identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039787AE392A8462FF6A5CC60CF2F8B2.text	039787AE392A8462FF6A5CC60CF2F8B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neustanthus Bentham, Pl. Jungh.	<div><p>Neustanthus Bentham, Pl. Jungh. 2: 234–235. 1852</p> <p>Type species: — Neustanthus phaseoloides (Roxburgh) Bentham, Pl. Jungh. 2: 235. 1852.</p> <p>Diagnosis: —Perennial herbaceous climber or creeper. Stems hirsute with spreading hairs. Leaves pinnately trifoliolate. Leaflets entire or lobed. Stipules basifixed, open and reflexed. Nodose (hereinafter refers to a highly compacted inflorescence branch or brachyblast; e.g. Figure 1P) pseudoracemes axillary, several flowers clustered at each node. Calyx 5-lobed, the lower lobe longer, the upper two connate from the middle. Corolla blue or purple or white with purple to pink stripe. Vexillum without callosities, apex slightly emarginate, base with incurved auricles. Vexillary stamen connate to the tube. Ovules many. Fruits subterete, valves twisting upon dehiscence. Seeds barrel-shaped to oblong-elliptic.</p> <p>Description: —Perennial herbaceous climber or creeper. Stems slender but strong, fibrous, up to 10 m long, rooting at the nodes, pubescent with spreading hairs. Stipules basifixed, triangular to ovate, 4–12 mm × 2–3 mm, acuminate, deciduous or persistent. Leaves pinnately trifoliolate; petiole 3–12 cm long, somewhat pubescent with adpressed and spreading hairs; leaflets broadly ovate, rhomboid, or ovate-rhomboid, the terminal one broader and symmetrical, entire or lobed, 3–18 cm × 2–16 cm, margin entire or sinuate, lateral leaflets sometimes oblique, 3–14 cm × 3–12 cm, mucronate, leaflets adpressed pubescent above, more densely below, veins conspicuous below, strongly pubescent, 3 from the base, 5 pairs of primary lateral veins, opposite or not; petiolules 2–6 mm long with spreading hairs; stipels 3–10 mm long, linear to lanceolate, setaceous. Inflorescences solitary, axillary pseudoracemes, unbranched, (4–)10– 35(–45) cm long, nodose (with brachyblasts), flowering above the middle with 4 or more flowers per node; bracts subtending the nodes, 2–5 mm long, caducous; pedicels 2–6 mm, pubescent; bracteoles 2 per flower, lanceolate, 3–4 mm × 1–2 mm. Calyx 4–9 mm long, hirsute on the outside, the tube 3–5 mm long, 5-lobed, the upper 2 lobes fused along half of their length or more but not entirely fused, 2–4 mm long, lateral lobes obtuse to acute, 1.5–3 mm long, the lower lobe acute to lanceolate-acuminate or subulate, longer than the others, 3–6 mm long. Corolla bluish-purple or white suffused with purple, pink, or bluish-purple, vexillum obovate, often with green or yellow patch at the base, 10–23 mm × 8–18 mm, base with 2 incurved auricles, without callosities; wing petals obovate-oblong, bluish-purple or white suffused with bluish-purple, slightly longer than the keel, 9–20 mm × 3–6 mm, one side of base with rounded auricle, claw slender, 3–4 mm long, basal margin lobed; keel petals falcate, bluish-purple or white with blue, green or purple tip, 10–21 mm long, base truncate, clawed, the claw 4–6 mm long. Ovary linear, hirsute, 10–15 mm long with ca. 20 ovules; style glabrous, 4–10 mm long, curving upward; stigma terminal, globose. Stamens diadelphous, the vexillary stamen adherent near the middle to the staminal column in bud but becoming free with age, 10–21 mm long; anthers basifixed, alternating on long and short filaments. Fruits leguminous pods, subterete, grey to black, first adpressed hirsute, glabrescent, 5–12 cm × 3–5 mm, valves twisting upon dehiscence, with papery partitions between the seeds. Seeds 15–20 per fruit, barrel-shaped to oblong-elliptic, 2.5–4 mm × 2–3 mm, 1.5–2 mm thick; funicle short, deltoid; arils elongate.</p> <p>Phenology: —Flowering and fruiting various, dependent upon locality (see Table 2 of van der Maesen 1985 for detailed information).</p> <p>Distribution: —Native to southern, eastern, and Southeast Asia: Australia, Bangladesh, Bhutan, Brunei, Cambodia, China, India, Indonesia, Laos, Malaysia, Myanmar, N. Borneo, Nepal, New Guinea, Philippines, Singapore, Solomon Islands, Sri Lanka, Taiwan, Thailand, Vietnam. Introduced widely to other tropical areas.</p> <p>Vernacular:—ΞṘḍƀ san lie ye ge (Chinese), thua sian pa (Thai), phak phit (Thai).</p> <p>Discussion: — Neustanthus is phylogenetically close to Sinodolichos (Egan et al., in prep.; Cagle 2013), a genus of similar geographic distribution. Both genera are perennial, herbaceous climbers or creepers, with spreading hirsute hairs and basifixed stipules. Both have long, acuminate calyx lobes and linear, somewhat compressed, subterete fruit. However, they differ in their inflorescence structure, with Neustanthus having a pseudoraceme whereas Sinodolichos has a true raceme. Neustanthus has ca. 20 seeds per pod that are usually barrel-shaped whereas Sinodolichos has 3–10 seeds that are oblong. The most recent treatment of P. phaseoloides by van der Maesen (2002) recognized three varieties: var. phaseoloides, var. javanica, and var. subspicata. These taxa have been variably recognized at the species level (e.g. as Neustanthus javanicus by Bentham (1832)), varietal levels (e.g. van der Maesen, 1985, 1994, 2002), or in synonymy under P. phaseoloides. A comprehensive sampling coupled with morphometric and/or molecular genetic work will be necessary to determine the best level at which to recognize these taxa. Pending further investigation, we choose to continue with the current recognition of these taxa at the varietal level. Resurrected names and new combinations are presented below. For a full list of synonyms and a key to the varieties and descriptions thereof, see van der Maesen (1985).