taxonID	type	description	language	source
0397507C833BFFE15BB76C1180FF4E3E.taxon	materials_examined	Type species. Armandia cirrhosa Filippi, 1861: 219, by monotypy.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE15BB76C1180FF4E3E.taxon	diagnosis	Diagnosis. Body elongated, not divided into distinct regions; ventral groove and two lateral grooves from chaetiger 2 to posterior end. Segments annulated. Prostomium conical, sometimes with terminal palpode; subdermal eyespots; a pair of large, eversible nuchal organs. Single and cirriform branchiae from chaetiger 2. Sometimes one or several posterior segments abranchiate. Segmental lateral eyes present between parapodia. Parapodia with prechaetal lobe, ventral lobe and sometimes a dorsal cirrus. Noto- and neuropodial lappets absent. Interramal ciliated organ present sometimes. Noto- and neuropodial small fascicles of simple capillary chaetae. Pygidium with conspicuous expansion — anal (pygidial) tube (funnel) — of variable shape, usually bearing internally attached midventral cirrus, marginal papillae and sometimes a basal pair of papillae. Diagnosis modified from Uebelacker (1984) and Blake (2000) by introducing a more detailed description of parapodial and anal tube characters, and confirming the presence of a parapodial ciliated sensorial organ in the parapodial prechaetal lobe.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE65BB76E6B83274F61.taxon	materials_examined	Material examined. Two specimens in two samples. Holotype: AM W. 44696, MI QLD 2366. Paratype: AM W. 44117, MI QLD 2422.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE65BB76E6B83274F61.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe and ventral lobe; anterior parapodial prechaetal lobe notoriously elongated and symmetrically biramous in chaetigers 1 to 3; following chaetigers lacking dorsal ramus and becoming asymmetrically uniramous towards body end. Anal tube somewhat narrower at base and slightly increasing in width towards distal end, becoming tube-like shape; tube opening slightly laterally compressed, directed posteroventrally; posterior border provided with 10 – 12 pairs of short paired cirri and an internal short anal cirrus.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE65BB76E6B83274F61.taxon	description	Description. Based on holotype. Specimen complete, 29.0 mm long and 2.5 mm wide, with 39 chaetigers. Body slender, slightly tapering towards anterior and posterior ends. Prostomium conical (Figs 2 A, 3 A) excluding palpode, longer than wide; palpode very short. A pair of small red eyes deeply embedded in prostomium. One pair of ring-shaped nuchal organs (Fig. 3 A); pharynx eversible (Fig. 2 A) provided with several oral tentacles. Branchiae present from CH 2 to last body chaetiger (CH 39), long, not decreasing in length towards posterior chaetigers, not meeting middorsally (Figs 2 A – B, 3 A – C); branchiae of last chaetigers with brown spots at midlength / distal third. Parapodia biramous, with prechaetal lobe and ventral lobe on each parapodium. Anterior parapodial prechaetal lobe notoriously elongated and symmetrically biramous in CH 1 – CH 3 (Fig. 3 A, D – E); following chaetigers lacking dorsal ramus and becoming asymmetrically uniramous towards body end (Fig. 3 F – I). Ventral ramus of prechaetal lobe with two conspicuous notches in CH 2 – CH 6 (Fig. 3 E – G). Elongated ventral lobe in all chaetigers; dorsal cirrus not present. Simple capillary chaetae in two bundles; notochaetae slightly longer than neurochaetae, of same length in all chaetigers and about as long as branchiae. Lateral eyespots anterior to parapodia on 11 chaetigers (CH 7 – CH 17), orange, horizontally oval; eyespots of CH 7 and CH 15 – CH 17 much smaller than others. Anal tube somewhat narrower at base and slightly increasing in width towards distal end, becoming tube-like shape (Figs 2 B, 3 B – C), as long as last 5 – 6 chaetigers. Anal tube opening slightly laterally compressed, directed postero-ventrally, appearing obliquely truncate in lateral view (Fig. 3 C). Posterior border provided with 10 – 12 pairs of short paired cirri, about 1 / 10 as long as anal tube, and an internal, unpaired anal cirrus, short and thin, not easily visible through tube wall (Fig. 3 C). Pair of short basal cirri not observed.