taxonID	type	description	language	source
0389D55CFFB682460AA0664CFEAFBEF4.taxon	materials_examined	Type species: Lyclene kepica Dubatolov & Bucsek, 2013.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB682460AA0664CFEAFBEF4.taxon	diagnosis	Diagnosis. Small moths with a typical ‘ miltochristoid’ forewing pattern. Body and wings coloration is yellow, forewing markings are blackish. Externally members of the new genus resemble Miltochrista obsoleta (Figs 30, 31, 52, 64), but can be distinguished by the less interrupted antemedial and medial transverse lines and the forewing discal spot situated closer to the costal margin. The male genitalia of Huangilene gen. n. are characterized by the following autapomorphic features that clearly distinguish them from those of the genus Miltochrista (Figs 34, 54, 65): (1) the anellus with two trigonal sclerotized lobes fused to each other basally and bearing series of denticles on their distal margins; (2) the presence of a lobe-like medial costal process, which is broad with a dentate ventral margin in the H. alikangiae species-group and bilobate with the distal lobe serrulate margin in the H. apoklinousa sp. n. species-group; (3) the distal section of costa present as a short, narrow and somewhat twisted lobe fused with the distal membranous lobe of valva. In addition, the male genitalia of the new genus are characterized by the large distal membranous lobe of valva being well-separated from the distal saccular process; and the aedeagus vesica curved dorsally and having a large medial ventral diverticulum bearing a robust spine-like cornutus, and the absence of a distal plate of vesica (= basal plate of vesica ejaculatorius). Also, in the H. alikangiae species-group the distal section of aedeagus is rugose with a long sclerotized string-like dentate dorsal carinal process protruding to the base of the vesica ejaculatorius, and a broad ventral area of numerous denticles protruding to the subbasal diverticulum, but in the H. apoklinousa sp. n. species-group the distal section of aedeagus is evenly serrulate without protrusions to the vesica. The female genitalia of Huangilene gen. n. are similar to those of Miltochrista, but differ by the area of numerous small spinules at the junction of the ductus bursae with the corpus bursae being protruding to the anterior section of the ductus bursae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB682460AA0664CFEAFBEF4.taxon	discussion	Remark. Species of the genus vary in their size and the clarity of forewing pattern, but the genitalia structures of various populations display no significant intraspecific variability: in males the width of valva (especially that of its distal lobe) may vary. However, the vesica structure is very stable within all species and is a reliable specific character.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB682460AA0664CFEAFBEF4.taxon	description	Description. External morphology of adults (Figs 1 – 29). Forewing length 7 – 11 mm in males and 9.5 – 14 mm in females. Male antenna ciliate, female antenna filiform. Body and wing coloration yellow. Thorax with two black dots. Forewing pattern elements thin, blackish. Subbasal spot. Subbasal line wavy, interrupted into several spots. Antemedial line strongly wavy, interrupted into a series of spots. Medial line sinuate or nearly straight, interrupted into a series of spots or continuous. Postmedial line S-like curved and irregularly dentate. Subterminal line wavy, interrupted into several spots. Terminal line absent. Discal spot as a dot. Cilia pure yellow. Hindwing pale yellowish, sometimes with slight dark greyish suffusion at apex. Male genitalia (Figs 35 – 51). Uncus elongate, thin, smoothly curved, apically pointed, slightly broadened medially and basally. Tuba analis moderately broad, scaphium thin, moderately sclerotized. Tegumen short and moderately broad. Anellus with two trigonal sclerotized lobes fused to each other basally and bearing series of denticles on their distal margins. Juxta broadly trapezoid with rounded corners. Vinculum well-sclerotized, U-shaped. Valva elongate and narrow. Costa with lobe-like medial process, which is broad with a dentate ventral margin (the H. alikangiae species-group) or bilobate with the distal lobe serrulate margin (the H. apoklinousa sp. n. species-group). Distal section of costa present as short, narrow and somewhat twisted lobe fused with distal membranous lobe of valva. Distal membranous