</p> </div>	https://treatment.plazi.org/id/039787AE392A8462FF6A5CC60CF2F8B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392D8464FF6A5ADF0E43FDBB.text	039787AE392D8464FF6A5ADF0E43FDBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neustanthus phaseoloides var. phaseoloides (Roxburgh) Bentham	<div><p>Neustanthus phaseoloides var. phaseoloides (Roxburgh) Bentham in Miquel, Pl. Jungh. 2: 235. 1852.</p> <p>TYPE: — INDIA. Calcutta Bot. Garden, grown from seeds received from Kerr at Canton, China (holotype: CAL).</p> <p>Basionym: — Dolichos phaseoloides Roxburgh, Fl. Ind., ed. 1832, 3: 316. 1832. Dolichos phaseoloides Roxburgh, Hort. Bengal.: 55. 1814, nom. nud.</p> <p>Selected Synonyms: — Pueraria phaseoloides (Roxburgh) Bentham, J. Linn. Soc. Bot. 9: 125. 1865.</p> <p>Images: —Illustration: Figure 1; Photo Plate: Figure 2.</p></div> 	https://treatment.plazi.org/id/039787AE392D8464FF6A5ADF0E43FDBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392E8467FF6A5A900954FE0A.text	039787AE392E8467FF6A5A900954FE0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neustanthus phaseoloides var. javanica (Bentham)	<div><p>Neustanthus phaseoloides var. javanica (Bentham) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — INDONESIA. Java, Merapi, R. Kuning, Junghuhn s.n. (holotype: K, not seen; Verdcourt (1971) describes Roxburgh drawing 1890 as a syntype at K, designated as a holotype by van der Maesen (1985)).</p> <p>Basionym: — Neustanthus javanicus Bentham in Miquel, Pl. Jungh. 2: 235. 1852.</p> <p>Selected Synonyms: — Pueraria javanica (Bentham) Bentham, J. Linn. Soc. Bot. London 9: 125. 1865. Pueraria phaseoloides var. javanica (Bentham) Baker in Hooker, Fl. Brit. India 2: 199. 1876.</p> </div>	https://treatment.plazi.org/id/039787AE392E8467FF6A5A900954FE0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392E8467FF6A5BD40991FD46.text	039787AE392E8467FF6A5BD40991FD46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neustanthus phaseoloides var. subspicata (Bentham)	<div><p>Neustanthus phaseoloides var. subspicata (Bentham) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — BANGLADESH. Mountains near Sylhet, Wallich 5557A (lectotype: K[barcode K000264080!]; isolectotypes: E[barcode E00301515!], G[barcode G00370593!]).</p> <p>Basionym: — Neustanthus subspicatus Bentham in Miquel, Pl. Jungh. 2: 234. 1852.</p> <p>Selected Synonyms: — Pueraria subspicata (Bentham) Bentham, J. Linn. Soc. Bot. London 9: 125. 1865. Pueraria phaseoloides var. subspicata (Bentham) Maesen, Agr. Univ. Wageningen Papers 85(1): 84–88. 1985.</p> </div>	https://treatment.plazi.org/id/039787AE392E8467FF6A5BD40991FD46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392E8466FF6A59180829FD46.text	039787AE392E8466FF6A59180829FD46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Teyleria Backer, Jard. Bot. Buitenzorg Bul.	<div><p>Teyleria Backer, Jard. Bot. Buitenzorg Bul. (ser 3) 16:107. 1939</p> <p>Type species: — Teyleria koordersii (Backer) Backer, Bull. Jard. Bot. Buitenzorg ser. 3 16:108. 1939 syn nov. (Teyleria tetragona (Merrill) Maesen, Agric, Univ, Wageningen Papers 85,1: 119. 1985.)</p> <p>Diagnosis: —Vine or scandent shrub, perennial. Stems angular to terete, sulcate in upper parts. Leaves pinnately trifoliolate. Leaflets entire. Stipules basifixed. Petioles angular to canaliculate. Nodose pseudoracemes axillary, rachis 4-angled, bract at the base of node persistent, pair of bracteoles at the base of the calyx. Calyx 4- or 5-lobed, the top two connate for more than half their length or entirely connate. Corolla white or bluish-purple. Stamens monadelphous, vexillary stamen connate to the tube in the middle. Fruits compressed, septate between the seeds. Seed coat tuberculate.</p> <p>Description: —Herbaceous or woody climber or scandent shrub, perennial. Young stems angular, pubescent, the older, woody stems becoming terete and glabrate. Stipules basifixed, lanceolate to broadly lanceolate, persistent. Leaves pinnately trifoliolate, petiole angular to canaliculate, pubescent; terminal leaflet rhomboid to ovate, lateral leaflets smaller, obliquely ovate, apices acute to acuminate, bases rounded to cuneate, veins prominent below, not in pairs, 4–6 laterals on each side; petiolules 2–7 mm long, pubescent; stipels lanceolate, persistent. Inflorescence either a terminal panicle or more commonly an axillary pseudoraceme with the rachis single or with one branch, peduncle and rachis strongly angular, pubescent, more densely hairy on the angles, nodose with