retrolatero-ventral process with bifid tip. Bulb simple, with short and wide embolus. LEGS. Without spines, with curved hairs dorsally on metatarsi 1 and 2 only (mostly on proximal half), with few vertical hairs; retrolateral trichobothrium on tibia 1 at 3.5 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 30 pseudosegments, distally fairly distinct. Male (variation) Tibia 1 in 4 other males: 6.1, 6.9, 7.1, 7.5; most males with distinct white marks on abdomen. Female In general similar to male; triads closer together (distance PME-PME 200 µm), not on stalks; clypeus less protruding; no curved hairs on metatarsi; abdomen with continuous ventral dark band between epigynum and spinnerets. Tibia 1 in 12 females: 4.2 – 5.8 (mean 5.1). Epigynum large brown plate (Figs 148, 172), slightly protruding, with distinctive lighter median area bordered by dark lines (parallel or converging anteriorly), with internal sclerites visible through cuticle. Internal genitalia as in Figs 149 and 174, apparently without sclerotized pockets, with pair of lateral membranous pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	biology_ecology	Natural history The spiders were found close to the ground in small holes and cavities. They barely reacted to disturbance and were very easy to take from their webs. They share the locality with A. kinabalu, which lives higher among the vegetation.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276012FFBA9594FB9346F2F98A.taxon	distribution	Distribution Known from two localities in Mt. Kinabalu area only (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	description	urn: lsid: zoobank. org: act: 12 E 0 B 867 - 3 A 77 - 4 B 63 - 92 FF- 97 DB 7302137 C Figs 109 – 110, 150 – 162, 175 – 177	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. poring Huber, sp. nov.) by shape of prolatero-ventral apophysis of male palpal femur (Fig. 150; pointed tip much longer), by distal cheliceral apophyses (Fig. 152; wide and distally rounded), and by female genitalia (Figs 153, 175; without pair of dark lines; posterior rim curved toward anterior); from all other congeners also by long apophysis on male palpal trochanter (in A. poring Huber, sp. nov. present but more slender and shorter).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	etymology	Etymology The species name is the Malay word for ‘ beautiful’; used here as noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	materials_examined	Material examined Holotype MALAYSIA-BORNEO: ♂, Sabah, Crocker Range between Kota Kinabalu and Tambuan, S-slope, forest along river (5.783 ° N, 116.338 – 116.340 ° E), 1430 – 1480 m a. s. l., near ground, 3 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13982). Other material MALAYSIA-BORNEO, Sabah: 3 ♂♂, 5 ♀♀, 3 juvs, same data as holotype, ZFMK (Ar 13983 - 84); 1 ♀, 2 juvs, in pure ethanol, same data, ZFMK (Bor 170). – 1 ♂, 3 ♀♀, Crocker Range between Kota Kinabalu and Tambuan, N-slope, forest along river (5.834 ° N, 116.336 ° E), 1600 m a. s. l., near ground, 3 Aug. 2014 (B. A. Huber, S. B. Huber), ZFMK (Ar 13985); 2 ♀♀, in pure ethanol, same data, ZFMK (Bor 168).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.8, carapace width 1.2. Leg 1: 36.8 (8.8 + 0.4 + 8.8 + 15.3 + 3.5), tibia 2: 5.2, tibia 3: 3.5, tibia 4: 5.3; tibia 1 L / d: 84. Distance PME-PME 500 µm, diameter PME 115 µm, distance PME-ALE 35 µm; AME absent. COLOR. Carapace pale ochre with black lateral bands and wide brown median band including ocular area. Clypeus pale ochre with indistinct pair of brown marks at rim. Sternum medially ochre, laterally slightly darker. Legs ochre to light brown, indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); tips of femora and tibiae whitish. Abdomen grey with dorsal and lateral pattern of black and indistinct white marks; ventrally with small brown mark near spinnerets and in genital area. BODY. Habitus as in Fig. 109; ocular area slightly raised, each triad on short stalk directed toward lateral; carapace without thoracic furrow (only dark line in anterior part); clypeus slightly more protruding than usual; sternum wider than long (0.65 / 0.50), unmodified. CHELICERAE. As