taxonID	type	description	language	source
03969B187A450319BC8620A8FD9FFD43.taxon	diagnosis	Diagnosis. Head with vertexal foveae devoid of setae, ventrally with prominent, parallel ridges, each bearing sharp spines at middle; maxillary palpi with three apical segments pedunculate at base, fourth segment triangular, apical pseudosegment present. Pronotum round, with median and lateral foveae of different sizes. Each elytron with one large basal foveae. Abdomen with first visible tergite (tergite IV) slightly longer than second (tergite V).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A450319BC8620A8FD9FFD43.taxon	description	Description. Body length 1.70 – 2.25 mm. Head with broad frontal rostrum (exception is A. spinipalpis which has narrower rostrum) antennal tubercles prominent, prolonged posteriad; rostral apex betweeen antennal tubercles with deep impression or sulcus (in A. spinipalpis); frontal fovea absent, two vertexal foveae well-defined but devoid of setae; eyes large, prominent, with 9 – 15 facets; temples short, rounded; head ventrally with prominent, parallel ridges, each bearing sharp spines at middle, large gular fovea present; maxillary palpi with segments II – IV pedunculate at base, segment II longest, with or without small inner tooth in apical third, segment III with or without large, lateral spine, segment IV of different shape, with large sensory area at apex bearing apical psedosegment; antennae with eleven antennomeres, covered with dense setae, scape long to very long, sometimes bent, about 3.0 – 7.5 times longer than pedicel, club with three to four segments; neck broad. Pronotum round, anterolateral corners completely absent, with median fovea devoid of setae, lateral foveae sparsely setose or also naked, foveae of different sizes, lacking antebasal sulcus; widest in apical third and convergent to apex. Thorax with median and lateral mesoventral foveae, with median and lateral metaventral foveae, metaventral apex broadly concave. Elytra short, each elytron triangular, with one basal fovea; with deep discal stria extending from basal fovea and reaching apex of elytron. Legs long and slender, protrochanter and profemora modified. Abdomen longer than elytra, with first visible tergite (tergite IV) longer than second (tergite V), convex, lacking basal depression, carinae, sulci or foveae; paratergites VII – X well-developed. Sexual dimorphism not apparent.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A450319BC8620A8FD9FFD43.taxon	discussion	Remarks. Anagonus, Paranagonus gen. nov. and Caledonogonus gen. nov. are closely related genera and well-isolated within the Tyrina. They are characterized by the combination of the following characters: 1) head triangular; 2) rostrum relatively broad and with large impression between antennal tubercles which are distinct and prominent; 3) scape long, usually bent, three or more times longer than pedicel; 4) each elytron short and triangular, strongly expanded from base to apex; 5) shape of maxillary palpi with triangular fourth segment. Anagonus can be separated from Caledonogonus by having one basal elytral fovea, presence of lateral pronotal foveae and apical pseudosegment of maxillary palpus. Anagonus can be readily distinguished from the more closely related Paranagonus by the shape of pronotum (lacking anterior angles and rounded anterior margin) and by rounded temples.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A450319BC8620A8FD9FFD43.taxon	distribution	Distribution. New Caledonia.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A460318BC86205EFC4AF926.taxon	materials_examined	Type material (ɗ). Holotype: NEW CALEDONIA: ɗ, [pink label] Coll. R. I. Sc. N. B. Nouvelle Calédonie Yahoué 2, rec Deplanche Fauvel / [red label] TYPE / Coll. et det. A. Fauvel Anagonus fracticornis Fvl. / [red ink] HOLOTYPE Anagonus fracticornis Fauvel, det. P. Hlaváč, 2007 (IRSNB). Additional material examined (2 ɗɗ, 2 ΨΨ). NEW CALEDONIA: Ψ, Mt. Koghi, primary forest, 400 – 500 m, 18 – 19. x. 1998, I. Löbl; ɗ, Ψ, Forêt Nord site 2, 22 o 19 ’ S 166 o 55 ’ E, 210 m, 21. IV. 2005, Monteith, berlesate; ɗ: Mt. Mou, base, 22 o 05 ’ S 166 o 20 ’ E, 350 m, 18. IV. 2005, GB Monteith, berlesate (ɗ, Ψ: QMBA, ɗ, Ψ: CPH).