thickened, condensed racemules persistent on the rachis, 3 or more flowers per node; bracts subtending nodes, lanceolate, curved upward towards apex of inflorescence, persistent; bracts subtending pedicels, lanceolate, curved upward towards apex of inflorescence, persistent to late-caducous; pedicels 1–3 mm long, pubescent; bracteoles 2 per flower, attached at base of the calyx. Calyx pubescent, 4- or 5-lobed, upper 2 lobes entirely or nearly entirely connate from the base. Corolla bluish-purple, white, or white suffused with purple; vexillum obovate to ovate, apex obtuse to emarginate, wing petals obovate, clawed; keel petals clawed. Ovary elongate, pubescent or not; style glabrous, stigma terminal, globular, pubescent at base. Stamens monadelphous, the vexillary stamen connate to staminal column in middle, free below; anthers basi-dorsifixed, alternately on long and short filaments. Fruits leguminous pods, flattened-oblong, glabrous to pubescent, apex acuminate, apiculate with persistent style, base cuneate, sutures thickened, septate. Seeds ovoid to nearly quadrate with rounded edges, compressed, reddish-brown to black, seed coat finely tuberculate; funicle deltoid; aril elongate.</p> <p>Distribution: — Cambodia, China (Yunnan and Hainan provinces), Indonesia, Laos, Malaysia, Myanmar, Thailand, and Vietnam.</p> <p>Discussion: — Teyleria stricta contains canavanine, a free amino acid not usually found in subtribe Glycininae, to which T. stricta is allied (Lackey, 1977a). Because of this, Lackey (1977b) suggested that T. stricta was anomalous in the genus Pueraria and likely allied with Neonotonia J.A.Lackey (J. Lackey, 1977). Van der Maesen (1985) acknowledged Lackey’s opinion, but disagreed with its removal from the genus, stating “morphologically the species fits better in Pueraria, as differences in habit, inflorescence size, calyx shape, flower size and shape, pod size and shape separate it from Neonotonia wightii (Arnott) J.A.Lackey.” Phylogenetic affiliations support Lackey’s hypothesis of a relationship between T. stricta and Neonotonia (Lackey, 1977b), with a clade comprised of Neonotonia and Teyleria, including T. stricta nested within Teyleria, strongly supported by both nuclear and chloroplast data (Egan et al., in prep.). This clade is supported by chemistry via the presence of canavanine in a subtribe otherwise devoid of it (Bell, Lackey, &amp; Polhill, 1978; Lackey, 1977 a, 1977b). Pueraria stricta is here transferred to Teyleria, a group sharing morphological features including angular petioles, quadrangular stems, small flowers along an erect rachis, monadelphous stamens, septate fruit, and a sculptured seed coat.</p> <p>Following the advice of (1977b), van der Maesen (1985) moved Pueraria tetragona Merrill from Pueraria to the genus Teyleria, creating the new combination Teyleria tetragona (Merrill) J.A.Lackey ex Maesen, stating in his key that T. tetragona has fruits 4–7 cm × 0.5 cm, whereas T. koordersii has fruits ca. 3 cm × 0.3 cm. However, in our review of several specimens, including all of the specimens examined by van der Maesen (1985), for Teyleria, we note that fruits from T. koordersii rarely approach 5 mm in width, but are more usually 4 mm. Furthermore, fruits range from 3–5 cm in length, with most between 3–4 cm for both T. koordersii and T. tetragona. In reality, no character states delineate the two. Based on the original descriptions of the two species as well as comparison of the holotype and isotypes of T. koordersii and isotypes (holotype not seen) of T. tetragona, we assert that the two entities are conspecific. Even though the type of the genus is T. koordersii Backer (1939) based on Glycine koordersii Backer (1911), Pueraria tetragona Merrill (1910) is the earliest name at specific rank within the genus Teyleria, a name later synonymized under Teyleria as T. tetragona (Merrill) Maesen (1985). We therefore recognize T. koordersii as a synonym of T. tetragona.</p> </div>	https://treatment.plazi.org/id/039787AE392E8466FF6A59180829FD46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392F8466FF6A5F260E43F904.text	039787AE392F8466FF6A5F260E43F904.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Teyleria stricta (Kurz)	<div><p>Teyleria stricta (Kurz) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — MYANMAR. Pegu, Kurz 2557 (lectotype: CAL; isolectotype: K[barcode K000264084]).</p> <p>Basionym: — Pueraria stricta Kurz, J. Asiatic Soc. Bengal, Pt. 2, Nat. Hist. 42-2: 254. 1873.</p> <p>Selected Synonyms: — Pueraria brachycarpa Kurz, J. Asiatic Soc. Bengal 42-2: 232,254. 1873. Pueraria hirsuta Kurz, J. Asiatic Soc. Bengal 42-2: 254. 1873. Pueraria collettii Prain, J. Asiatic Soc. Bengal 66-2: 420. 1897 (lectotype: CAL; isolectotype: K[barcode K000797408!]). Pueraria siamica Craib, Kew Bull. 1911 (1):40–41. 1911 (holotype: K[barcode K000797404!]); isotypes: E[barcode E00275914!], K[barcodes K000797402!, K000797403!], TCD[barcode TCD0016063!]). Pueraria longicarpa Thuan, Adansonia Ser. 2, 16-4: 509. 1977 (holotype: P[barcode P00507989]).</p> <p>Images: —Illustration: Figure 3; Photo Plate: Figure 4.