lobe of valva large, wellseparated from the distal saccular process. Sacculus narrow, its distal process elongate, spur-like, slightly curved dorsally. Aedeagus short and moderately broad, ventral part of carina longitudinally rugose and weakly dentate, this dentation continues to the basal section of vesica. Dorsal part of carina as weakly sclerotized cord continuing to the basal section of vesica. Vesica curved dorsally, with large medial ventral diverticulum bearing a robust spine-like cornutus and two or three small medial diverticula. Distal plate of vesica absent. Female genitalia (Figs 55 – 63). Papillae anales broadly trapezoid with rounded corners, weakly setose. Apophyses long and thin, apophyses anteriores equal in length to apophyses posteriores or slightly shorter. Ostium bursae broad. Antrum broad, granulated, with rugose thick-walled membranous ostial margin. Ductus bursae short, dorso-ventrally flattened, moderately sclerotized. Corpus bursae sack-like, weakly spinulose scobinated, with area of numerous small spinules at junction with ductus bursae protruding to anterior section of ductus bursae. Appendix bursae conical, membranous, situated postero-laterally, directed posteriorly.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB682460AA0664CFEAFBEF4.taxon	distribution	Distribution. The genus is widespread from Nepal and northeastern India through Myanmar and southwest and east China to Indochina except Malay Peninsula.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB682460AA0664CFEAFBEF4.taxon	etymology	Etymology. The generic name is a combination of the surname Huang and the genus group name Lyclene. The genus is dedicated to Mr Huang Si-yao (Guangzhou, China), the authors’ colleague and an expert in Chinese Lithosiini.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB582460AA06339FDD6BA54.taxon	diagnosis	Diagnosis. The male genitalia of the species-group are characterized by the broad, lobe-like medial costal process having a dentate ventral margin, the short, thin and not serrulate distal section of costa (Fig. 50), the distal membranous lobe of valva of moderate size, the strongly elongate distal saccular process, the distal section of aedeagus with two sclerotized protrusions to the vesica, and the moderately long medial ventral diverticulum of vesica bearing a cornutus with a narrow base. The female genitalia are characterized by the elliptical corpus bursae of moderate size and the short and conical appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06599FBFBBE12.taxon	description	(Figs 1 – 3, 35, 36, 50, 55)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06599FBFBBE12.taxon	materials_examined	Type material examined. Holotype (Figs 1, 35): male, “ Cambodia, Kep Province, Kep, Treetop Bungalow, 10.494 ° N, 104.296 ° E, 51 m, O. Kosterin [leg.], 5. XII. 2010 ” / red handwritten label “ Holotypus ♂ Lyclene kepica Dubatolov et Bucsek ”, slide AV 1713 Volynkin, ex prep. in sugar by V. V. Dubatolov (Coll. SZMN). Additional material examined. CAMBODIA: 1 male, 1 female, Kirirom Nat. Res., 650 m, 7 – 9. XI. 2007, leg. S. Murzin, slides ZSM Arct. 2019 - 587 (male), 2019 - 588 (female) Volynkin (Coll. MWM / ZSM); 1 male, 2 females, Kampot Prov., Bokor N. P., Hill Station, 1025 m, 10 ° 37 ’ N 104 ° 01 ’ E, 19 – 21. I. 2006, leg. G. Csorba & G. Ronkay, slides MWM 33522 (male), MWM 33524, ZSM Arct. 2020 - 105 (females) Volynkin (Coll. MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06599FBFBBE12.taxon	diagnosis	Diagnosis. Forewing length is 7 – 8 mm in males and 9.5 – 10 mm in females. The species differs externally from other congeners by its more diffuse forewing markings. In the male genitalia, the narrow valva with a narrow distal membranous lobe, and the narrow vesica with a relatively small cornutus are characteristic. The differences between H. kepica and the closely related H. odontotilepida sp. n. are listed in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06599FBFBBE12.taxon	distribution	Distribution. Southern Cambodia (Kep, Kampot and Kampong Speu Provinces).