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE65BB76E6B83274F61.taxon	discussion	Remarks. Armandia bifida n. sp. is characterised by: 1) the large body size and the large number of chaetigers (39); 2) the bifid shape of the prechaetal lobe in CH 1 – CH 3; 3) the presence of branchiae from CH 2 to the last chaetiger, and 4) the distinct tubular, distally-truncated shape of the anal tube. The photograph of the paratype AM W. 44696 (Fig. 2 A – B) when still alive shows the pharynx extruded with at least 12 oral tentacles of same length in a continuous series. Once the specimen was fixed, only three tentacles could be seen externally. Gallardo (1967) illustrates the oral tentacles of A. leptocirris (Grube, 1878) and A. longicaudata (Caullery, 1944) found at the Nha Trang Bay, South Vietnam. That paper was also unique in providing a key for opheliids mostly based on the shape and number of the afore-mentioned tentacles instead of using the shape of the anal tube (Gallardo 1967). However, the usefulness of oral tentacles as discriminating character at the species level is doubtful because either needs the observation of live specimens or dissection of fixed material. On the other hand, the paratype AM W. 44696 also shows the same number of chaetigers (39), bears brown spots on branchiae in the last segments and the lateral eyes have the same distribution as in the holotype. The specimen bears an internal long unpaired anal cirrus which seems to have been lost in the holotype (dotted line in Fig. 3 C).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833BFFE65BB76E6B83274F61.taxon	etymology	Etymology. The epithet bifida (L.) refers to the conspicuous bifid shape of the prechaetal lobes of the first three chaetigers. Habitat / Distribution. Intertidal (0 – 1.5 m) in sandy bottoms off Casuarina beach, in front of Lizard Island Research Station (LIRS) (Fig. 1 B).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833CFFEB5BB76F9B859B4BD9.taxon	materials_examined	Material examined. Eight specimens in four samples. Holotype: AM W. 44241, MI QLD 2373. Paratypes: AM W. 43898, MI QLD 2333 (2); AM W. 47320, MI QLD 2373 (2); AM W. 47321, MI QLD 2373, on SEM stub; AM W. 45399, MI QLD 2440; AM W. 44557, MI QLD 2330, in EtOH.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833CFFEB5BB76F9B859B4BD9.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe and ventral lobe; prechaetal lobe symmetrical, with a notch giving distal border a bilobed appearance. Anal tube barrel-shaped, straight at base, increasing width at midline and tapering again towards distal end; anal tube opening directed posteriorly, not laterally compressed; posterior border provided with 10 pairs of short paired anal cirri of different length, about 1 / 5 as long as anal tube length; internal unpaired anal cirrus and pair of basal cirri present.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833CFFEB5BB76F9B859B4BD9.taxon	description	Description. Based on holotype. Specimen complete, 8.0 mm long and 1.0 mm wide, with 29 chaetigers. Body slender, slightly tapering towards anterior and posterior ends. Prostomium conical, palpode short and clavate (Figs 4 A, 5 A). A pair of small red eyes deeply embedded in prostomium (Fig. 4 A). One pair of ring-shaped nuchal organs (Fig. 5 A – B); pharynx eversible (Fig. 4 A), oral tentacles not seen. Branchiae present from CH 2 – CH 26, long, almost reaching dorsal midline, not decreasing in length in posterior chaetigers. Parapodia biramous, with prechaetal lobe and ventral lobe (Figs 4 C – H, 5 C – D); dorsal cirrus from CH 4. Prechaetal lobe symmetrical, with a notch giving distal border a bilobed appearance (Fig. 4 C – H). Lateral eyespots anterior to parapodia on 11 chaetigers (CH 7 – CH 17); orange, horizontally oval, those of CH 7 and CH 16 – CH 17 smaller than others. Simple, very long capillary chaetae in two bundles; notochaetae generally longer than neurochaetae (Fig. 5 A, E). Anal tube barrel-shaped, straight at base, increasing width at midline and tapering again towards distal end, as long as last four chaetigers (Figs 4 B, 5 E – F). Anal tube opening directed posteriorly, not laterally compressed. Posterior border provided with 10 pairs of short paired anal cirri of different length, about 1 / 5 as long as anal tube length (Figs 4 B, 5 F); internal unpaired anal cirrus and pair of basal cirri not seen (but present in several paratypes, see below).