in Figs 152 and 161, with pair of proximal lateral apophyses and distinctive pair of wide distal apophyses in very lateral position, with additional pair of small processes on frontal side of distal apophyses (Fig. 160); without modified hairs; without stridulatory ridges. PALPS. As in Figs 150 – 151, coxa unmodified; trochanter with long ventral apophysis with small teeth prolaterally (Fig. 158); femur with rounded retrolatero-ventral apophysis, very long prolatero-ventral apophysis with two side branches, with small prolateral apophysis proximally close to trochanter. Procursus complex (Figs 155 – 158); retrolatero-ventral process with bifid tip. Bulb simple, with short and wide embolus (Fig. 159), weakly sclerotized. LEGS. Without spines, with curved hairs dorsally on metatarsi 1 and 2 only (mostly on proximal half), with few vertical hairs; retrolateral trichobothrium on tibia 1 at 2.5 %; prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 with ~ 35 pseudosegments, fairly distinct. Male (variation) Tibia 1 in 4 other males: 8.7, 8.8, 8.9, 9.0. Female In general similar to male; triads closer together (distance PME-PME 210 µm), not on stalks; clypeus less protruding; no curved hairs on metatarsi; abdomen with continuous ventral dark band between epigynum and spinnerets. Tibia 1 in 8 females: 6.3 – 7.2 (mean 6.7). Epigynum large brown plate, slightly protruding, with internal pockets and other sclerites visible through cuticle (Figs 153, 175). Internal genitalia as in Figs 154 and 177, with pair of membranous pockets near posterior margin.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	biology_ecology	Natural history The spiders were found very close to the ground in small holes and cavities and barely reacted to disturbance (similar only to A. poring Huber, sp. nov.; see above). They share the locality with A. kinabalu which lives higher among the vegetation.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276010FFA19588F9D4479DFD94.taxon	distribution	Distribution Known from Crocker Range only (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	description	urn: lsid: zoobank. org: act: C 0 C 92230 - 1 A 67 - 45 FC-BCC 4 - 3 B 66 E 2 E 82 EC 0 Figs 178 – 179, 186, 189 – 193, 195 – 197, 213 – 215	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. omayan) by male clypeus modification (Fig. 191; apophyses closer together), distinct ventro-distal apophysis on male palpal femur (Fig. 190; only indistinct hump in A. omayan), and shape of epigynum (Fig. 213; whitish areas smaller and wider apart). Distinguished from other congeners by bipartite retrolatero-ventral process on procursus (Fig. 190), male palpal trochanter with prolateral apophysis (Fig. 189; other species with only ventral apophysis), and pair of internal sclerotized pockets in female genitalia (Figs 192 – 193).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	etymology	Etymology Named for Philippine-born cosmopolitan artist Pacita Abad (1946 – 2004), famous for her vibrant, colorful abstract work, but also for her paintings of tropical flowers and animal wildlife.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Negros Isl., Negros Oriental Prov., Twin Lakes N. P. (9.365 – 9.368 ° N, 123.181 ° – 123.182 ° E), 850 – 950 m a. s. l., forest above Baliansasayao Crater Lake, 9 Mar. 2014 (B. A. Huber), ZFMK (Ar 13986). Other material PHILIPPINES, Negros Isl., Negros Oriental Prov.: 5 ♂♂, 13 ♀♀, same data as holotype, ZFMK (4 ♂♂, 12 ♀♀; Ar 13987 - 88) and MSU-IIT (1 ♂, 1 ♀); 1 ♀, 4 juvs, in pure ethanol, same data, ZFMK (Phi 193). – 1 ♂, Casaroro Falls (9.281 ° N, 123.208 ° E), 550 m a. s. l., forest along river below waterfall, 10 Mar. 2014 (B. A. Huber), ZFMK (Ar 13989); 1 ♀, in pure ethanol, same data, ZFMK (Phi 189).