</p></div> 	https://treatment.plazi.org/id/039787AE392F8466FF6A5F260E43F904	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE392F8468FF6A5CDA0E9EFE2E.text	039787AE392F8468FF6A5CDA0E9EFE2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Teyleria tetragona (Merrill) Maesen, Agric. Univ. Wageningen Papers	<div><p>Teyleria tetragona (Merrill) Maesen, Agric. Univ. Wageningen Papers 85(1): 119. 1985.</p> <p>TYPE: — PHILIPPINES. Palawan, near Puerto Princesa, Bermejos, Bur. Sci. 295. January 1906.</p> <p>Basionym: — Pueraria tetragona Merrill, Philipp. J. Sci. 5: 122. 1910 (holotype: PNH; isotype: NY[barcode NY00026869!], US [barcode US 00004642!]).</p> <p>Selected Synonyms: — Glycine koordersii Backer syn. nov., Schoolflora voor Java 358. 1911. Teyleria koordersii (Backer) Backer syn. nov., Bulletin du Jardin Botanique de Buitenzorg, sér. 3, 16: 108. 1939 (holotype: BO; isotypes: L[barcodes L0906816!, L0906817!, L0906818!]; paratypes: Koorders 28959B L[barcode L0906820!], Beumee 1799 L[barcode L0906822!], Koorders 21260B L[barcode L0906817!]). Glycine hainanensis Merrill &amp; P.F.Metcalf, Lingnan Sci. Jour. 16: 194. 1937 (holotype: SYS[barcode SYS00040713!]; isotypes: NY[barcode NY00011891!], BM[barcode BM001118522!]; paratypes: Liang 65132 PE[barcode PE00217043], Liang 65132 IBSC[barcode IBSC0734204!], Liang 66411 ISBC[barcodes ISBC0197748!, ISBC0197749!], DAO, not seen).</p> </div>	https://treatment.plazi.org/id/039787AE392F8468FF6A5CDA0E9EFE2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE3923846AFF6A5A900D9BFE9C.text	039787AE3923846AFF6A5A900D9BFE9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haymondia Egan & Pan 2015	<div><p>Haymondia A.N.Egan &amp; B.Pan, gen. nov.</p> <p>Type species: — Haymondia wallichii (de Candolle) A.N.Egan &amp; B.Pan (Pueraria wallichii de Candolle, Ann. Sci. Nat. (Paris) 4: 97. 1825.)</p> </div>	https://treatment.plazi.org/id/039787AE3923846AFF6A5A900D9BFE9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE3923846CFF6A5B6409D6FC12.text	039787AE3923846CFF6A5B6409D6FC12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haymondia wallichii (de Candolle)	<div><p>Haymondia wallichii (de Candolle) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — NEPAL. 1821, Wallich 5353a (holotype: G!; isotypes: BM[barcode BM000958610!], C[barcodes C 10012332!, C 10012333!, C 10012334!], K[barcode K001120653!]).</p> <p>Basionym: — Pueraria wallichii de Candolle, Ann. Sci. Nat. (Paris) 4: 97. 1825.</p> <p>Selected Synonyms:— Pueraria composita Graham ex Wallich nom. nud., Cat. Herb. Ind. no. 5570, based on Burma, Taong Dong, Wallich 5570. Neustanthus wallichii (de Candolle) Bentham, in Miquel, Pl. Jungh. 2: 234. 1852. Pueraria wallichii de Candolle var. composita (Graham ex Wallich) Bentham, J. Linn. Soc. Bot. London 9: 124. 1867 (holotype: K[barcode K001121317!]; isotypes: BM[barcode BM000958611!], G[barcodes G00370594!, G00370591!], K[barcodes K000264070!, K000264071!]). Dolichos frutescens Hamilton, in Don, Prodr. 240. 1825.</p> <p>Images: —Illustration: Figure 5; Photo Plate: Figure 6.</p> <p>Diagnosis: —Scandent shrub, erect, or climbing when in shade or associated with other trees or shrubs. Roots not tuberous. Leaves pinnately trifoliolate. Stipules basifixed, lanceolate and caducous. Stipels bristle-like. Pseudoracemes or panicles with many flowers on short branches, ascending or pendulous. Calyx campanulate, lobes 5, the upper two fused, short and blunt or minutely bifid. Corolla large. Vexillum apex rounded, without callosities, sides not reflexed, green spot and white rays in the center. Vexillary stamen connate to the staminal column. Staminal column moves up to touch vexillum in late blooming. Fruits coriaceous, glabrous, valves twisting upon dehiscence. Seeds orbicular and compressed.</p> <p>Description: —Perennial scandent shrub, erect or climbing. Stems woody, up to 7 cm in diameter, 2–4(–7) m tall, sparsely pubescent with adpressed hairs, glabrescent with age. Stipules basifixed, linear-acuminate, 4–11 mm × 2 mm, quickly caducous, stipule scar narrowly elliptic. Leaves pinnately trifoliolate; petiole striate, 5–18 cm long; terminal leaflets broadly ovate, rhomboid-elliptic, to suborbicular, 8–28 cm × 8–26 cm, lateral leaflets smaller, oblique, 7–23 cm × 4–16 cm, apex acuminate, rarely obtuse, base cuneate, leaflets green and glabrous above, grey-green and sparsely adpressed pubescent below, veins conspicuous below, in ca. 7 pairs; petiolules 4–10 mm long with spreading hairs; stipels small, 1–3 mm long, bristle-like, falling with age. Inflorescences solitary, axillary or terminal pseudoracemes, branched or not, (4–)10–40(–55) cm long, nodose (with brachyblasts) to short branched, 4 or more flowers per node; bracts subtending the nodes, 2–5 mm long, caducous; pedicels 2–6 mm, pubescent; bracteoles 2 per flower, ovate to lanceolate, 0.5–3 mm long, caducous. Calyx 4–6 mm long, short appressed hairs on the outside, the tube 3–5 mm long, 5-lobed, the upper two lobes fused entirely or nearly so, other lobes obtuse, 0.5–1 mm long. Corolla white to pink; vexillum obovate, green spot and white rays