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833CFFEB5BB76F9B859B4BD9.taxon	discussion	Remarks. The most remarkable diagnostic feature of Armandia dolio n. sp. is the unique shape of the anal tube. Observations on live specimens showed that the anal tube seems to be somewhat flexible allowing some degree of variation in its shape, i. e. sometimes looking either inflated or deflated (Fig. 2 C). The anal tube of fixed specimens does, however, always show a conspicuous inflated shape (Figs 4 B, 5 E). Paratypes measure about 6 – 10 mm in length and 0.5 – 0.9 mm in width; these specimens have the same of numbers of chaetigers and the presence of prostomial eyes as the holotype. In contrast, two paratypes (AM W. 47320 and AM W. 43898) show an unpaired anal cirrus and fewer paired anal cirri (<7). The first chaetiger and the last three ones (CH 27 – CH 29) also lack branchiae and the parapodia show a small dorsal elevation at the level of the postchaetal lobe as well (Fig. 4 C, H). Furthermore, the anal tube apparently lacks paired basal cirri. The closest species to A. dolio n. sp. is A. paraintermedia n. sp. (see below) according to general body shape but they differ in the shape and features of the anal tube. Thus, A. dolio n. sp. has a large, barrel-shaped anal tube while in A. paraintermedia n. sp. the paired anal cirri are comparatively longer and larger when compared to the size of the anal tube.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C833CFFEB5BB76F9B859B4BD9.taxon	etymology	Etymology. The epithet dolio (L.) refers to the barrel-shaped anal tube. Habitat / Distribution. Sublittoral (5 – 14 m) on muddy sand. Most specimens (87.5 %) were collected at Vicki’s reef, in front of Palfrey Island and remaining ones at NW of Watson’s Bay (Fig. 1 B).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8331FFEB5BB76BC383A14ECE.taxon	materials_examined	Material examined. One specimen in one sample. Holotype: AM W. 44115, MI QLD 2364.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8331FFEB5BB76BC383A14ECE.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe and ventral lobe; prechaetal lobe asymmetrical, with short ventrally displaced tip; ventral lobe wide, becoming longer than prechaetal lobe in posterior chaetigers. Branchiae present from chaetiger 2 to last body chaetiger, thin, much longer than chaetae, conspicuously twisted after fixation and not decreasing in length in posterior chaetigers. Anal tube funnel-like; narrow at base, increasing in width at distal end and opening directed posterio-dorsally, appearing obliquely truncate in lateral view. Posterior border with numerous (~ 25) pairs of small paired anal cirri; paired basal cirri and internal unpaired cirrus not observed.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8331FFEB5BB76BC383A14ECE.taxon	description	Description. Based on holotype. Specimen complete, 8.0 mm long and 0.8 mm wide, with 29 chaetigers. Body slender, slightly tapering towards anterior end; posterior end wide and truncated. Prostomium conical provided with a pair of small red eyes (Figs 2 D, 6 A); palpode well developed but short and narrow, weakly clavate (Fig. 6 A). One pair of ring-shaped nuchal organs (Fig. 6 A); pharynx eversible, provided with about 10 oral tentacles (only observed when alive; Fig. 2 D), hidden after fixation. Branchiae present from CH 2 to last body chaetiger (CH 29), thin, much longer than chaetae (Fig. 2 D), conspicuously twisted after fixation (Fig. 6 A), not decreasing in length in posterior chaetigers (Fig. 6 B). Parapodia biramous, with prechaetal lobe and ventral lobe on each parapodium; dorsal cirrus not seen (Fig. 6 C – F). Prechaetal lobe asymmetrical, with short ventrally displaced tip; ventral lobe wide, becoming longer than prechaetal lobe in posterior chaetigers. Lateral eyespots anterior to parapodia on chaetigers CH 7 – CH 17, orange, horizontally oval; those of CH 7 and CH 15 – CH 17 smaller than others. Simple capillary chaetae in two bundles; notochaetae generally longer than neurochaetae in posterior chaetigers (Fig. 6 B). Anal tube funnel-like; narrow at base, increasing in width at distal end (Figs 2 D, 6 B). Anal tube opening directed posterio-dorsally, appearing obliquely truncate in lateral view. Ventral side as long as about 7 chaetigers, and about twice as long as dorsal side, with V-shaped ventral incision (Fig. 6 B). Posterior border with numerous (~ 25) pairs of small paired anal cirri (Fig. 6 B); paired basal cirri and internal unpaired cirrus not observed.