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.4, carapace width 1.4. Leg 1: 44.3 (10.6 + 0.6 + 10.6 + 19.2 + 3.3), tibia 2: 6.6, tibia 3: 4.5, tibia 4: 6.5; tibia 1 L / d: 80. Distance PME-PME 430 µm, diameter PME 140 × 155 µm, distance PME-ALE ~ 40 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including ocular area and clypeus; sternum ochre-yellow, with darker triangular mark posteriorly and dark labium; legs ochre-yellow with slightly darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, ventrally with dark mark behind gonopore. BODY. Habitus as in Fig. 178; ocular area raised, each triad on additional short hump directed toward lateral, without process below ALE (Fig. 191); carapace with very shallow median furrow in anterior part only; clypeus with distinctive pair of apophyses (Fig. 191); sternum wider than long (0.95 / 0.65), unmodified. CHELICERAE. As in Fig. 191, with pair of lateral processes proximally and pair of long lateral apophyses distally; without modified hairs; without stridulatory ridges. PALPS. As in Figs 189 – 190; coxa unmodified; trochanter with ventral and prolateral apophyses; femur with retrolateral hump, large prolateral apophysis, and ventro-distal apophysis; patella triangular in lateral view; tibia with retrolateral trichobothrium in very distal position; proximal part of procursus with bipartite retrolatero-ventral process, with complex and apparently partly hinged distal elements; bulb with only one process (weakly sclerotized embolus), distally with indistinct hump, without small knobs. LEGS. Without spines; with curved hairs on metatarsi 1 – 3; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, only distally distinct. Male (variation) Tibia 1 in six other males: 9.8 – 10.6 (mean: 10.2). Dark spot behind gonopore absent in two males. Abdomen with or without additional white spots in dorso-lateral rows. Male from Casaroro Falls with large light brown mark on sternum posteriorly. Female In general similar to male but clypeus unmodified and more homogeneously dark brown; eye triads much closer together (distance PME-PME 165 µm); with indistinct stridulatory apparatus between carapace and abdomen: small modified area medially on carapace (Fig. 186) versus barely distinguishable hairless area on abdomen. Tibia 1 in 13 females: 7.8 – 8.8 (mean: 8.1). Epigynum large sclerotized plate with pair of light lateral humps (Figs 192, 195, 213), with pair of very indistinct membranous pockets behind epigynum in weakly modified cuticle (weak transversal ridges) (Figs 192, 195 – 196). Internal genitalia as in Figs 193 and 215, with distinct pair of sclerotized pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	biology_ecology	Natural history At both localities the spiders were found in domed sheet webs close to the ground, usually in well protected dark spaces under large rocks.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827600BFFA39590FDD64053F9F2.taxon	distribution	Distribution Known from two localities on Negros Island only (type locality and nearby locality; Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	description	Figs 180 – 181, 187, 194, 198 – 212, 216 – 218	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	discussion	Note The original description was based on a single male and two females. Here we present data on new material from the type locality and a nearby locality, as well as an amended diagnosis to account for the newly described congeners.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	diagnosis	Diagnosis Distinguished from closest known relative (A. abadae Huber, sp. nov.) by male clypeus modification (apophyses wider apart; cf. Huber 2005 a: fig. 112), very indistinct ventro-distal apophysis on male palpal femur (distinct in A. abadae Huber, sp. nov., cf. Fig. 190), and shape of epigynum (whitish areas larger and closer together; Fig. 216). Distinguished from other congeners by bipartite retrolatero-ventral process on procursus (Huber 2005 a: fig. 111), male palpal trochanter with prolateral apophysis (other species only with ventral apophysis), and pair of internal sclerotized pockets in female genitalia (Fig. 194).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	materials_examined	New material examined PHILIPPINES, Luzon Isl., Benguet Prov.: 9 ♂♂, 16 ♀♀, 2 juvs, Baguio, Crystal Cave (16.396 ° N, 120.572 ° E), 1360 m a. s. l., 2 Mar. 2014 (B. A. Huber), ZFMK (8 ♂♂, 15 ♀♀; Ar 13990) and MSU-IIT (1 ♂, 1 ♀); 1 ♀, 6 juvs, in pure ethanol, same data, ZFMK (Phi 206). 