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06652FBC8BA06.taxon	description	(Figs 4 – 6, 37 – 39, 56)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06652FBC8BA06.taxon	materials_examined	Type material. Holotype (Figs 4, 37): male, “ SE Thailand, Chantaburi prov., 265 m, Bar Ta Moon, Soi Dao Dist., 13 ° 3 ’ 40 ’’ N 102 ° 14 ’ 45 ’’ E, 21. XI. 2005, leg. K. Černý ”, slide AV 2650 Volynkin (Coll. MWM / ZSM, ex coll. CKC). Paratypes (42 males and 30 females in total). THAILAND: 6 males, 1 female, C Thailand, Saraburi, Khao Yai NP, Mt. Khao Kaew, 1244 m, 14 ° 21 ’ 56 ’’ N 101 ° 24 ’ E, 31. V. 2005, leg. K. Černý, slide AV 2648 (male) Volynkin (Coll. CKC); 1 male, 5 females, C Thailand, Saraburi, Khao Yai NP, Mt. Khao Khieo, 752 m, 14 ° 24 ’ 20 ’’ N 101 ° 22 ’ 14 ’’ E, 1. VI. 2005, leg. K. Černý, slide AV 2649 (female) Volynkin (Coll. CKC); 1 male, 21 – 22. VIII. [20] 09, Thailand, Nakhon Tai, Province Phitsanulok, h- 214 m, at light, N 17 ° 06 ’ 13 ’’ E 100 ° 50 ’ 28 ’’, A. V. Korshunov leg., slide AV 1630 Volynkin (Coll. CAV); 1 male, Thailand, Korat Plateau, Korat, 14 ° 48 ’ N 102 ° 07 ’ E, VI. 1996, leg. Steinke & Lehmann, slide ZSM Arct. 2019 - 981 Volynkin (Coll. MWM / ZSM); LAOS: 32 males, 24 females, 10 – 20. VIII. 2014, South Laos, Champasak prov., 27 km ENE of Pakse, near Tad Fane waterfall, N 15 ° 11 ’ 01.1 ’’ E 106 ° 07 ’ 40.5 ’’, 950 m, leg. local collector (Coll. CAV); CAMBODIA: 1 male, Mondolkiri Prov., Seima Biodiv. Cons. Area, 360 m, 12 ° 15 ’ N 107 ° 03 ’ E, 27 – 29. I. 2006, leg. G. Csorba & G. Ronkay, slide MWM 33523 (male) Volynkin (Coll. MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06652FBC8BA06.taxon	diagnosis	Diagnosis. Forewing length is 8 – 8.5 mm in males and 9.5 mm in females. Huangilene odontotilepida sp. n. is very similar to H. kepica, and differs externally by its slightly sharper forewing markings only. The male genitalia of the new species can be distinguished from those of H. kepica by the larger spinules on the lobes of anellus, more robust denticles on the medial costal lobe-like process, somewhat broader distal membranous lobe of valva, more massive distal saccular process, and larger cornutus in aedeagus vesica. The female genitalia of the two species are similar, but in H. odontotilepida sp. n. the apophyses anteriores and posteriores are shorter than those of H. kepica, and the spinules at the junction of the corpus bursae and ductus bursae are smaller.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06652FBC8BA06.taxon	distribution	Distribution. The species is found from central and southeastern Thailand (Saraburi, Phitsanulok and Korat Provinces) (Černý & Pinratana 2009, as Lyclene unguifera, part), southern Laos (Champasak Province) and eastern Cambodia (Mondolkiri Province).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482470AA06652FBC8BA06.taxon	etymology	Etymology. ‘ Odontoti lepida’ is the Latin transliteration of the Greek ‘ οδοντωτή λεπίδα’ meaning ‘ dentate lobe’. The specific epithet refers to the strongly dentate medial costal lobe-like process.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	description	(Figs 7 – 10, 40, 41, 57, 58)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	materials_examined	Type material. Holotype (Figs 7, 40): male, [China, Fujian] “ Kuatun, Prov. Fukien, 12. VIII. [19] 38 Höne ” / “ undulosa Wkr. ”, slide AV 1976 Volynkin (Coll. ZFMK). Paratypes (16 males and 13 females in total). CONTINENTAL CHINA: 1 female, [Fujian] Kuatun (2300 m), 27,40 n. Br. [N] 117,40 ö. L. [E], J. Klapperich (Fukien), 18. VII. [19] 34 / undulosa Wkr., slide AV 1975 Volynkin (Coll. ZFMK); 1 female without abdomen, China / Wuy Shan, Jiangxi – Fujian border, 50 km SE of Yingtan, 27 ° 56 ’ N, 117 ° 25 ’ E, 1600 m, V. 2002, leg. Siniaev & local coll. (Coll. MWM / ZSM); TAIWAN: 1 male, Prov. Taipei, Pi Hu, 450 m, 24 ° 54 ’ 02 ’’ N 121 ° 45 ’ 27 ’’ E, 4 – 5. IV. 1997, leg. L. Peregovits & A. Kun, slide ZSM Arct. 2019 - 980 (Coll. MWM / ZSM); 1 male, Prov. Taipei, Pi Hu, 410 m, 26. III. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 1 male, the same locality, but, 3. VII. 1997, leg. B. Herczig & S. Kovács (Coll. MWM / ZSM); 5 males, 1 female, Prov. Taipei, Pi Hu, 450 m, 24 ° 54 ’ 02 ’’ N, 121 ° 45 ’ 27 ’’ E, 4 – 5. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 2 males, Prov. Taipei, Hsiapen, 650 m, 11. VII. 1997, leg. B. Herczig & Y. Y. Lien (Coll. MWM / ZSM); 2 males, Prov. Taipei, 10 km SE of Pinglin, Pihou, 450 m, 26. III. 1997, leg. Csorba & Ronkay, slide ZSM Arct. 2019 - 979 Volynkin (Coll. MWM / ZSM); 1 male, Prov. Ilan, 700 m, Fu Shan Botanical Garden, 27 – 28. III. 1997, leg. Csorba & Ronkay, slide ZSM Arct. 2019 - 978 Volynkin (Coll. MWM / ZSM); 6 females, Prov. Ilan, 3 km SS of Suao, 121 ’ 51 ’’ E – 24 ’ 34 ’’ N, 400 m, 26. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 1 male, Prov. Ilan, Fu Shan Bot. Garden, LTER site, 700 m, 24 ° 45 ’ 13 ’’ N, 121 ° 35 ’ 39 ’’ E, 8 – 9. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 1 female, Prov. I-lan, 5 km SW of Ilan, 290 m, 25. II. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 1 female, Prov. Nantou, 5 km SW of Tayüling, 121 ’ 17 ’’ E – 24 ’ 09 ’’ N, 2900 m, 21. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 1 male, Prov. Tai-Tung, 5 km W of Chipen, 350 m, 09. III. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 1 male, Prov. Ping-Tung, 10 km SE of Mutan, 470 m, 03 – 04. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 1 female, Prov. Pingtung, 500 on the Road No. 199, 22 ° 14 ’ 38 ’’ N, 120 ° 51 ’ 51 ’’ E, 19. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 1 female, Prov. Kao-Hsiung, 3 km E Chiashen, 460 m, 18. X. 1996, leg. Gy. Fábián & F. Németh (Coll. MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	discussion	Remark. Daniel (1952) erroneously recorded this species from eastern continental China as Lyclene (Asura) undulosa (Walker, 1854). The true Miltochrista undulosa (Figs 32, 33, 53) is clearly different from members of Huangilene in its coloration and genitalia structures.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	diagnosis	Diagnosis. Forewing length is 9 – 10 mm in males and 11 – 12 mm in females. Huangilene kutzscheri sp. n. is very similar externally to H. pseudolutara, H. kepica and H. odontotilepida sp. n., but, however, males of the new species can be easily recognized by the yellow distal part of abdomen, which is covered with blackish hair-like scales in the three similar species. The male genital capsule of the new species differs from those of other congeners by the strongly elongate distal membranous lobe of valva. In addition, compared to those of the most similar H. pseudolutara, the male genitalia of H. kutzscheri sp. n. have the more elongate distal saccular process, narrower aedeagus, conspicuously broader distal chamber of aedeagus vesica, and conspicuously smaller distal diverticulum. The female genitalia of H. kutzscheri sp. n. differ from those of H. pseudolutara by the narrower ductus bursae, broader, more globular corpus bursae, smaller spines at the junction of the corpus bursae with ductus bursae, more heavily scobinated anterior section of corpus bursae, and membranous basal section of appendix bursae (in H. pseudolutara that is sclerotized).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	distribution	Distribution. The new species is found from eastern continental China (Zhejiang Province) (Daniel 1952, as Lyclene (Asura) undulosa, part) and the island of Taiwan.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB482400AA0627EFC4DBB48.taxon	etymology	Etymology. The species is named after Mr Christian Kutzscher (SDEI, Müncheberg, Germany), who kindly located the unlabeled syntypes of Asura obsoleta Form alikangiae Strand, 1917, what helped us to separate the Taiwanese populations of the new species from the sympatric H. alikangiae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB382410AA063ACFA29BAA3.taxon	description	(Figs 11 – 18, 42, 43, 59)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB382410AA063ACFA29BAA3.taxon	materials_examined	Material examined. NEPAL: 1 male, 4 females, Pokhara, 01. VII. 1995, leg. V. Sinyaev & E. Afonin (Coll. MWM / ZSM); 3 females, Dhumre, Bimal Nager, 500 m, 29 – 30. III. 1995, 84 ° 26 ’ E, 27 ° 55 ’ N, leg. László Bódi & György Makranczy (Coll. MWM / ZSM); 1 male, 3 females, 500 m, Dhumre, Bimal Nager, 26 – 28. III. 1995, 84 ’ 26 ’ E, 27 ’ 55 ’ N, leg. M. Hreblay & L. Nemeth, slides ZSM Arct. 2019 - 395 (male), 2019 - 396 (female) Volynkin (Coll. MWM / ZSM); 2 males, 1 female, Tanahoun distr., Bimalnager village, 530 m, 11 – 12. IV. 1995, leg. Gy. M. László & G. Ronkay, slides ZSM Arct. 2019 - 393, 2019 - 394 (males) 2019 - 412 (female) Volynkin (Coll. MWM / ZSM); INDIA: 1 female, SE Sikkim, 1500 – 1800 m, VIII. 94, exp. Cadio, coll. Schintlmeister (Coll. MWM / ZSM); 3 males, NE. India, Assam, Nambor Reserve Forest, Garampani, H = 100 m, 26 ° 30 ’ N, 93 ° 55 ’ E, 21 – 29. XI. 1997, lg. V. Sinyaev & M. Murzin, slides ZSM Arct. 2019 - 448, 2019 - 453 Volynkin (Coll. MWM / ZSM); 3 males, NE India, Assam, Kaziranga Wild Life, Pan Bari Reserve Forest, 26 ° 45 ’ N, 93 ° 10 ’ E, 100 m, 12 – 21. XI. 1997, lg. V. Sinyaev & M. Murzin, slides ZSM Arct. 2019 - 449, 2019 - 450 Volynkin (Coll. MWM / ZSM); 