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8331FFEB5BB76BC383A14ECE.taxon	discussion	Remarks. This new species is unique within the genus Armandia because of the thin and extremely long branchiae, which become conspicuously twisted after fixation. The anal tube is particularly fragile and apparently deciduous because it is lacking in the holotype after fixation; the truncate shape of the posterior end in this only preserved specimen resembles that of Ophelina, namely that of Ophelina abranchiata Støp-Bowitz, 1948 (see Parapar et al. 2011, Fig. 9 a). Furthermore, the ecological distribution of A. filibranchia n. sp. differs from the other species described herein because its has only been collected on the outer barrier of the GBR.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8331FFEB5BB76BC383A14ECE.taxon	etymology	Etymology. The epithet filibranchia (L.) refers to the long and thin parapodial branchiae. Habitat / Distribution. Only one specimen was collected in sublittoral sand at 9 m depth at North Direction Island (outer reef SE Lizard Island) (Fig. 1 C).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8330FFEE5BB76E3782994D69.taxon	materials_examined	Material examined. Thirty-seven specimens in ten samples. Holotype: AM W. 44702, MI QLD 2421. Paratypes: AM W. 43896, MI QLD 2334 (4); AM W. 44103, MI QLD 2334; AM W. 44109, MI QLD 2340 (8); AM W. 44236, MI QLD 2366 (3); AM W. 47322, MI QLD 2373 (2); AM W. 44296, MI QLD 2373; AM W. 44294, MI QLD 2376 (2); AM W. 47323, MI QLD 2376 (2 on SEM stub); AM W. 44699, MI QLD 2415 (3); AM W. 45122, MI QLD 2429 (3); AM W. 45127, MI QLD 2432 (3); AM W. 47324, MI QLD 2432, in EtOH; AM W. 45126, MI QLD 2433; AM W. 45129, MI QLD 2433, in EtOH; AM W. 45133, MI QLD 2433.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8330FFEE5BB76E3782994D69.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe and ventral lobe on each parapodium; dorsal cirrus not present. Prechaetal lobes highly asymmetrical; wide and foliose from chaetiger 1 to chaetiger 3, then becoming progressively smaller towards last chaetigers; prechaetal lobe tip ventrally displaced; conspicuous bilobed appearance from midbody to last chaetigers. Anal tube square-shaped, as long as last 2 chaetigers; posterior and ventral margins open, provided with long unpaired anal cirrus, thick at base and distally tapered, projecting outwardly. Posterior border provided with 5 – 6 pairs of elongate, finger-like, paired anal cirri, almost as long as anal tube, shorter and thinner than pair of clavate basal cirri.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8330FFEE5BB76E3782994D69.taxon	description	Description. Based on holotype. Specimen complete, 9.0 mm long and 0.8 mm wide, with 27 chaetigers. Body slender, slightly tapering towards anterior end, posterior end truncated (Fig. 8 A). Prostomium conical (Figs 2 F, 7 A), palpode well-developed, large, clavate (Fig. 8 B). Eyes not seen. One pair of ring-shaped nuchal organs (Figs 7 A, 8 B); pharynx eversible, oral tentacles not seen. Branchiae present from CH 2 to last body chaetiger (CH 27), long, reaching chaetal bundle of next chaetiger, not decreasing in length in posterior chaetigers, not meeting middorsally. Parapodia biramous, with prechaetal lobe and ventral lobe on each parapodium; dorsal cirrus not present. Prechaetal lobes highly asymmetrical; wide and foliose from CH 1 – CH 3, then becoming progressively smaller towards last chaetigers; prechaetal lobe tip ventrally displaced (Figs 7 A, C – F, 8 C – D, 9 E); conspicuous bilobed appearance from midbody to last chaetigers (Figs 7 G – M, 8 E – F, 9 A – B). Lateral eyespots anterior to parapodia on 11 chaetigers (CH 7 – CH 17), orange, horizontally oval; those of CH 7 and CH 16 – CH 17 smaller than others. Simple capillary chaetae in two bundles, notochaetae generally longer than neurochaetae. Anal tube squareshaped, about as long as wide (Figs 7 B, 9 C – D, F); tube as long as last 2 chaetigers; posterior and ventral margins open, provided with long unpaired anal cirrus, thick at base and distally tapered, projecting outwardly (Fig. 7 B). Posterior border provided with 5 – 6 pairs of elongate, finger-like, paired anal cirri, almost as long as anal tube (Figs 7 B, 9 C – D, F), shorter and thinner than pair of clavate basal cirri (Figs 7 B, 9 F).