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A460318BC86205EFC4AF926.taxon	description	Description. Body length 2.15 – 2.25 mm, glossy, with sparse and erect golden setae; head and pronotum light brown, elytra, abdomen, legs and antennae yellowish brown, maxillary palpi yellow. Head 1.1 times as wide as long, with large frontal rostrum, antennal tubercles well-defined, rostral apex with deep triangular depression between antennal tubercles; eyes with 13 – 15 facets; maxillary palpi (Fig. 9) long, their segment I minuscule, II with small inner tooth, III pedunculate, lacking lateral spine, partly with wrinkled surface; antennae very long, scape more than seven times as long as pedicel and nearly three times as long as terminal antennomere, slightly shorter than funicule, antennomeres III – VII elongate, club foursegmented, VIII rhomboic, slightly longer than wide, IX 1.2 times as wide as long, X 1.7 times as wide as long, terminal antennomere pointed at apex, 1.65 times as long as wide and three times as long as X. Pronotum as long as wide, 1.25 times as wide as head, widest in apical third, lateral foveae large, slightly setose. Legs long, protrochanter with two spines, basal spine longer and more robust than apical one, almost as long as single basal spine on profemora. Abdomen 1.6 times longer than elytra. Aedeagus as in Figs. 13, 14.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A460318BC86205EFC4AF926.taxon	discussion	Remarks. Anagonus fracticornis is closely allied to A. spinipalpis sharing with this species a very long scape, but is readily separated by the shape of third segment of maxillary palpus which lacks a lateral protuberance.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A460318BC86205EFC4AF926.taxon	biology_ecology	Biology. All recent material was collected by sifting in primary forest.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A47031EBC8622D1FC7FFD7A.taxon	materials_examined	Type material (ɗ). Holotype: NEW CALEDONIA: ɗ, New Caledonia, above Serraméa creek, leaf litter, 24. x. 1998, I. Löbl / [red label], HOLOTYPE Anagonus breviscapus sp. nov., P. Hlaváč det., 2007 (MHNG).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A47031EBC8622D1FC7FFD7A.taxon	description	Description. Body length about 1.85 mm, shining, with sparse long erect golden setae, head and pronotum light brown, elytra, abdomen, maxillary palpi, antennae and legs lighter, yellowish brown. Head 1.3 times as long as wide, with wide frontal rostrum; rostral apex with deep, elongate impression between antennal tubercles which are well-defined; eyes with about 10 facets; maxillary palpi (Fig. 10) long, segment I minuscule, II with weakly defined inner tooth, III pedunculate at base, lacking lateral protuberance and lacking wrinkled surface; antennae very long, reaching middle of abdomen, scape about 4 times as long as pedicel and about 1.4 times as long as terminal antennomere, about half length of funicule, antennomeres II – VIII elongate, club three-segmented, antennomeres IX and X about as long as wide, terminal antennomere pointed at apex, about twice as long as wide and about 2.8 times as long as X. Pronotum slightly longer than wide, and 1.3 times as long as head, widest before middle, lateral foveae small, lacking setae. Legs long, protrochanter with two spines; basal spine longer and more robustr than apical one, almost as long as single basal spine on profemora. Abdomen 1.5 times longer than elytra, first visible tergite (tergite IV) 1.8 times as long as second (V).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A47031EBC8622D1FC7FFD7A.taxon	discussion	Remarks. Anagonus breviscapus can be easily separated from both congeners by the short scape and by the first visible tergite (tergite IV) 1.8 times as long as second (tergite V). Adeagus was lost during the preparation. An additional specimen, a female collected from Col des Roussettes by Ivan Löbl (MHNG), is very similar to A. breviscapus, but has the first visible tergite (tergite IV) much shorter than in the holotype of A. breviscapus, not more that 1.5 times as long as the second visible tergite (tergite V). Further, this female specimen has a much larger body, about 2.2 mm long. For the time being I decided not to name this specimen.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A47031EBC8622D1FC7FFD7A.taxon	biology_ecology	Biology. The holotype was sifted from leaf litter in a humid ravine in tropical forest and extracted by Winkler-Mocsarski eclectors (Löbl, personal communication).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A47031EBC8622D1FC7FFD7A.taxon	etymology	Etymology. The name of the new species refers to its short scape.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A41031EBC86273EFE6DF8E9.taxon	materials_examined	Type material (2 &&). Holotype: NEW CALEDONIA: &, NEW CALEDONIA (S), 22 o 05 ’ S 166 o 37 ’ E, 260 – 280 m, Haute Rivière Bleue, Sentier de Kaoris, virgin humid forest, sifted, 26.01.2004, leg. M. Wanat / [red label] HOLOTYPE Anagonus spinipalpis sp. nov., P. Hlaváč det., 2007 (MNHW). Paratype: NEW CALEDONIA: &, NEW CALEDONIA, Rivière Bleue, Mois de Mai / QN Berlesate 1089, 22 o 06 ’ S 166 o 38 ’ E, rainforest, 800 m, sieved litter / [red label] PARATYPE Anagonus spinipalpis sp. nov., P. Hlaváč det., 2007 (QMBA).