10 females, NE India, W Meghalaya, Umran, 33 km N Shillong, 26 ° 06 ’ N, 92 ° 23 ’ E, 800 m, 14 – 23. VII. 1997, leg. Sinyaev & Afonin (Coll. MWM / ZSM); MYANMAR: 1 male, Myanmar (Burma), 40 km N Myitkyina, Chanc Kand Village, 235 m, 23 – 24. IV. 1998, leg. Murzin & Sinyaev, slide ZSM Arct. 2019 - 397 Volynkin (Coll. MWM / ZSM); 1 male, Myanmar (Burma), 21 km E Putao, Nan Sa Bon village, 550 m, 1 – 5. V. 1998, leg. Murzin & Sinyaev, slide ZSM Arct. 2019 - 982 Volynkin (Coll. MWM / ZSM); CHINA: 1 male, 1 female, P. Yunnan, Simao District, Mangxi Ba Mts., 18 km S of Simao, 1280 m, 26. II. – 20. III. 1999, leg. Dr. R. Brechlin, slides ZSM Arct. 2019 - 410 (male), 2019 - 411 (female) Volynkin (Coll. MWM / ZSM); 1 female, the same locality and collector, but 15 – 21. VI. 1999 (Coll. MWM / ZSM); THAILAND: 1 male, Changwat Chiang Mai, 12 km NW Chiang Dao, 750 m, 12. XI. 1998, leg. Tibor Csővári & László Mikus (Coll. MWM / ZSM); 1 male, Changwat Chiang Mai, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 8. I. 1999, leg. A. Szabó & Z. Czere (Coll. MWM / ZSM); 1 female, Changwat Chiang Mai, 20 km NW of Mae Ai, 1650 m, 7. I. 1999, leg. A. Szabó & Z. Czere (Coll. MWM / ZSM); 1 male, N Thailand / Chiangmai Prov., NE Chiangmai, Doi Saket Distr., ESE Hang vic., Doi San Yao, ca. 1260 m NN, 20 / 18 ° C, LF, 12. III. 1986 / 18.30 – 0.30 h, leg. W. Nässig, H. Bänziger / C 2 (Coll. MWM / ZSM); 2 males, Changwat Chiang Mai, Mt. Doi Inthanon, NP, 2300 m, 17. X. 2000, leg. local collector, slides ZSM Arct. 2019 - 418, 2019 - 419 Volynkin (Coll. MWM / ZSM); 2 males, Changwat Chiang Mai, 4 km SE of Pang Faen, 1100 m, 13. I. 2004, leg. Péter Hentzshel & Atilla Szabó, slides ZSM Arct. 2019 - 437, 2019 - 438 Volynkin (Coll. MWM / ZSM); 1 female, Mt. Doi Phahompok, 14 km NW of Fang, 1700 m, 16. X. 2000, leg. local collector, slide ZSM Arct. 2019 - 420 Volynkin (Coll. MWM / ZSM); 1 male, 1 female, Changwat Mae Hong Son, 10 km NE of Pai, 1560 m, 3. XII. 1998, leg. M. Hreblay, Y. Sherpa & I. Soós (Coll. MWM / ZSM); 1 female, Changwat Mae Hong Son, 21 km NW of Pai, 1360 m, 7. II. 1998, leg. Márton Hreblay & Czaba Szabóky (Coll. MWM / ZSM); 1 male, 1 female, Mae Hong Son prov., 10 km E Khun Yuam, 950 m, 18 ° 47 ’ 10 ’’ N 98 ° 3 ’ 4 ’’ E, forest edge, 1. V. 2006, leg. K. Černý, slides ZSM Arct. 2019 - 600 (male), 2019 - 601 (female) Volynkin (Coll. MWM / ZSM); 1 male, 1 female, Changwat Nan, 25 km N of Bo Luang, 1150 m, 14. I. 1999, leg. A. Szabó & Z. Czere (Coll. MWM / ZSM); 1 female, the same locality, but, 11. XI. 1999, leg. Márton Hreblay (Coll. MWM / ZSM); 1 female, Changwat Nan, 22 km N of Bo Luang, 1120 m, 15. I. 1999, leg. A. Szabó & Z. Czere (Coll. MWM / ZSM); 1 male, Changwat Nan, 30 km E of Pua, 1700 m, 10. XI. 1999, leg. Márton Hreblay (Coll. MWM / ZSM); 1 male, the same locality, but 1. III. 1998, leg. Márton Hreblay & Csaba Szabóky, slide ZSM Arct. 2019 - 416 Volynkin (Coll. MWM / ZSM); 1 female, Lampang, Mae Chaem, 1453 m, 18 ° 56.119 ’ N, 99 ° 23.481 ’, 9. V. 2008, leg. K. Černý (Coll. CKC); VIETNAM: 1 male, North Vietnam, Tam Dao (secondary forest), 60 km NW Hanoi, 1200 m (21.34 ’ N, 105.20 ’ E), 1 – 15. XI. 1992, leg. Sinyaev & Simonov (Coll. MWM / ZSM); 7 males, 5 females, the same locality and collectors, but 1 – 5. V. 1993, leg. Sinajev & Simonov (Coll. MWM / ZSM); 1 female, North Vietnam, Mai-chau, 25 km S Moc-chau, 1400 m, 20.50 ’ N / 104.40 ’ E, 14 – 18. XI. 1994, secondary forest, leg. V. Sinyaev (Coll. MWM / ZSM); 1 female, N Vietnam / Tonkin, Mt. Fan-si-pan (North), Cha-pa, cloud forest (22.15 ’ N, 103.45 ’ E), 2 – 4. III. 1995, 2240 m, leg. Dr. R. Brechlin (Coll. MWM / ZSM); 1 female, N Vietnam, 1600 m, Mt. Fan-si-pan (North), Cha-pa, primary forest, 22 ° 17 ’ N 103 ° 44 ’ E, 25 – 30. III. 1995, leg. V. Sinyaev & A. Schintlmeister, slide ZSM Arct. 2019 - 641 Volynkin (Coll. MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB382410AA063ACFA29BAA3.taxon	diagnosis	Diagnosis. Forewing length is 8.5 – 10 mm in males and 10.5 – 12 mm in females. The species is highly variable in its size and forewing width. However, the male genitalia are uniform in all the populations examined. The male genitalia of H. pseudolutara are most similar to those of H. kutzscheri sp. n., but differ by the less elongate distal saccular process, more massive aedeagus, narrower distal chamber of aedeagus vesica, and much larger distal diverticulum. The female genitalia of H. pseudolutara differ from those of H. kutzscheri sp. n. by the broader ductus bursae, narrower corpus bursae, larger spines at the junction of the corpus bursae with ductus bursae, weaker scobination of the anterior section of corpus bursae, and sclerotized basal section of appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB382410AA063ACFA29BAA3.taxon	distribution	Distribution. Nepal, northeastern India (Mizoram and Assam), northern Myanmar (Kachin State), southwestern China (Yunnan Province), North Thailand (Chiang Mai, Lampang, Nan and Mae Hong Son Provinces) (Černý & Pinratana 2009, as Lyclene unguifera, part) and North Vietnam (Hòa Bình, Vĩnh Phúc and Lào Cai Provinces).