in the center, (10–)16–19(–23) mm × 8–15(–18) mm, apex rounded, auricles not reflexed, without callosities at base; wing petals white or pink, darker than vexillum or keel petals, 15–17 mm × 2–3 mm; keel petals strongly curved, ventrally and basally fused, white to light pink, 14–18 mm × 3 mm. Ovary elongate, finely hirsute, 8–12 mm long; style 4–6 mm long, the last 2–3 mm strongly upcurved, glabrous, stigma terminal, globose. Stamens monadelphous, the vexillary stamen connate to staminal column in the middle, free below, 14–15 mm long, the free end 1–2 mm, upcurved, stamens and style move up to touch vexillum in late blooming; anthers dorsifixed, alternately on long and short filaments. Fruits leguminous pods, flattened, oblanceolate, tan to medium brown, glabrous, coriaceous, ca. 5–8 ovuled, not septate, 6–13 cm × 0.8–1.2 cm, acuminate at both ends, style persistent, dehiscent when mature, valves twisting. Seeds orbicular to oblong, ca. 5–7 mm × 3–6 mm, ca. 2.5 mm thick, compressed, brown or with black mottling; funicle broad, triangular in shape; aril elongate.</p> <p>Phenology: —Flowering July to October (to February at lower elevations). Fruiting October to February.</p> <p>Distribution and Ecology: — Bangladesh, Bhutan, China (Tibet, Yunnan), India (E Himalayas, Meghalaya), Myanmar, Nepal, Thailand. Elevation 180–2000 (–2300) m; in hills and forest margins of dry evergreen forests where it is often associated with Dipterocarpaceae, particularly Shorea robusta, or with Quercus; in open grassy vegetation, on slopes, along rivers.</p> <p>Conservation: — Haymondia is fairly common in Thailand and Burma throughout dry dipterocarp forests and occurs within or near the borders of several national parks. It is less common in China and areas at the edge of its range. It is assessed here as Least Concern (LC) according to the criteria of IUCN (2001) based on frequency within its range and presence within protected areas.</p> <p>Etymology: —This genus is named after Welby Dean Haymond and Mildred Winona Davies Haymond, maternal grandparents of author Ashley N. Egan, who instilled and cultivated a love of nature and science in her by the simple act of allowance.</p> <p>Vernacular:—ẾËƀ xu mi ge (Chinese), บะแปบวอ ma paep wo (Thai)</p> <p>Discussion: — Haymondia wallichii, which commemorates Dr. Wallich who sent it, along with a number of other plant specimens, to de Candolle from Nepal (its type locality), was originally described by de Candolle in 1825 along with Pueraria tuberosa (the type species for Pueraria). The two species resemble each other superficially, both having large trifoliolate leaves, twining habit, and long, pendulous inflorescences. However, careful examination confirms that the two species are strikingly different. Lackey (1977b) tentatively separated members of genus Pueraria into four groups based on morphological differences such as number of flowers per node, stipule type, calyx type, callosities on the vexillum, and fruit type, with P. wallichii, P. peduncularis, and P. stracheyi comprising his group D. Lackey’s (1977b) group D admittedly comprised those species which he felt were “surely anomalous in the genus but fits nowhere else.” Lackey also found that P. wallichii contains the free amino acid canavanine, a chemical that most species of subtribe Glycininae lack. In fact, Lackey (1977b) went so far as to suggest that P. wallichii was not only anomalous in the genus, but perhaps in the tribe as well: “all Pueraria species studied have paraveinal mesophyll, except for P. peduncularis and P. wallichii. This substantiates the morphological data which indicates that these two species are probably generically, and perhaps subtribally or tribally misplaced”. Van der Maesen (1985) revised the genus Pueraria into three sections: Pueraria, Schizophyllon, and Breviramulae, placing wallichii into the latter, a motley group including the two new genera presented here. In spite of their observations, neither Lackey nor van der Maesen took action to describe new genera for or move the anomalous species, largely due to uncertainty as to true relationships. Lee and Hymowitz (2001) were the first to substantiate the hypotheses of these authors using phylogenetic evidence. Their rps16 phylogeny included P. pulcherrima, P. phaseoloides, P. lobata, P. stricta, and P. wallichii. Their findings support the exclusion of P. wallichii from Pueraria s.s., but they, also, did not take revisionary action, suggesting that “more taxa of Pueraria that represent all four groups should be included in a rigorous molecular investigation.” Cagle (2013) and Egan et al. (in prep.) included a comprehensive sampling of Pueraria as well as many other genera to ascertain how these anomalous species were related to other phaseoloid genera and found that Haymondia is a distinct phylogenetic lineage at the base of tribe Phaseoleae, but found no clear affinity of Haymondia for any other genera. A key feature of Haymondia is the position of stamens throughout flowering. At the onset of anthesis, the staminal column is positioned within the keel petals (Figure 5A), but later moves upward until the stamens and stigma are touching the vexillum or nearly so and are fully reflexed from the wing and keel petals (Figure 5C). This feature is shared with several other genera to which Haymondia is loosely affiliated in phylogenies, including Apios, Mucuna, Cochlianthus, and several members of tribe Desmodieae (Egan et al., in prep.).