3 ♀♀, near Baguio, Mt. Kabuyao, N slope (16.374 ° N, 120.557 ° E), 1200 – 1400 m a. s. l., among rocks, 2 Mar. 2014 (B. A. Huber), ZFMK (Ar 13991); 1 ♀, 5 juvs, in pure ethanol, same data, ZFMK (Phi 205).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	description	Description (amendments to Huber 2005 a) MALE. Tibia 2 slightly shorter than tibia 4 (e. g., 7.1 / 7.3); curved hairs on all tibiae and metatarsi; tibia 1 in 8 males: 9.7 – 10.9 (mean: 10.4). FEMALE. Eye triads much closer together than in male (distance PME-PME ~ 180 – 200 µm vs. 300 – 400 µm); indistinct stridulatory apparatus between carapace and abdomen: small modified area medially on carapace (Fig. 187) versus light brown hairless area on abdomen; with pair of very indistinct membranous pockets behind epigynum in unmodified cuticle (Figs 194, 218). Tibia 1 in 16 females: 7.6 – 8.5 (mean: 8.2).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	biology_ecology	Natural history The type locality is a highly degraded and polluted cave in the midst of a suburb of Baguio City. The cave is actually a natural tunnel of about 50 m length, open on both sides. Within the cave, the spiders were found in high numbers, building their typical domed sheet webs mainly along lower parts of the cave walls near the ground, in crevices and small holes. The finding of the same species among rocks on nearby Mt. Kabuyao suggests that the species is actually widespread in the area and not in danger of extinction by further degradation of the cave.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276009FFAE958DF9FC4059FD3E.taxon	distribution	Distribution Known from two localities on Luzon Island only (type locality and nearby locality; Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	description	urn: lsid: zoobank. org: act: 087 C 21 AC-F 569 - 43 EE-A 817 - 3 B 0407 A 1 C 1 EA Figs 182 – 183, 188, 219 – 223, 229, 234 – 236	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. banahaw Huber, sp. nov., A. lozadae Huber, sp. nov.) by combination of smaller retrolatero-ventral process and smaller dorso-distal sclerite on procursus (compare Figs 229 – 231) and by narrower epigynum with anterior half wider and more heavily sclerotized than posterior half (Figs 222, 234). Distinguished from A. lozadae Huber, sp. nov. also by smaller projections at ALE (Fig. 221). Distinguished from other congeners by presence of projections at ALE, by long epigynum, and by posterior membranous pockets close together (Fig. 222).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	etymology	Etymology Named for the Filipino painter and illustrator Vicente Silva Manansala (1910 – 1981), most famous for his ‘ Madonna of the Slums’.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Luzon Isl., Quezon Prov., between Lucban and Tayabas (14.063 ° N, 121.567 ° E), 330 m a. s. l., degraded forest along river, near ground, 26 Feb. 2014 (B. A. Huber), ZFMK (Ar 13992). Other material PHILIPPINES, Luzon Isl.: Quezon Prov., 5 ♂♂, 13 ♀♀, same data as holotype, ZFMK (4 ♂♂, 12 ♀♀) (Ar 13993 - 94) and MSU-IIT (1 ♂, 1 ♀); 3 ♀♀, 1 juv., in pure ethanol, same data, ZFMK (Phi 213). – Laguna Prov., 2 ♂♂, 2 ♀♀, Mt. Banahaw, forest near Taytay Falls (14.110 ° N, 121.507 ° E), 560 m a. s. l., near ground, 26 Feb. 2014 (B. A. Huber), ZFMK (Ar 13995).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.2, carapace width 1.3. Leg 1: 36.5 (8.5 + 0.5 + 8.5 + 15.5 + 3.5), tibia 2: 5.2, tibia 3: 3.5, tibia 4: 5.3; tibia 1 L / d: 67. Distance PME-PME 350 µm, diameter PME 125 × 150 µm, distance PME-ALE ~ 45 µm; AME absent. COLOR. Carapace ochre-yellow with narrow lateral marginal bands and wide dark brown median band including posterior part of ocular area; clypeus ochre yellow with indistinct light brown pattern; sternum light brown to orange, laterally paler, labium darker; legs greenish ochre with slightly darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, ventrally with dark mark behind gonopore and larger less distinct mark in front of spinnerets. BODY. Habitus as in Fig. 182; ocular area raised, each triad on additional short hump directed toward lateral, with small process below ALE (Fig. 221); carapace with very shallow median furrow