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB1824C0AA065B4FF43B9DC.taxon	description	(Figs 19 – 22, 44, 45, 60)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB1824C0AA065B4FF43B9DC.taxon	materials_examined	Type material examined. Photographs of the lectotype (designated herein) (Fig. 19): female, “ Karapin, VIII. 11. Japan, H. Sauter ” / “ Asura obsoleta Mr. ♀ Strand det. ” (Coll. SDEI). Paralectotypes: 1 female, “ Kosempo Formosa H. Sauter, X. 11 ” / “ Asura obsoleta Mr. ♀ Strand det. ” / “ Asura undulosa Wkr. det F. Daniel ” (Coll. SDEI); 2 females, “ Kosempo Formosa H. Sauter, I. 10 ” / “ Asura obsoleta Mr. ♀ Strand det. ” (Coll. SDEI). Additional material examined. TAIWAN: TAOYUAN: 15 males, 49 females, Ming Chyr Forest Recreation Area, 1160 m, 13 – 14. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 1 male, the same locality, 29. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 1 female, the same locality, 05 – 06. X. 1996, leg. Gy. Fábián & F. Nemes (Coll. MWM / ZSM); 2 males, 2 females, the same locality, 17 – 18. IV. 1997, leg. Gy. Fábián & S. T. Kovács (Coll. MWM / ZSM); 1 male, 2 females, the same locality, 24 – 26. VI. 1997, leg. S. T. Kovács (Coll. MWM / ZSM); 11 males, 21 female, 14 km E of Fuhsing, 121 ’ 23 ’’ E – 24 ’ 50 ’’ N, 800 m, 18. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 2 males, 4 females, the same locality and collectors, 24 – 25. V. 1995 (Coll. MWM / ZSM); 1 male, the same locality and collectors, 31. V. 1995 (Coll. MWM / ZSM); 4 females, 14 km E of Fuhsing, 800 m, 4. X. 1995, 121 ’ 23 ’ E, 24 ’ 50 ’ N, lg. Csovary & Steger (Coll. MWM / ZSM); 1 male, the same locality and collectors, 28. X. 1995 (Coll. MWM / ZSM); 1 female, the same locality and collectors, 4. X. 1995 (Coll. MWM / ZSM); 1 male, 3 females, 7 km E of Fuhsing, 121 ’ 23 ’’ E – 24 ’ 49 ’’ N, 600 m, 20. XII. 1997, leg. S. Simonyi & A. Szabo (Coll. MWM / ZSM); 1 male, the same locality and collectors, 25. XII. 1997 (Coll. MWM / ZSM); 1 male, 5 females, the same locality and collectors, 28 – 29. XI. 1997 (Coll. MWM / ZSM); 1 male, the same locality, 17. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 1 male, 1 female, 16 km E of Fuhsing, 121 ’ 27 ’’ E, 24 ’ 50 ’’ N, 30. XI. – 1. XII. 1997, leg. S. Simonyi & A. Szabo (Coll. MWM / ZSM); 2 females, 16 km E of Fuhsing, 870 m, 27. X. 96 (121 ° 24 ’ E, 24 ° 50 ’ N), leg. T. Csovari & C. Szaboky (Coll. MWM / ZSM); TAIPEI: 1 male, Hsiapen, 650 m, 11. VII. 1997, leg. B. Herczig & Y. Y. Lien, slide ZSM Arct. 2019 - 975 Volynkin (Coll. MWM / ZSM); 1 female, Pi Hu, 410 m, 22. VI. 1997, leg. B. Herczig & L. Ronkay (Coll. MWM / ZSM); 1 male, 1 female, the same locality, 26. III. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 1 male, 4 females, Pi Hu, 450 m, 24 ° 54 ’ 02 ’’ N, 121 ° 45 ’ 27 ’’ E, 4 – 5. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 1 female, 10 km SE of Pingling, Pihou, 450 m, 3. V. 1997, leg. Gy. Fábián & S. T. Kovács (Coll. MWM / ZSM); 1 male, 2 females, the same locality and collectors, 16. IV. 1997 (Coll. MWM / ZSM); 2 females, the same locality, 26. III. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 1 female, the same locality, 04. X. 1996, leg. Gy. Fábián & F. Nemes (Coll. MWM / ZSM); 1 male, 10 km NE Pingling, Pihou, 550 m, 07. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 2 males, 6 females, the same locality and collectors, 01. VI. 1997 (Coll. MWM / ZSM); 1 male, 1 female, Wulai, 24 ° 50 ’ N, 121 ° 32 ’ E, 300 m, 13. II. 1999, leg. L. Peregovits & G. Ronkay (Coll. MWM / ZSM); 2 females, Formosa / Taiwan, Berge bei Taipei, 1980, ex coll. Kamper / Lödl (Coll. MWM / ZSM); KAOHSIUNG: 6 males, 11 females, 15 km NE Taoyuan, 1850 m, 07. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 1 male, 1 female, the same locality and collectors, 12. III. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 2 females, 3 km E Chiashen, 460 m, 18. X. 1996, leg. Gy. Fábián & F. Németh (Coll. MWM / ZSM); 1 male, 2 females, 700 m, Liu-Kuei, San-Ping Forest Res. Stat., 22 ° 58 ’ 16 ’’ N, 120 ° 41 ’ 15 ’’ E, 14 – 15. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 1 male, 2 females, 26 km SE of Taoyüan, 120 ‘ 52 ‘‘ E – 23 ‘ 17 ‘‘ N, 1370 m, 19. III. 1996, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 1 male, 1 female, the same locality, 4. XI. 96, leg. T. Csovari & C. Szaboky (Coll. MWM / ZSM); 3 males, the same locality, 19. III. 96, leg. T. Csovári & P. Stéger (Coll. MWM / ZSM); 1 female, Alikang, Formosa, Sauter, XI. 09, Samml. F. Daniel (Coll. MWM / ZSM); HUALIEN: 1 male, Taroko N. P., Hsipan, 270 m, 5. IV. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 2 males, 1 female, 6 km SE Juisui, 200 m, 22. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 1 male, 10 km NE of Tienchih, 121 ’ 20 ’’ E – 24 ’ 04 ’’ N, 2100 m, 23. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); ILAN: 1 male, 1 female, 150 km SE Taipei, 400 m, 24 ° 23 ’ 43 ’’ N, 121 ° 46 ’ 74 ’’ E, 10. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 17 mals, 25 females, 1200 m, Ming Chyr Forest Recreation Area, 8 – 9. VII. 1997, leg. B. Herczig & S. Kovács (Coll. MWM / ZSM); 6 males, 4 females, the same locality, 4 – 5. VI. 1997, leg. B. Herczig & L. Ronkay (Coll. MWM / ZSM); 2 males, the same locality, 30 – 31. III. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 1 female, 700 m, Fu Shan Botanical Garden, 27 – 28. III. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 4 females, 2 km N Suao, 130 m, 01. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 1 male, 3 females, 5 km SW of Ilan, 290 m, 25. II. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 2 females, Fu Shan Bot. Garden, LTER site, 700 m, 24 ° 45 ’ 13 ’’ N, 121 ° 35 ’ 39 ’’ E, 8 – 9. IV. 1997, leg. L. Peregovits & A. Kun (Coll. MWM / ZSM); 2 males, 13 females, 3 km SS of Suao, 121 ‘ 51 ‘‘ E – 24 ‘ 34 ‘‘ N, 400 m, 26. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 9 females, 5 km S of Suao, 400 m, 6. X. 1995, 121 ‘ 51 ‘ E, 24 ‘ 34 ‘ N, lg. Csovary & Steger (Coll. MWM / ZSM); 1 male, 3 km E of Suao, 121 ‘ 51 ‘‘ E – 24 ‘ 34 ‘‘ N, 400 m, 14. III. 1996, leg. T. Csovary & P. Steger (Coll. MWM / ZSM); 1 male, 1150 m, Ming Chyr, 24 ° 39 ’ N 121 ° 28 ’ E, 11 – 12. IV. 1999, leg. G. Csorba & B. Herczig, slide AV 5484 (male) Volynkin (Coll. CKC); TAITUNG: 1 female, Chihpen Hot Springs, 400 m, 6. VII. 1997, leg. B. Herzig & S. T. Kovács (Coll. MWM / ZSM); 6 females, the same locality, 6. IV., 9. IV. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 1 female, Mutien, 1500 m, 21. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 9 males, 1 female, 5 km W of Chipen, 350 m, 09. III. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 3 males, the same locality, 05. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 1 female, the same locality, 15 – 16. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 1 male, 1 female, Hsianguang, Police station, 2320 m, 7. VII. 1997, leg. S. T. Kovács (Coll. MWM / ZSM); 3 females, 2 km N of Tupan, 500 m, 16. X. 1995, 120 ’ 52 ’ E, 22 ’ 29 ’ N, lg. Csovari & Steger (Coll. MWM / ZSM); 2 females, the same locality, 29. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 2 males, 1 female, 5 km W Hungyeh vill., 120 ’ 51 ’’ E, 22 ’ 28 ’’ N, 350 m, 3. XII. 1997, leg. S. Simonyi & A. Szabo (Coll. MWM / ZSM); 5 males, 1 female, 5 km NW of Lirao, 120 ’ 59 ’’ E – 23 ’ 13 ’’ N, 1760 m, 28. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 4 males, 2 females, 7 km N of Tupan, 120 ’ 52 ’’ E – 22 ’ 29 ’’ N, 500 m, 9 – 10. XII. 1997, leg. S. Simonyi & A. Szabo (Coll. MWM / ZSM); 1 male, 4 km N of Tupan, 390 m, 1. VII. 1997, 120 ’ 52 ’’ E, 22 ’ 28 ’’ N, lg. Csövári & Mikus (Coll. MWM / ZSM); 1 female, 350 m, 3 km W village Hungyeh, 30. VI. 1997, 120 ’ 52 ’’ E, 22 ’ 28 ’’ N, lg. Csövari & Mikus (Coll. MWM / ZSM); 1 female, 3 km W Hungyeh village, 350 m, 5. XII. 96, (120 ° 52 ’ E, 22 ° 28 ’ N), leg. T. Csovari & C. Szaboky (Coll. MWM / ZSM); 12 males, 20 females, 7 km N of Tupan, 500 m, 03 – 06. II. 1997, leg. S. Simonyi & P. Steger (Coll. MWM / ZSM); 7 males, 7 females, the same locality and collectors, 14 – 16. II. 1997 (Coll. MWM / ZSM); 1 female, the same locality, 20. III. 1996, leg. T. Csovári & P. Stéger (Coll. MWM / ZSM); CHIAYI: 1 male, 4 km NE Alishan, 2200 m, 12. