taxonID	type	description	language	source
0395153B9C2C4B6709D3AE90FBA7FA6A.taxon	diagnosis	Diagnosis: Length 9 – 10 mm, width 4 – 5 mm; body large, broad, 2.0 ¥ longer than wide, densely punctured but with overall surface smooth; pronotum transverse, 1.5 – 1.6 ¥ wider than long, apex deeply concave, broadest at middle, distinctly narrowed at base, punctures coarse, sparse on disc; elytra evenly punctured throughout, striae barely impressed; dorsal tooth on right mandible absent. Distribution: San Cristóbal Island: various sites around Baquerizo Moreno (Wreck Bay), playa north of Punta Lido, Playa Lobos, Playa at Bahia Rosa Blanca. Bionomics: In littoral and arid zones, from sea level to 10 m elevation, on beach sand and ash substrates, under litter of beach plants and at sea lion dung. Adults collected January, February and May. Examined: 177 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C2C4B600A5EAFD1FBDDF9CE.taxon	diagnosis	Diagnosis: Length 8 – 10 mm, width 4 – 5 mm; body large, broad, and often flattened, 2 ¥ longer than wide; pronotum transverse, 1.4 – 1.5 ¥ wider than long, coarsely, but shallowly punctured, apical margin nearly straight, widest at middle, nearly as wide at base, but sometimes constricted; elytra with striae formed by deep punctures, single irregular row of fine punctures in the intervals between the striae; dorsal tooth of right mandible present but weak. Distribution: Floreana Island, and its nearby satellite islets of Champion, and Gardner at Floreana. On Floreana it is known from: sites around Black Beach, Post Office Bay, inland from Las Cuevas, at the Ritter Farm, the Cruz Farm, Punta Cormoran and the summit of Cerro Pajas. Bionomics: The species occurs in littoral and arid zone habitats, from sea level up to 550 m on summit of Cerro Pajas. Found under shrubs, in mangrove litter, at sea lion dung, and under stones on ash and soil, and on calcareous beach sand at night. Adults collected from January to August, and October and December. Examined: 395 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C2B4B610A41AC66FE9DF96D.taxon	diagnosis	Diagnosis: Length 8 – 10 mm, width 4 – 5 mm; body large, broad, and flattened, 2 ¥ longer than wide; pronotum transverse, 1.6 ¥ wider than long, coarsely punctured, apical margin very convex, widest at middle, nearly as wide at base; elytra with striae formed by deep punctures and a double row of offset punctures in the intervals between the striae, the punctures nearly as coarse as those in the striae; dorsal tooth of right mandible weak. Distribution: Española Island: Bahia Manzanilla, Punta Suarez, Bahia Gardner. Gardner at Española Island. Bionomics. The species is found in the littoral and arid zones, from sea level to 30 m elevation, on calcareous sands, in litter under Cryptocarpus shrubs and Cordia and Prosopis trees, in seabird rookeries, in carrion and dung bait pitfall traps. Adults have been collected in February, and from April to June. Examined: 210 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C2A4B6109C7ACD9FB3CF964.taxon	diagnosis	Diagnosis: Length 7 – 9 mm, width 3 – 4 mm; body elongate, 2.2 ¥ longer than wide; pronotum nearly quadrate, 1.2 – 1.3 ¥ wider than long; pronotum widest at middle, slightly constricted at base; pronotum with coarse, shallow punctures, elytra with parallel ridges, coarsely punctured both between and on top of the ridges; dorsal tooth of right mandible present but weak. Distribution: Darwin and Wolf Islands. Bionomics: Occurring in the arid zone from sea level to 50 m elevation, in leaf litter and under low vegetation. Adults have been collected in January, February, June, September and December. Abundant in dense Croton thickets on the north plateau of Wolf Island, probably feeding on abundant sea bird guano or nest debris. Examined: 364 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C2A4B620AA9ACDFFD6AFCD6.taxon	diagnosis	Diagnosis: Length 6 – 8 mm, width 3 – 4 mm; body broad, 2.0 ¥ longer than wide; body coarsely and rugosely punctured; pronotum slightly transverse, 1.5 ¥ wider than long, widest behind middle, nearly as wide at base; elytra with rugose ridges, very coarsely punctured both between and on top of ridges, some punctures anastomosing; dorsal tooth on right mandible present, moderately well developed. Distribution: Marchena and Pinta Islands. Bionomics: The species occurs from the littoral and arid zone from sea level up to transition zone forest at 400 m elevation, and is found in litter under shrubs and in forests. Adults have been collected in February, March and September. Examined: 176 specimens. THE ‘ B GROUP’, A MONOPHYLETIC GROUP ON CENTRAL AND WESTERN ISLANDS, WHICH ARE MIDDLE AGED OR	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C294B6209C5A98FFD1EF923.taxon	diagnosis	Diagnosis: Length 7 – 9 mm, width 3 – 4 mm; body elongate, 2.4 ¥ longer than wide, surface shiny; sides of abdomen are parallel; pronotum quadrate, 1.3 – 1.4 ¥ wider than long, with fine punctures, widest at middle, narrowed toward base; elytra marked by distinct parallel ridges; fine punctures arranged irregularly in intervals between ridges, single row of coarse punctures adjacent to ridges; dorsal tooth on right mandible present, moderately well developed. Distribution: Widespread and common on Isabela (around Villamil, Volcan Alcedo, Cerro Azul) and Santa Cruz (CDRS, Academy Bay, Tortuga Bay). Bionomics: The species