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A41031EBC86273EFE6DF8E9.taxon	description	Description. Body length about 1.8 mm, shining, with sparse, erect golden setae, uniformly coloured dark brown to almost black, maxillary palpi much lighter. Head about as long as wide, with narrow frontal rostrum; rostral apex with deep elongate impression between antennal tubercles which are well-defined; eyes somewhat small, with about 9 facets; maxillary palpi long (Fig. 11), segment I minuscule, II with small inner tooth, III pedunculate at base, with large lateral spine, lacking wrinkled surface; antennae very long, reaching middle of abdomen, scape about seven times as long as pedicel and 2.15 times as long as terminal antennomere, slightly shorter than funicule, antennomeres II – VII elongate, club four-segmented, VIII rhombic, slightly longer than wide, IX and X about as long as wide, terminal antennomere pointed at apex, about 2.7 times as long as wide and about four times as long as X. Pronotum about as long as wide, 1.15 times as long as head, widest before middle, each lateral fovea prolonged anteriorly by deep sulcus. Legs long, protrochanter and profemora with long, subequal and outwardly curved basal spines. Abdomen 1.4 times as long as elytra, first visible tergite (tergite IV) slightly longer than second (tergite V).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A41031EBC86273EFE6DF8E9.taxon	discussion	Remarks. Anagonus spinipalpis is readily distinguished from its congeners by its entirely dark brown colour, and the presence of a lateral protuberance on the third maxillary palpus as well as with lateral fovea prolonged anteriorly by deep sulcus	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A41031EBC86273EFE6DF8E9.taxon	biology_ecology	Biology. Both specimens were sifted in virgin humid forest and extracted by Berlese.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A41031EBC86273EFE6DF8E9.taxon	etymology	Etymology. The name of the new species refers to the presence of the lateral protuberance on the third segment of maxillary palpi.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	materials_examined	Type species. Caledonogonus loebli sp. nov.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	diagnosis	Diagnosis. Head with vertexal foveae present, ventrally lacking ridges and spines, maxillary palpi with segments II and III pedunculate at base, fourth segment triangular, apical pseudosegment absent. Pronotum with distinct median fovea, lacking lateral foveae, median metaventral fovea present. Each elytron with two basal foveae. Abdomen with first visible tergite (tergite IV) longest.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	description	Description. Length 1.4 – 1.6 mm. Head with broad rostrum bearing large triangular depression between distinct and prominent antennal tubercles, frontal foveae absent, vertexal foveae punctiform; eyes large, prominent, with 5 – 8 facets, temples short, rounded; ventrally head with median gular fovea transverse, two anterior foveae located before eyes well-defined, lacking ridges and spines; maxillary palpi (Fig. 12) with segments II – III pedunculate, segment II pedunculate in basal half, longest, about twice as long as III, segment III almost oval, gradually widening to apex from very shortly pedunculate base, segment IV triangular, with large sensory area at apex and lacking apical pseudosegment; antennae (Fig. 8) covered with dense setae, scape bent and very long, about as long as funicule, much longer than pedicel, club three-segmented; neck broad. Pronotum round, anterolateral angles completely absent, with median fovea devoid of setae, lacking lateral foveae and antebasal sulcus. Thorax with median mesoventral foveae, without lateral procoxal foveae, but with median and lateral metaventral foveae. Elytra short, each with two basal foveae; each fovea with stria extending from foveae through nearly three-quarters of elytron length. Abdomen longer than elytra, with first visible tergite (tergite IV) longest, but only slightly longer than second visible tergite (tergite V) and distinctly shorter than second and third visible tergites combined; convex, lacking basal depression, carinae and sulci; tergite IV with lateral foveae adjacent to paratergite present, paratergites VII – X well developed. Sexual dimorphism not apparent.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	discussion	Remarks. Caledonogonus can be readily separated from Anagonus and Paranagonus by the presence of two basal foveae on each elytron, the lack of ridges and spines on the ventral side of the head, the absence of the lateral pronotal foveae and the apical palpal pseudosegment.