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8330FFEE5BB76E3782994D69.taxon	discussion	Remarks. Armandia laminosa n. sp. is mostly characterised by the large size and the foliose shape of the prechaetal lobe in CH 1 – CH 3 (Figs 7 A, C – E, 8 D, 9 E); this feature is unique among all other species described in Lizard Island and described or reported in Australasia and the South-East Indian region as well. The anal tube is, however, not very distinctive and quite similar to that of A. paraintermedia n. sp. described herein, both considering the shape and size of the tube and the size of the anal cirri (compare Figs 7 B and 10 B). The paired anal cirri and the pair of basal cirri are similar in shape and width being the last ones slightly longer than the former. Paratypes measure 3 – 10 mm in length with 27 – 28 chaetigers. They still bear the anal tube and all cirri which suggests that those structures are not easily lost; the anal unpaired cirrus also show a conspicuous hook-like shape when is not broken (Fig. 7 B). The length of the palpophore seems highly variable, sometimes as long as the prostomium (paratype AM W. 43896). Observations on live specimens revealed that they conspicuously coil themselves when disturbed (Fig. 2 E, F).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8330FFEE5BB76E3782994D69.taxon	etymology	Etymology. The epithet laminosa refers to the foliose shape of the parapodial prechaetal lobes of the first three chaetigers (CH 1 – CH 3). Habitat / Distribution. All specimens were found in the West coast of Lizard Island, mostly in front of Casuarina Beach (59.5 %) and Vicki’s Reef (32.4 %), from the intertidal to 15 m depth in several types of sediments, mainly in sand but also associated with Halophila seagrass (Fig. 1 D).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8334FFF15BB76D9381E24C6D.taxon	materials_examined	Material examined: Thirty-seven specimens in sixteen samples. Holotype: AM W. 44243, MI QLD 2370. Paratypes: AM W. 43901, MI QLD 2337; AM W. 44107, MI QLD 2355; AM W. 44113, MI QLD 2356; AM W. 44112, MI QLD 2360; AM W. 47325, MI QLD 2370 (2); AM W. 44238, MI QLD 2374; AM W. 45131, MI QLD 2379, in EtOH; AM W. 44546, MI QLD 2381, in EtOH; AM W. 44550, MI QLD 2389; AM W. 44556, MI QLD 2397 (3); AM W. 45130, MI QLD 2432, in EtOH; AM W. 45217, MI QLD 2440 (15); AM W. 47326, MI QLD 2440 (2 on SEM stub); AM W. 45218, MI QLD 2440 (3 in EtOH); AM W. 45409, MI QLD 2444 (2).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8334FFF15BB76D9381E24C6D.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe, ventral lobe and small dorsal cirrus on each parapodium; prechaetal lobe asymmetrical provided with a ventrally displaced tip. Anal tube about as long as wide, squareshaped; opening at posterior margin with a long ventral incision, provided with one long internal anal cirrus, thick at base and distally tapered, projecting outwardly; posterior margin provided with 3 pairs of elongate, finger-like, paired anal cirri, almost as long as total anal tube length, thinner than pair of basal cirri.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8334FFF15BB76D9381E24C6D.taxon	description	Description. Based on holotype. Specimen complete, 7.0 mm long and 0.5 mm wide, with 29 chaetigers. Body slender, slightly tapering towards anterior end and truncated at posterior end (Fig. 11 A). Prostomium conical provided with a pair of small red eyes and one larger dorsal one (Fig. 10 A), palpode well-developed but short (Figs 10 A, 11 B). One pair of ring-shaped nuchal organs (Fig. 10 A); pharynx eversible, oral tentacles not seen (Fig. 11 C – E). Branchiae present from CH 2 – CH 26; last three chaetigers (CH 27 – CH 29) abranchiate; branchiae long surpassing parapodium of following chaetiger (Fig. 11 F), slightly decreasing in length and width in CH 24 – CH 26. Parapodia biramous, with prechaetal lobe, ventral lobe and small dorsal cirrus on each parapodium. Prechaetal lobe asymmetrical provided with a ventrally displaced tip present from CH 1 – CH 29 (Figs 10 D – I, 12 A – C). Lateral eyespots anterior to parapodia on CH 7 – CH 17, orange, circular; those of CH 16 and specially CH 17 smaller than others. Simple capillary chaetae in two bundles, notochaetae generally longer than neurochaetae, those of posterior chaetigers very long (Figs 10 C, 12 D – E). Anal tube about as long as wide, square-shaped (Figs 10 B, 12 D – E); as long as last 2 chaetigers; opening at posterior margin with a long ventral incision (Fig. 10 C), provided with one long internal anal cirrus, thick at base and distally tapered, projecting outwardly (Fig. 10 B, not present in holotype). Anal tube posterior margin provided with 3 pairs of elongate, finger-like, paired anal cirri, almost as long as total anal tube length (Figs 10 B, 12 E – F), thinner than pair of basal cirri (Figs 10 B – C, 12 E).