</p> </div>	https://treatment.plazi.org/id/039787AE3923846CFF6A5B6409D6FC12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE3925846FFF6A59EC0C3DFDD6.text	039787AE3925846FFF6A59EC0C3DFDD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxicopueraria	<div><p>Toxicopueraria A.N.Egan &amp; B. Pan, gen. nov.</p> <p>Type species: — Toxicopueraria peduncularis (Graham ex Bentham) A.N.Egan &amp; B. Pan (Neustanthus peduncularis Graham ex Bentham, in Miquel, Pl. Jungh. 2: 235. 1852).</p> <p>Diagnosis: —Woody lianas, roots not tuberous. Leaves pinnately trifoliolate. Leaflets entire. Stipules basifixed, lanceolate, open. Pseudoracemes long and pendulous, 3-several flowers clustered at each node. Bracts and bracteoles setaceous, quickly caducous. Pedicels slender. Calyx 4- or 5-lobed. Vexillum without callosities. Vexillary stamen adherent to the tube, but detaching with age. Fruits linear, flat, subchartaceous, valves not twisting upon dehiscence. Seeds elliptic, compressed.</p> <p>Description: —Perennial twining, woody climber. Roots not tuberous. Stems to 40 cm in diameter when mature. Branches robust, up to 10 m long. Stipules basifixed, ovate to lanceolate, 4–10 mm × 1–3 mm, striate, persistent to caducous, leaving a raised scar. Leaves pinnately trifoliolate; petiole striate, pubescent or glabrous, 4–13 cm long; leaflets ovate to rhomboid, lateral leaflets obliquely so, glabrous to strigulose on both sides, 5–14(–23) cm × 2–8(– 14) cm, base rounded-cuneate, apex long-acuminate, margins entire, veins prominent below, pubescent, in 6 or 7 unequal pairs with basal pair opposite; petiolules pubescent, 3–7 mm long; stipels short, 1–3 mm long, persistent. Inflorescences axillary, solitary pseudoracemes, 1 or 2 per axil, pendulous, 10–40(–60) cm long, slightly nodose (with swollen nodes or brachyblasts), with (2–)4–7 flowers per node; bracts subtending the nodes, 1–3 mm long, quickly caducous; pedicels slender, thickening in fruit, to 14 mm long; bracteoles 2 per flower, hirsute, minute, caducous. Calyx 4- or 5-lobed, with short, adpressed hairs on outside, glabrous on inside, tube 3–5 mm long, gibbous above base, lobes shorter than the tube, acute to broadly so, upper two lobes connate or only almost entirely so, 1–3 mm, lateral lobes triangular, 1–2 mm long, lower lobe narrowly triangular, 1–1.5 mm long. Corolla purplish-blue to violet or white suffused with purple or pink at the tips; vexillum orbicular-ovate to obovate, apex emarginate, base clawed, auricles inflexed or truncate, without callosities; wing petals oblong; keel petals ventrally fused. Ovary elongate, pubescent, 5–8 mm long, ca. 7 ovules; style glabrous, 2–5 mm, with terminal 2–3 mm inclined towards the vexillum; stigma terminal, globose, pubescent at the base. Stamens diadelphous, the vexillary stamen adherent to staminal column at first, becoming free with age, 10–14 mm long, the free part ca. 2–3 mm, inclined upward; anthers basi-dorsifixed on alternately long and short filaments. Fruits leguminous pods, flattened-oblong, black, purple-brown, or tan, glabrous, chartaceous, (3–) 5–7 cm × 0.5–1 cm, not septate, (1–)4–7 seeded, cuneate at base, acuminate at apex, style persistent, sutures robust. Seeds compressed; funicle elongate-triangular in shape; arils elongate.</p> <p>Etymology: — Toxicopueraria is derived from the latin toxicus (“poisoned”) due to the use of ground stems and roots as an insecticide and fish poison in Yunnan, China (Perry &amp; Metzger, 1980) and in deference to its former name, Pueraria.</p> <p>Discussion: —As with Haymondia wallichii, several botanists have recognized the anomalous placement of T. peduncularis within Pueraria (see discussion under Haymondia). Lackey (1977b) placed P. peduncularis in his group D, citing the absence of paraveinal mesophyll as evidence that the species did not belong to Pueraria, in addition to the minute bracteoles, a puckered calyx base and flat papery fruit. In spite of this, he did not create a new genus or offer any suggestions as to its affinity. Van der Maesen (1985) acknowledged these differences, but distinctly stated that he favored keeping P. peduncularis within Pueraria, stating “Even if biosystematic research would establish more distance from the other species, or even an anomalous position in the genus, it is not at all uncommon to admit within a genus a more or less anomalous species.” Although keeping P. peduncularis within Pueraria would mean fewer nomenclatural issues, it does not represent a natural or evolutionary grouping, and so we remove it and provide a new generic name here.</p> </div>	https://treatment.plazi.org/id/039787AE3925846FFF6A59EC0C3DFDD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE3926846FFF6A59810C8FF829.text	039787AE3926846FFF6A59810C8FF829.