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	distribution	Distribution: New Caledonia.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031DBC8624ABFBBFFAE9.taxon	etymology	Etymology. The name is a combination of „ Caledono- “ referring to New Caledonia and „ - gonus “ showing the relationship of the new genus to Anagonus. Gender masculine.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031CBC8622EEFE2BF91B.taxon	materials_examined	Type material (38 specimens). Holotype: NEW CALEDONIA: ɗ, New Caledonia, Col des Roussettes, ravine, forest litter, 25. ix. 1998, I. Löbl / [red label] HOLOTYPE Caledonogonus loebli sp. nov. P. Hl a vá č det. 2007, (MHNG). Paratypes: NEW CALEDONIA: 10 ɗɗ, 7 ΨΨ, the same locality as holotype; 7 ɗɗ, 10 ΨΨ, 3 specimens: FM (HD) 78 - 256, NEW CALEDONIA: Col de Rousettes nr. Bourail, 500 m, VIII- 7 - 1978, Leg. S & J Peck, Berlese rainforest litter. All paratypes bear the following red label: PARATYPE Caledonogonus loebli sp. nov., P. Hlaváč det., 2007 (6 ɗɗ, 4 ΨΨ, 1 specimen: MHNG, 5 ɗɗ, 8 ΨΨ, 2 specimens: FMNH, 2 ɗɗ, 2 ΨΨ: QMBA, 3 ɗɗ, 3 ΨΨ: CPH).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031CBC8622EEFE2BF91B.taxon	description	Description. Body length 1.4 – 1.6 mm, shining, dark reddish-brown, with long golden setae except on disc of pronotum; elytra and first visible abdominal tergite (tergite IV), legs, maxillary palpi and apical antennomeres lighter, yellowish brown. Head about as long as wide and slightly shorter than pronotum, pronotum widest in apical third; antenna (Fig. 8) short and stout, scape about five times as long as pedicel, slightly shorter than funicule, antennomeres III – VI quadrate, VII – VIII transverse, three-segmented antennal club very clearly defined, antennomeres IX and X about equal in size, X about 1.5 times as wide as long, terminal antennomere pointed apically, about 1.7 times as long as wide. Legs shorter, protrochanter with two minuscule spines, lateral one longer. Aedeagus as in Figs. 15, 16.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031CBC8622EEFE2BF91B.taxon	discussion	Remarks. Anagonus loebli is readily distinguished from its congener by smooth disc of pronotum and shorter terminal antennomere.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031CBC8622EEFE2BF91B.taxon	biology_ecology	Biology. All specimens collected by Ivan Löbl were sifted from leaf litter in a humid ravine in tropical forest and extracted by Winkler-Mocsarski eclectors (Löbl, personal communication).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A42031CBC8622EEFE2BF91B.taxon	etymology	Etymology. The species is named after my friend Ivan Löbl (Geneva) who collected the holotype and some of the paratypes.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A430313BC862334FC79FDEE.taxon	materials_examined	Type material (ɗ, Ψ). Holotype: NEW CALEDONIA: ɗ, NEW CALEDONIA 1 1960, 20 o 47 ’ S x 164 o 35 ’ E, 980 m, Mt. Taom summit, Site 1. 4 Jan 2005, G. Monteith, Berlesate, sieved litter, RF / [red label] HOLOTYPE Caledonogonus pilosus sp. nov. P. Hl av á č det. 2007 (MNHN). Paratype: NEW CALEDONIA: Ψ, the same data as holotype, with the following red label (p) PARATYPE Caledonogonus pilosus sp. nov. P. Hl a vá č det. 2007 (QMBA).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A430313BC862334FC79FDEE.taxon	description	Description. Body length 1.5 – 1.6 mm, shining, entirely dark brown, with long and dense setation on whole body; legs, maxillary palpi and antennae slightly lighter. Head slightly wider than long and slightly shorter than pronotum, pronotum globular; antenna short and stout, scape about five times as long as pedicel, about as long as funicule, antennomeres III – VI quadrate, VII – VIII transverse, three-segmented antennal club very clearly defined, antennomeres IX and X of about same size, transverse, terminal antennomere pointed apically, 1.9 times as long as wide. Legs shorter; protrochanter with one laterally bent spine; profemur close to base with well defined spine. Aedeagus as in Figs. 17, 18.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A430313BC862334FC79FDEE.taxon	biology_ecology	Biology. All specimens were sifted from leaf litter in rain forest and extracted by Berlese.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A430313BC862334FC79FDEE.taxon	etymology	Etymology. The name of the species relates to its pubescent body.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	materials_examined	Type species. Paranagonus excavatus sp. nov.