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8334FFF15BB76D9381E24C6D.taxon	discussion	Remarks. Examination of paratypes shows some variability for several characters. Lateral eyes are present from CH 7 to CH 15 – CH 17, being smaller the first and the last 1 – 2. The chaetae from the last three chaetigers appear more deciduous than others. The number of papillae on the anal tube ranges from 1 – 3 pairs in small specimens (4 mm in length) to 11 pairs in larger specimens (8 mm). The pair of basal cirri may be short and therefore difficult to distinguish from the other cirri (Fig. 12 D – F). The eversible pharynx has digitiform protuberances which bear abundant ciliature and projects beyond the border of the mouth (Fig. 11 D – E). The branchiae are present from CH 2 to CH 27, showing two abranchiate chaetigers rather than three. In abranchiate chaetigers, there is a small dorsal elevation in the postchaetal lobe in the same place where the branchia are present in branchiate chaetigers (Fig. 10 D, I), similarly as it happens in A. dolio n. sp. The shape and size of the anal tube of Armandia paraintermedia n. sp. (Fig. 10 B) is similar to that of other species of Armandia (Figs 13, 17 A, C); some of these species have been reported in Australia but their type localities are far away such as it happens for A. intermedia Fauvel, 1902 (Fig. 13 B) and A. maculata Webster, 1884 (Fig. 13 F). Other related species from SE Asia are A. cf. melanura Gravier, 1905 (Fig. 13 G), A. amakusaensis (Fig. 13 A) and A. lanceolata Willey, 1905 (Fig. 13 E). Armandia intermedia was described by Fauvel (1902) from West Africa and later reported in South Africa (Day 1967; Fig. 13 C) and Australia (Hutchings 2000; Fig. 13 D); this species bears an anal tube similar to that of A. paraintermedia n. sp. which has paired cirri, one pair of basal cirri and ventral unpaired cirrus (“ 2 grosses papilles ventrales courtes, séparées par une longue papille impaire, médiane, ventrale ”; Fauvel 1902). Armandia intermedia differs, however, from A. paraintermedia n. sp. by having the last three chaetigers which are abranchiate instead of two, lateral eyes in 13 chaetigers (CH 7 – CH 19) instead of 11 (CH 7 – CH 17) and a constant number of paired cirri (9 pairs) instead of 1 – 11 pairs. Specimens from South Africa as described by Day (1967) have 27 – 29 chaetigers, branchiae start in CH 2, the last 23 chaetigers are abranchiate, lateral eyes are present in 12 chaetigers (CH 7 – CH 18), the unpaired anal cirrus is comparatively longer (see Fig. 13 C) and shows a high degree of variability in the number of paired anal cirri (10 – 20; “ dorsal clavate papillae ” sensu Day (1967 )). Day (1967) does not mention the presence of the pair of ventral cirri but they seem to be present according to the drawing provided in that paper (Day 1967, Fig. 25.2. g; see also Fig. 13 C).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C8334FFF15BB76D9381E24C6D.taxon	etymology	Etymology. The name of the new species refers to its morphological similarity to Armandia intermedia Fauvel, 1902 from West Africa (see Remarks below). Habitat / Distribution. Armandia paraintermedia n. sp. was the most abundant and widespread opheliid at Lizard Island, and was found both all around LI (89.2 %) and outer sites (10.8 %) (Fig. 1 E). Present from the intertidal to 24 m depth on a wide variety of bottom types (sand, coral rubble, calcareous algae, sponges and coral sand).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C832BFFF55BB76C9E80F048B8.taxon	materials_examined	Material examined. Seven specimens in three samples. Holotype: AM W. 47327, MI QLD 2370. Paratypes: AM W. 47328, MI QLD 2337; AM W. 47329, MI QLD 2340; AM W. 47330, MI QLD 2370 (3); AM W. 47331, MI QLD 2370, on SEM stub.