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxicopueraria peduncularis (Bentham)	<div><p>Toxicopueraria peduncularis (Graham ex Bentham) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — NEPAL. Graham, Wallich Cat.No. 5354 (holotype:K[barcode K001120656!]; isotypes: K[barcode K000264081!], BM[barcodes BM000958608!, BM000521674!, BM000958607!], CAL, G[barcodes G00370586!, G00370595!]).</p> <p>Basionym: — Neustanthus peduncularis Graham ex Bentham in Miquel, Pl. Jungh. 2: 235. 1852.</p> <p>Selected Synonyms:— Pueraria peduncularis (Graham ex Bentham) Bentham, J. Linn. Soc. Bot. London 9: 124. 1867; Pueraria peduncularis (Graham ex Bentham) Bentham var. violacea Franchet, Pl. Delav. 182. 1890. (lectotype: P[barcode P00500995]; isolectotype: A[barcodes A00228287!, A00228289!], K[barcode K000264082!]).</p> <p>Images: —Illustration (T. peduncularis): Figure 7; Photo Plate (T. peduncularis &amp; T. yunnanensis): Figure 8.</p> <p>Description: —Woody climber to 10 m. Leaves and stems densely pubescent, becoming glabrate with age. Petioles 4–13 cm; stipules without small spur-like projections beneath; leaflets hirsute on both surfaces, rarely glabrous adaxially, apex acuminate, base acute. Pseudoracemes slightly nodose, 20–50 cm long. Flowers white, tinged with purple to pink or flowers purple, blue, or mauve; 4–7 flowers per node; pedicels slender, 6–9 mm long. Calyx papyraceous, adpressed pubescent; upper 2 lobes connate or nearly so. Vexillum obovate, 11–15 mm × 8–10 mm, auricles truncate, inflexed; wings oblong, 8–14(–15) mm × 3–5 mm, claw ca. 3 mm long, apex obtuse; keel petals obovate, 8–12(–14) mm × 3–5 mm, claw 3–4 mm long. Fruits linear, tan to black when mature, 3–8 cm × 6 mm. Seeds elliptic, 2–3 × 3–4 mm, compressed, dark mahogany to black, sometimes with a red streak.</p> <p>Phenology: —Flowering June to October; fruiting August to December.</p> <p>Distribution and Ecology: — Bangladesh, Bhutan, India, Myanmar, Nepal, Pakistan, and S &amp; SW China. Forests or forest margins or in thickets. Elevation 1000–4300 m.</p> <p>Conservation: — Toxicopueraria peduncularis is assessed here as Least Concern (LC) according to the criteria of IUCN (2001) due to its fairly wide distribution, although a detailed study would be beneficial.</p> <p>Vernacular:—ẫƀ ku ge (Chinese), ȓħƀŭ Yun nan ge teng (Chinese), ting khla (Khasi).</p> <p>Discussion: — Toxicopueraria peduncularis has been documented for use by various indigenous peoples. The Nyishi (Daffla) tribe of Arunachal Pradesh in northeastern India are said to eat the fruits, either fresh or boiled (Srivastava et al. 2010), whereas the roots are crushed up and thrown into lakes and streams a fish poison in Yunnan, China (Perry &amp; Metzger, 1980).</p> </div>	https://treatment.plazi.org/id/039787AE3926846FFF6A59810C8FF829	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
039787AE39398473FF6A5A900E8AFDD6.text	039787AE39398473FF6A5A900E8AFDD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxicopueraria yunnanensis (Franch.)	<div><p>Toxicopueraria yunnanensis (Franch.) A.N.Egan &amp; B.Pan, comb. nov.</p> <p>TYPE: — CHINA. Yunnan. woods near Tapintze. April 1883 Delavay 506 (lectotype: P[barcode P00507990!]; isolectotypes: P[barcodes P02961048!, P00507992]) Lectotype designated by Z. F. Le and X. Y. Zhu (Ann. Bot. Fenn. 46: 419–424. 2009).</p> <p>Basionym: — Pueraria yunnanensis Franchet, Pl. Delav. 181. 1890.</p> <p>Selected Synonyms: — Derris bonatiana Pampanini, Nuovo Giorn. Bot. Ital., n.s. 17(1): 8. 1910 (lectotype: NY[barcode NY00007612!]; paratypes: Maire 210 P[barcode P02961022!])</p> <p>Images: —Photo Plate (T. peduncularis &amp; T. yunnanensis): Figure 8.</p> <p>Description: —Woody climber. Leaves and stems glabrous or thinly pubescent or glabrescent. Petioles 3–8 cm; stipules with two small spur-like projections below the stipules, 1–2 mm, these becoming woody with age; leaflets with sparse hairs, apex caudate-acuminate, base rounded. Pseudoracemes 10–25 cm long. Flowers white or cream, not suffused with purple or pink, (2–)3–6(–7) flowers clustered at nodes of rachis; pedicels slender, 2–8 mm. Calyx membranaceous, subglabrate, sparsely villous; upper 2 lobes connate entirely, apex of lobes obtuse. Vexillum orbicular-ovate; 11–13 mm × ca. 8 mm; auricles inflexed; wings oblong, (7–) 9–13 mm × 3–4 mm, claw ca. 4 mm long; keel petals (6–) 7–12 mm × 3–4 mm, claw ca. 4 mm long, apex obtuse. Fruits linear, tan when mature, 4–8 cm × 7–11 mm. Seeds kidney-shaped to elliptic, ca. 5 × 3 mm, compressed, red-brown to black.</p> <p>Phenology: —Flowering April to June; fruiting May to July.</p> <p>Distribution and Ecology: —Endemic to Southwest China (Chongqing, Guangxi, Guizhou, Sichuan, Yunnan). Forests and forest margins or in thickets. Elevation 800–2300 m.</p> <p>Conservation: — Toxicopueraria yunnanensis is endemic to five provinces in southwestern China and is less common than its congener. It is assessed here as Least Concern (LC) according to the criteria of IUCN (2001) due to its fairly wide distribution, but a more detailed study would be beneficial to accurately determine the conservation status.