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	diagnosis	Diagnosis. Head with vertexal foveae devoid of setae, ventrally lacking prominent parallel ridges but with sharp spines, maxillary palpi with segments II – IV pedunculate at base, segment IV triangular, apical pseudosegment present. Pronotum with sharp, very prominent anterior angles and concave anterior margin, with median fovea devoid of setae and large lateral setose fovea. Each elytron with one large basal fovea. Abdomen with first visible tergite (tergite IV) almost twice as long as second (tergite V).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	description	Description. Length 1.9 mm. Head triangular, with broad rostrum which bears large, triangular depression between distinct and prominent antennal tubercles, frontal foveae absent, vertexal foveae well defined, deep, devoid of setae; eyes large, prominent, with about seven facets, temples very short, angular; head ventrally with median gular fovea transverse, with two anterior pits located before eyes well defined, without prominent parallel ridges but with sharp spines; maxillary palpi with segment II pedunculate in basal half; segment III pedunculate at base and longest, almost oval, gradually widening to apex from very abruptly pedunculate base; fourth segment also slightly peduculate at base, with apical pseudosegment. Antennae with all antennomeres covered with dense setae, scape bent and very long, much longer than pedicel, club foursegmented. Neck broad. Pronotum with anterior angles sharp and very prominent, median fovea distinct and devoid of setae, lateral foveae large and setose; pronotum lacking antebasal sulcus, anterior margin concave. Thorax with median and lateral metaventral foveae. Elytra short, each elytron triangular with one basal fovea; with striae extending from basal fovea and reaching apex of elytron. Abdomen longer than elytra, with first visible tergite (tergite IV) longest, almost twice as long as second visible tergite (tergite V), lacking basal depression, sulci or foveae; paratergites VII – X well developed. Sexual dimorphism. Female unknown.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	discussion	Remarks. See discussion under Anagonus.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	distribution	Distribution: New Caledonia.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4C0313BC862689FBCCF8B3.taxon	etymology	Etymology. The name is formed as a combination of Greek ” para - ”, meaning beside or near, and ” Anagonus ”, showing the relation of the new genus to Anagonus. Gender masculine.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4D0312BC8620B9FA8FF840.taxon	materials_examined	Type material (ɗ): Holotype: NEW CALEDONIA: ɗ, (p) NEW CALEDONIA, Col d’Amieu, W slope, 27 January 2004, G. B. Monteith / QM Berlesate 1120, 21 o 37 ’ S x 165 o 49 ’ E, rainforest, 470 m, sieved litter / red label (p) HOLOTYPE Paranagonus excavatus sp. nov., P. Hlaváč det., 2007, (MNHN).	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4D0312BC8620B9FA8FF840.taxon	description	Description. Body length 1.85 mm, shining and with dense, erect golden setae; head and pronotum brown, elytra, abdomen, legs and antennae lighter, yellowish brown, maxillary palpi yellow. Head triangular, frontal rostrum about half as wide as base of head, rostral apex with deep but short triangular depression between well defined antennal tubercles; eyes with six facets; antennae long, scape seven times as long as pedicel and about 1.7 times as long as terminal antennomere, pedicel short, 1.5 times shorter than antennomere IV, antennomeres III – V elongate, IV – V of about the same length, 1.3 times as long as III, antennomere VII slightly longer than globular antennomere VI, antennomeres VIII – X transverse, IX – X about same size and larger than asymmetric antennomere VIII, terminal antennomere pointed at apex, about twice as long as wide. Pronotum about as long as wide and 1.3 times as long as head, widest just behind anterior corners, lateral foveae setose. Abdomen 1.4 times longer than elytra, first visible tergite (tergite IV) twice as long as second (tergite V). Legs shorter; mesofemur with large and deep excavation at apex; protrochanter with large truncate protuberance; profemur close to base with large spine. Aedeagus as in Figs. 19, 20.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4D0312BC8620B9FA8FF840.taxon	biology_ecology	Biology. Holotype was sifted from leaf litter in rainforest and extracted by Berlese.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
03969B187A4D0312BC8620B9FA8FF840.taxon	etymology	Etymology. The species name refers to the deep excavation in the apical part of each mesofemur.	en	Č, Peter Hlavá (2009): The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. II. Revision of the tribe Tyrini Reitter, 1882. Zootaxa 2010: 46-56, DOI: 10.5281/zenodo.185787