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C832BFFF55BB76C9E80F048B8.taxon	diagnosis	Diagnosis. Parapodia biramous, with prechaetal lobe, ventral lobe and dorsal cirrus on each parapodium; prechaetal lobe short and asymmetrical, with a ventrally displaced tip. Anal tube long and narrow, tube-like. Tube opening directed posterio-ventrally, appearing obliquely truncate in lateral view; not laterally compressed; posterior border provided with 6 pairs of finger-like anal cirri of about 1 / 5 as long as dorsal tube length; paired basal cirri and internal unpaired anal cirrus.	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C832BFFF55BB76C9E80F048B8.taxon	description	Description. Based on holotype. Specimen complete, 11.0 mm long and 1.0 mm wide, with 29 chaetigers. Body slender, slightly tapering towards anterior end and truncated at posterior end. Prostomium conical, provided with a pair of lateral, small red eyes and a dorsal one (Fig. 14 A); palpode well-developed but short and narrow, twisted (Figs 14 A, 15 A). One pair of ring-shaped nuchal organs (Fig. 15 B); pharynx eversible (Fig. 14 A), oral tentacles not seen (but present in paratypes; Fig. 15 C). Branchiae present from CH 2 – CH 26; last three chaetigers (CH 27 – CH 29) abranchiate; branchiae short, slightly surpassing parapodium of following chaetiger, not decreasing in length in posterior chaetigers. Parapodia biramous, with prechaetal lobe, ventral lobe and dorsal cirrus on each parapodium (Figs 14 D – I, 15 E – F, 16 A – D). Prechaetal lobe short and asymmetrical, with a ventrally displaced tip. Lateral eyespots anterior to parapodia on CH 7 – CH 17, orange, horizontally oval; those of CH 7 and CH 15 – CH 17 smaller than others. Simple capillary chaetae in two bundles, notochaetae generally longer than neurochaetae. Anal tube long and narrow, tube-like, of same width along its length (Figs 14 B, 16 E – F). Anal tube opening directed posterio-ventrally, appearing obliquely truncate in lateral view; not laterally compressed; ventrally as long as about 2 chaetigers, dorsally as long as about 3 chaetigers (Figs 14 C, 16 E). Posterior border provided with 6 pairs of finger-like anal cirri of about 1 / 5 as long as dorsal tube length (Fig. 14 B); paired basal cirri and internal, unpaired anal cirrus not observed (but present in one paratype, see below).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C832BFFF55BB76C9E80F048B8.taxon	discussion	Remarks. The shape of the anal tube of A. tubulata n. sp. resembles that of A. leptocirris Grube, 1878 and A. andamana Eibye-Jacobsen (2002). Armandia leptocirris was described from the Philippine Islands, and later reported in Sri Lanka (Willey 1905), the Solomon Islands (Gibbs 1971) and South Africa (Day 1967, Fig. 25.2. h, as A. leptocirrus). The drawing of A. leptocirris presented by Day (1971) (Fig. 17 F) shows an anal tube long and obliquely truncated in lateral view, quite similar to that of A. tubulata. Nevertheless, in A. leptocirris the anal tube is opened posterio-dorsally instead of postero-ventrally. Armandia andamana has not been reported since it was described by Eibye-Jacobsen (2002). This species is similar to A. longicaudata but it lacks an unpaired anal cirrus which is present instead in A. longicaudata and A. tubulata n. sp. The paratypes of A. tubulata n. sp. measure 9 – 11 mm in length, and show that the size and the number of chaetigers are constant. One paratype (AM W. 47329) has an internal unpaired ventral cirrus. The branchiae are lacking in the last three chaetigers; those and the first chaetiger show a small dorsal elevation in the postchaetal lobe, as it happens in A. dolio n. sp. and A. paraintermedia n. sp. (Fig. 14 D, I).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
0397507C832BFFF55BB76C9E80F048B8.taxon	etymology	Etymology. The epithet tubulata (L.) refers to the tubular shape of the anal tube. Habitat / Distribution. A sublittoral species only found in SW LI coast, and almost restricted to Vicki’s Reef sampling sites (85.7 %) (Fig. 1 F). Found from 1 to 10 m depth on different bottom types (dead coral rubble, fine sand and calcareous algae).	en	Parapar, Julio, Moreira, Juan (2015): Six new species of the genus Armandia Filippi, 1861 (Polychaeta, Opheliidae) from Lizard Island (Great Barrier Reef, Australia). Zootaxa 4019 (1): 577-603, DOI: 10.11646/zootaxa.4019.1.19