in anterior part only; clypeus medially slightly projecting, with distinctive lateral plates bordered by sclerotized ridges (Fig. 221); sternum wider than long (0.85 / 0.60), unmodified. CHELICERAE. As in Fig. 221, with pair of lateral processes proximally and pair of very long lateral apophyses; without modified hairs; without stridulatory ridges. PALPS. As in Figs 219 – 220; coxa unmodified; trochanter with ventral apophysis; femur with small ventro-distal apophysis and retrolateral ridge ending in small hump; patella triangular in lateral view; tibia with retrolateral trichobothrium in very distal position; proximal part of procursus with simple retrolatero-ventral process, with complex and apparently partly hinged distal elements (Fig. 229); bulb with only one process (weakly sclerotized embolus), distally with several small knobs. LEGS. Without spines; with curved hairs on metatarsi 1 – 3; few vertical hairs; retrolateral trichobothrium on tibia 1 at 3 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, only distally distinct. Male (variation) Tibia 1 in 5 other males: 8.0 – 8.8 (mean: 8.3). Abdomen with or without additional white spots in dorsolateral rows. Female In general similar to male but clypeus unmodified and with pair of dark brown bands below ALE; eye triads much closer together (distance PME-PME 150 µm), without processes at ALE; with indistinct stridulatory apparatus between carapace and abdomen: modified area medially on carapace (Fig. 188) versus barely distinguishable hairless area on abdomen. Tibia 1 in 13 females: 5.8 – 6.5 (mean: 6.3); dark and light rings on legs often more distinct than in males. Epigynum long, anterior half of plate wider and more heavily sclerotized than posterior half, anterior half with pair of low humps (Figs 222, 234); area behind epigynum with pair of very indistinct membranous pockets in weakly modified cuticle (weak transversal ridges). Internal genitalia as in Figs 223 and 236, without sclerotized pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	biology_ecology	Natural history At both localities the spiders were found in domed sheet webs close to the ground, usually in well protected dark spaces under large rocks. The type locality suggests that the species does not depend on well preserved forests and is probably widespread in the area.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276004FFA89577FD404059F97A.taxon	distribution	Distribution Known from two localities on Luzon Island only (type locality and nearby locality; Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	description	urn: lsid: zoobank. org: act: 3526 A 886 - 2551 - 4 FBE- 900 A- 603 D 92 A 43 E 23 Figs 184 – 185, 224 – 228, 230, 237 – 239	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. manansalai Huber, sp. nov., A. banahaw Huber, sp. nov.) by combination of long projections at ALE (Fig. 226), large retrolatero-ventral process on procursus (Fig. 230; similar to A. banahaw Huber, sp. nov.), and oval shape of epigynum (Figs 227, 237). Distinguished from other congeners by presence of projections at ALE, by epigynum longer than wide, and by posterior membranous pockets close together (Figs 227 – 228).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	etymology	Etymology Named for Filipino violinist Carmencita Lozada (1940 – 2006), prize winner of the Paganini International Violin Competition in Italy.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Luzon, Camarines Sur Prov., Mt. Isarog, W slope (13.664 ° N, 123.34 – 123.35 ° E), ~ 600 – 900 m a. s. l., forest, near ground, 23 Feb. 2014 (B. A. Huber), ZFMK (Ar 13996). Other material PHILIPPINES, Luzon Isl., Camarines Sur Prov.: 3 ♂♂, 13 ♀♀, same data as holotype, ZFMK (Ar 13997 - 98); 1 ♀, in pure ethanol, same data, ZFMK (Phi 221). – 1 ♂, 1 ♀, in pure ethanol, Mt. Isarog (13.665 ° N, 123.354 ° E), 9.3 km E of Naga City, 920 m a. s. l., 31 May – 2 June 2011 (M. Yngente et al.), CAS (9042055).