occurs in littoral habitats, is most abundant in arid zone habitats, especially around the Darwin Research Station, and as high as the inversion zone on the summit of Cerro Azul, from sea level to 1500 m elevation. It occurs in coastal dunes, and especially under litter on ash and basalt rock substrates. Adults have been collected in every month of the year. Examined: 569 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C294B620835AD1DFC21FC14.taxon	diagnosis	Diagnosis: Length 7 – 9 mm, width 3 – 5 mm; body broad and convex, 2.0 – 2.1 ¥ longer than wide; pronotum slightly transverse, 1.5 – 1.6 ¥ wider than long, coarsely punctured, some punctures near margins elongate, pronotum widest at middle, nearly as wide at base, sides rounded; elytra with parallel ridges, single row of weak punctures between ridges; thorax very convex, sides rounded; dorsal tooth on right mandible present, well developed. Distribution: On the widely separated, small, low islands of Pinzón, Santa Fe and Tortuga. Seemingly limited to small satellite islands south of or between Santa Cruz and Isabela. Bionomics: The species occurs in the littoral and arid zones of these small and low islands, from sea level up to 240 m on Pinzón, under litter and rocks. Adults have been collected in February, from May to July, and October to December. Examined: 150 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C294B630A59AE2FFE9DFCEC.taxon	diagnosis	Diagnosis: Length 6 – 8 mm, width 3 – 4 mm; overall body surface smooth and somewhat shiny, 2.1 ¥ longer than wide; fine sparse punctures on pronotum and elytra; pronotum strongly convex at the apex; pronotum widest at middle, roughly the same width at the base as at the apex, constricted slightly at the base, 1.5 – 1.6 ¥ wider than long; elytra smooth with fine but distinct irregularly arranged punctures; sides of abdomen parallel; dorsal tooth on right mandible present, moderately well developed. Distribution: Widespread. Fernandina (Cabo Hammond), Isabela (Punta Garcia, Tagus Cove, Urvina Bay, Volcan Alcedo, near Villamil), Santa Cruz (Tortuga Bay) and Santiago (Cerro Cowan, James Bay, Playa Espumilla, Los Guayabillos) Islands. Also Cowley, Daphne Major and Caamano islets. Bionomics: The species occurs from the littoral up to the humid forest zone, from sea level to 500 m on Volcan Wolf and 1100 m on the summit of Volcan Alcedo, both on Isabela. Collected at many sites, and in all months from January to August. Examined: 521 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C284B630A7AA827FBE1F9C3.taxon	materials_examined	Holotype: In CMN collection. Type data: Galápagos Islands, Genovesa Island. Holotype data: Darwin Bay, 26 – 28. III. 92, J. Cook and S. B. Peck, under Cryptocarpus and Bursera litter, 92 - 68. Paratypes: 99 with same data, and 200 with: Genovesa Island, Prince Phillips Steps, 27. III. 92, arid zone, 20 m, Cook and Peck, 92 - 61. Etymology: The name is a noun used in apposition, and is the name of the island to which the species is restricted. Diagnosis: Length 8 – 10 mm, width 4 – 5 mm; body broad, convex, 2.0 ¥ longer than wide; striae and intervals of elytra nearly the same height, giving this species an overall smooth surface; pronotum transverse, 1.5 ¥ wider than long, coarsely, shallowly punctured, widest behind middle, slightly constricted at base; elytral punctures coarse along striae, smaller in intervals; right mandible with well-developed dorsal tooth as in Figure 13. The etymology of the name comes from the distribution of this species, which is restricted to Genovesa Island. Distribution. Genovesa Island: Darwin Bay and Prince Philip’s Steps. Bionomics. Found in the arid zone, under rocks and in litter of Cryptocarpus shrubs and Bursera trees. Abundant in nesting areas of red-footed boobies. Adults collected from February to June and November. Examined: 341 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
0395153B9C284B63081AA946FC35FE0F.taxon	diagnosis	Diagnosis: Length 7 – 9 mm, width 4 – 5 mm; body broad, <2.0 ¥ longer than wide; overall body surface smooth; fine, sparse punctures on pronotum and elytra; pronotum apical margin nearly straight; pronotum transverse, 1.7 ¥ wider than long, noticeably narrow at apex and broadening continually toward the base; elytra smooth with punctures very fine, sometimes obsolete, arranged in parallel rows; abdomen broad and convex, giving a robust shape; dorsal tooth of right mandible present, well developed. Distribution: Widespread as the nominate subspecies (Figs 32, 33) on Baltra, Bartolomé, Beagle, Daphne Major, Eden, Rabida, Santiago, Seymour and South Plazas islands, and as S. linelli leleupi (Figs 34, 35) in the uplands and north slope of Santa Cruz Island. Not Genovesa (Tower) Island. Bionomics: The species occurs in the littoral and arid zones, on coastal dunes and in mangrove debris and shrub litter and at carrion from sea level to upper transition zone forests at 600 m elevation, on both ash and soil substrates. Adults collected in January, March to July, and November. Examined: 571 specimens.	en	Finston, Terrie L., Peck, Stewart B. (2004): Speciation in Darwin’s darklings: taxonomy and evolution of Stomion beetles in the Galápagos Islands, Ecuador (Insecta: Coleoptera: Tenebrionidae). Zoological Journal of the Linnean Society 141 (1): 135-152, DOI: 10.1111/j.1096-3642.2004.00115.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00115.x