</p> <p>Discussion: — Toxicopueraria yunnanensis has been synonymized with T. peduncularis by a number of taxonomists (Lackey, 1977b; van der Maesen, 1985, 1994, 2002; D. Wu &amp; Thulin, 2010; T. L. Wu, 1995) because of their morphological similarities. However, they differ in indumentum, flower color, petal shape, projections below the stipules, and phenology (Figure 8). In addition, Le and Zhu (2009) found leaf epidermis and seed coat micro-characters to support the separation of these two species. From experience, the authors can easily determine T. yunnanensis from T. peduncularis in both live and pressed specimens, particularly in flower. Toxicopueraria yunnanensis tends to have a more upright vexillum with sides reflexed backwards whereas in T. peduncularis the whole vexillum is strongly reflexed, often to touch the calyx. Toxicopueraria yunnanensis has wing petals that are equal to or slightly longer than the keel, with the wings often displayed in a plane more perpendicular to the sides of the keel and with the ends curving backwards, presenting a more open display. In comparison, T. peduncularis has wing petals that are equal to or slightly shorter than the keel, and that are usually straight or slightly curved outward but more often in a plane parallel to the sides of the keel (compare Figure 8A and 8C). Furthermore, T. yunnanensis has small, spur-like projections that flank the stipule or stipular scar, these often becoming woody with age (see Figure 8C). These are lacking in T. peduncularis.</p> <p>Several issues concerning types of T. yunnanensis or its synonyms exist. Van der Maesen (1985) designated one collection at P as the holotype of Pueraria yunnanensis (barcode P00507990), listing another as an isotype. Le &amp; Zhu (2009) found issue with van der Maesen’s designation of a holotype because there is no specification by the author/ collector, J.M. Delavay, as to which of his collections, all designated under number 506 but collected during April or August 1883 or 1885, was to act as the holotype. Therefore, Le &amp; Zhu (2009) correctly determined the specimen selected by van der Maesen as the lectotype, relegating all other sheets of Delavay 506 (albeit with different dates) as isolectotypes.</p> <p>In addition, there is some confusion surrounding the types of Derris bonatiana. The protologue states the type as “Yunnan-sen, source du Pe-long-tan. 8 May, 1904 (Ducloux, n. 377); [sine loco] (Maire, n. 210).” A search of Ducloux collections at Paris showed that Ducloux numbers 362–399 were collected from “Environs de Yun Nan Sen” during August and September of 1897, suggesting that Ducloux 377 would have been collected during this time frame as well. While doing research for this manuscript, A.N. Egan came across a specimen from Paris [P02961736] collected by Ducloux (no. 2301) on 8 May 1904 from “Yun-Nan: environs de Yun nan-sen”, written in Ducloux’s own hand. A survey of Paris’ Ducloux numbers from 2273 to 2504 includes specimens collected from January to May of 1904, a range that includes that stated in the protologue. Because the institution of deposition of the specimen cited in the protologue was not named (Pampanini, 1910), van der Maesen (1985) designated a lectotype: “ China, Yunnan-sen, source of the Pe-long-tan river, 8 May 1904, Ducloux 377 (FI, holo, not seen).” This is the same information stated in the protologue, but with the addition of a location (FI, a place of employment of R. Pampanini, author of Derris bonatiana). Note that van der Maesen did not actually see the specimen he designated as the lectotype. Attempts by author A.N. Egan to contact curators at FI concerning a Ducloux 377 collection in their herbarium were unsuccessful. As of 18 May 2015, JStor Global Plants includes a type specimen of Derris bonatiana from NY [NY00007612], but the kind of type is not specified. The typed label of the NY Ducloux 377 collection states: “ Derris botaniana Pampanini n.sp., Yunnan Sen, source de Pé, 377 (E.E. Maire?)” The typed font of “E.E. Maire?” is crossed out in pencil with “ Ducloux! ” handwritten above it. E.E. Maire was a French contemporary of F. Ducloux who spent time in Yunnan as a missionary and began collecting plants in 1905. Thus, there is confusion concerning the type information stated in the protologue and how its matches with various collections in herbaria around the world. The above body of research suggests that Ducloux and Maire collections may have been mixed up or either the date of collection or the collection number or the collector where misstated in the protologue. Until it can be determined whether a Ducloux 377 specimen exists in FI, the type designations of D. bonatiana are dubious.</p> </div>	https://treatment.plazi.org/id/039787AE39398473FF6A5A900E8AFDD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Egan, Ashley N.;Pan, Bo	Egan, Ashley N., Pan, Bo (2015): Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria. Phytotaxa 218 (3): 201-226, DOI: 10.11646/phytotaxa.218.3.1, URL: http://dx.doi.org/10.11646/phytotaxa.218.3.1