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.9, carapace width 1.3. Leg 1: 40.1 (9.2 + 0.5 + 9.2 + 17.7 + 3.5), tibia 2: 5.8, tibia 3: 4.0, tibia 4: 5.8; tibia 1 L / d: 71. Distance PME-PME 390 µm, diameter PME 135 × 155 µm, distance PME-ALE ~ 45 µm; AME absent. COLOR. Carapace ochre-yellow with narrow dark lateral marginal bands and wide dark brown median band including posterior part of ocular area; clypeus ochre yellow with indistinct light brown pattern; sternum light brown to orange, labium darker; legs greenish ochre with slightly darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, ventrally with dark mark behind gonopore and larger less distinct mark in front of spinnerets. BODY. Habitus as in Figs 184 – 185; ocular area raised, each triad on additional hump directed toward lateral, with long process at ALE (Fig. 226); carapace with very shallow median furrow in anterior part only; clypeus medially not projecting, with distinctive lateral plates bordered by sclerotized ridges (Fig. 226); sternum wider than long (0.85 / 0.65), unmodified. CHELICERAE. As in Fig. 226, with pair of lateral processes proximally and pair of very long lateral apophyses, without modified hairs; without stridulatory ridges. PALPS. As in Figs 224 – 225; very similar to A. manansalai Huber, sp. nov. and A. banahaw Huber, sp. nov.; procursus as in Fig. 230. LEGS. Without spines; with curved hairs on metatarsi 1 – 2; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, only distally distinct. Male (variation) Tibia 1 in 2 other males: 8.8, 9.2. Abdomen with or without additional white spots in dorso-lateral rows (white spots seem to get partly lost or indistinct in alcohol). Female In general similar to male but clypeus unmodified and most females with pair of dark brown bands below ALE; eye triads much closer together (distance PME-PME 155 µm), without processes near ALE; with indistinct stridulatory apparatus between carapace and abdomen: modified area medially on carapace (smaller than in A. manansalai Huber, sp. nov.) versus barely distinguishable hairless area on abdomen. Tibia 1 in 9 females: 6.7 – 7.7 (mean: 7.2); dark and light rings on legs often more distinct than in males. Epigynum oval, longer than wide (Figs 227, 237), anterior half with pair of low humps; area behind epigynum with pair of very indistinct membranous pockets in weakly modified cuticle (weak transversal ridges). Internal genitalia as in Figs 228 and 239, without sclerotized pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	biology_ecology	Natural history The spiders were found in domed sheet webs close to the ground, usually in well protected dark spaces under large rocks and logs, but apparently not as deeply hidden in these cavities as the sympatric A. ocampoi Huber, sp. nov.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E8276002FFD4959EF8844632FC20.taxon	distribution	Distribution Known from type locality on Luzon Island only (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	description	urn: lsid: zoobank. org: act: 6 D 270 C 60 - BBF 0 - 4 DFE- 88 BC- 1 F 90 D 87 C 5 D 8 F Figs 231 – 233, 240 – 242	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	diagnosis	Diagnosis Distinguished from closest known relatives (A. manansalai Huber, sp. nov., A. lozadae Huber, sp. nov.) by shape of procursus (large retrolatero-distal sclerite; compare Figs 229 – 231) and by pentagonal epigynum (Figs 232, 240); from A. manansalai Huber, sp. nov. also by larger retrolatero-ventral process on procursus (Fig. 231) and narrower apophysis of male palpal trochanter; from A. lozadae Huber, sp. nov. also by smaller projections at ALE (similar to A. manansalai Huber, sp. nov.; cf. Fig. 221). Distinguished from other congeners by presence of projections at ALE, by longer than wide epigynum, and by posterior membranous pockets close together (Fig. 232).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	etymology	Etymology Named for the type locality; noun in apposition.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	materials_examined	Material examined Holotype PHILIPPINES: ♂, Luzon Isl., Laguna Prov., Mt. Banahaw, forest near Taytay Falls (14.110 ° N, 121.507 ° E), 560 m a. s. l., near ground, 26 Feb. 2014 (B. A. Huber), ZFMK (Ar 13999). Other material PHILIPPINES, Luzon Isl., Laguna Prov.: 7 ♂♂, 4 ♀♀, same data as holotype, ZFMK (Ar 14000 - 01); 1 ♀, 4 juvs, in pure ethanol, same data, ZFMK (Phi 217). – 1 ♀, 1 juv., in pure ethanol, Mt. Banahaw (14.103 ° N, 121.518 ° E), 4.38 km W of Lucban, 790 m a. s. l., 16 May 2011 (H. Wood et al.), CAS (9045550).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.7, carapace width 1.35. Leg 1: 36.7 (8.6 + 0.5 + 8.6 + 15.4 + 3.6), tibia 2: 5.5, tibia 3: 3.8, tibia 4: 5.6; tibia 1 L / d: 63. Distance PME-PME 395 µm, diameter PME 140 µm, distance PME-ALE ~ 70 µm; AME absent. COLOR. Carapace ochre-yellow with narrow dark lateral marginal bands and wide dark brown median band including posterior part of ocular area; clypeus ochre yellow, small marks below each eye triad; sternum monochromous light brown, labium darker; legs greenish ochre with very indistinct darker rings on femora (subdistally, with light tip), and tibiae (proximally and subdistally, the latter followed by light tip); abdomen ochre-gray, dorsally and laterally covered with many black marks, ventrally with dark mark behind gonopore and larger, less distinct mark in front of spinnerets. BODY. Habitus very similar to A. lozadae Huber, sp. nov. (cf. Figs 184 – 185); ocular area raised, each triad on additional short hump directed toward lateral, with small process below ALE (only slightly longer than in A. manansalai Huber, sp. nov.; cf. Fig. 221); carapace with very shallow median furrow in anterior part only; clypeus with distinctive lateral plates bordered by sclerotized ridges; sternum wider than long (0.9 / 0.7), unmodified. CHELICERAE. As in close relatives (cf. Figs 221, 226), with pair of lateral processes proximally and pair of very long lateral apophyses; without modified hairs; without stridulatory ridges. PALPS. In general as in A. manansalai Huber, sp. nov. and A. lozadae Huber, sp. nov. (cf. Figs 219 – 220, 224 – 225); coxa unmodified; trochanter with ventral apophysis slightly narrower than in A. manansalai Huber, sp. nov.; femur with ventro-distal apophysis and retrolateral ridge ending in small hump; patella triangular in lateral view; tibia with retrolateral trichobothrium in very distal position; proximal part of procursus, with simple retrolatero-ventral process, with complex and apparently partly hinged distal elements; bulb with only one process (weakly sclerotized embolus), distally with one small knob. LEGS. Without spines; with curved hairs on metatarsi 1 – 3; few vertical hairs; retrolateral trichobothrium on tibia 1 at 3 %; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~ 25 pseudosegments, distally fairly distinct. Male (variation) Tibia 1 in 7 other males: 8.2 – 9.4 (mean: 8.9). Female In general similar to male but clypeus unmodified and with pair of dark brown bands below ALE; eye triads much closer together (distance PME-PME 185 µm), without processes at ALE; with indistinct stridulatory apparatus between carapace and abdomen: modified area medially on carapace versus barely distinguishable hairless area on abdomen. Tibia 1 in 4 females: 6.6, 6.8, 7.1, 7.3; dark and light rings on legs mostly more distinct than in males. Epigynum as in Figs 232 and 240, anterior large plate pentagonal, with transversal anterior bulge bordered posteriorly by shallow indentation; area behind epigynum with pair of very indistinct membranous pockets in weakly modified cuticle (weak transversal ridges). Internal genitalia as in Figs 233 and 242, without sclerotized pockets.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	biology_ecology	Natural history The spiders were found in domed sheet webs close to the ground, usually in well protected dark spaces under large rocks. Males and females were sometimes found together in one web. When disturbed, the spiders ran to the rock, vibrated only for a moment and then remained motionless, pressed against the rock surface.	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
0390E827607EFFD19564FC2A47A9F9F2.taxon	distribution	Distribution Known from the type locality only (Fig. 5).	en	Huber, Bernhard A., Nuñeza, Olga M., Ung, Charles Leh Moi (2015): Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae). European Journal of Taxonomy 162: 1-78, DOI: 10.5852/ejt.2015.162