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); 4 males, 5 females, 1 km E Yungsing, 500 m, 13. V. 1997, leg. Gy. M. László & G. László (Coll. MWM / ZSM); NANTOU: 1 female, Tayuling, 2750 m, 28. VI. 1997, leg. S. T. Kovács (Coll. MWM / ZSM); 7 males, 7 females, 15 km N of Puli, 500 m, 04. III. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 1 female, the same locality and collectors, 14 – 15. X. 1996 (Coll. MWM / ZSM); 4 females, the same locality and collectors, 28. II. 1996 (Coll. MWM / ZSM); 2 females, the same locality, 29 – 30. VI. 1997, leg. S. T. Kovács (Coll. MWM / ZSM); 4 males, 11 females, 15 km N Puli, Uisun Forest Area, 500 m, 09 – 10. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 5 males, 8 females, 3 km S Hoshe, Yu Shan Nat. Park, 1400 m, 16. X. 1996, leg. Gy. Fábián & F. Nemes, slide ZSM Arct. 2019 - 977 (female) Volynkin (Coll. MWM / ZSM); 1 female, 10 km SE of Shenmu, Yu-Shan Nat. Park, 2200 m, 14. III. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 1 female, Huisun Forest Area, 500 m, 15 km N Puli, 12 – 13. IV. 1997, leg. Csorba & Ronkay (Coll. MWM / ZSM); 2 females, the same locality, 7 – 8. VI. 1997, leg. B. Herczig & I. Ronkay (Coll. MWM / ZSM); 1 male, 1 female, Tungpu, YuShan Nat. Park, 1400 m, 24 – 25. IV. 1997, leg. Gy. Fábián & S. T. Kovács (Coll. MWM / ZSM); 1 male, 1 female, 5 km SW of Tayüling, 121 ’ 17 ’’ E – 24 ’ 09 ’’ N, 2900 m, 21. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 4 females, 3 km E of Tili, 120 ’ 58 ’’ E – 23 ’ 47 ’’ N, 500 m, 17. III. 1996, leg. T. Csovary & P. Steger (Coll. MWM / ZSM); 1 female, the same locality, 27 – 28. V. 1997, C. Szaboky & I. Soos (Coll. MWM / ZSM); 4 females, the same locality, 2. XI. 1996, leg. T. Csovari & C. Szaboky (Coll. MWM / ZSM); 2 females, 3 km E of Tili, 555 m, 08 – 09. II. 1997, leg. S. Simonyi & P. Steger (Coll. MWM / ZSM); 2 females, 3 km SW of Tsuifeng, 2100 m, 25 / 26. VI. 1997, 121 ’ 10 ’’ E, 24 ’ 06 ’’ N, lg. Csővary & Mikus (Coll. MWM / ZSM); 3 males, 2 females, 17 km E of Kuchsing, 121 ’ 00 ’’ E – 24 ’ 05 ’’ N, 560 m, Ursun Forest Camping, 6 / 7. VI. 97, lg. C. Szaboky & I. Soos (Coll. MWM / ZSM); 1 male, the same locality and collectors, 16. VI. 97 (Coll. MWM / ZSM); 2 females, the same locality and collectors, 8. VI. 1997 (Coll. MWM / ZSM); 1 female, 3 km SW of Tsuifeng, 2100 m, 7. VII. 1997, 121 ° 10 ’ E, 24 ° 06 ’ N, Tibor Csővári & László Mikus (Coll. MWM / ZSM); 1 female, the same locality, 31. X. 96, leg. T. Csovary & C. Szaboky (Coll. MWM / ZSM); 1 male, the same locality, 26 – 27. X. 1995, Tibor Csővári & Pál Stéger (Coll. MWM / ZSM); 3 males, 5 females, the same locality and collectors, 11. X. 1995 (Coll. MWM / ZSM); 1 female, the same locality and collectors, 16. III. 1996 (Coll. MWM / ZSM); 1 male, the same locality, 22. V. 1995, leg. M. Hreblay & P. Steger (Coll. MWM / ZSM); 1 male, 2 females, Ursun F. [orest], 16 km E of Kuohsing, 560 m, 29 – 30. X. 96 (121 ° 00 ’ E, 24 ° 05 ’ N), leg. T. Csovari & C. Szaboky (Coll. MWM / ZSM); 1 male, the same locality and collectors, 18. XI. 96 (Coll. MWM / ZSM); 2 females, the same locality, 15. VII. 1997, leg. Tibor Csővári & László Mikus (Coll. MWM / ZSM); 1 female, Central Taiwan, near Sun Moon Lake, Hosher, 800 m, 23 ° 35 ’ 32 ’’ N 120 ° 63 ’ 16 ’’ E, 20. V. 2001, leg. U. Buchsbaum, slide AV 5485 (female) Volynkin (Coll. CKC); PINGTUNG: 11 males, 25 females, 10 km SE of Mutan, 470 m, 03 – 04. VII. 1996, leg. G. Csorba & L. Németh (Coll. MWM / ZSM); 7 males, 12 females, the same locality and collectors, 07 – 10. III. 1996 (Coll. MWM / ZSM); 2 females, the same locality, 22. X. 1996 (Coll. MWM / ZSM); 2 males, 2 females, the same locality, 26. IV. 1997 (Coll. MWM / ZSM); 1 male, the same locality, 17. V. 1997 (Coll. MWM / ZSM); 1 male, the same locality, 5. VII. 1997 (Coll. MWM / ZSM); 7 males, 2 females, the same locality, 7 – 8. IV. 1997 (Coll. MWM / ZSM); 2 males, 2 females, Huang-Lion Forest Recreation Area, 210 m, 06. III. 1996, leg. Gy. Fábián & L. Németh (Coll. MWM / ZSM); 1 female, 5 km W of Chipen, 470 m, 27 – 28. IV. 1997, leg. Gy. Fábián & S. T. Kovács (Coll. MWM / ZSM); TAICHUNG: 1 female, An-ma-Shan, Hooping, 2000 m, 1 – 2. VII. 1997, leg. S. T. Kovács (Coll. MWM / ZSM); 1 female, Hui Sun Exp. Forest, Guandashi LTER site, 950 m, 24 ° 04 ’ 49 ’’ N, 121 ° 02 ’ 08 ’’ E, 12 – 13. IV. 1997, leg. Peregovits & Kun (Coll. MWM / ZSM); 1 male, 4 females, Anmashan, lower forest zone, 1650 m, 20. VI. 1997, leg. B. Herczig & L. Ronkay (Coll. MWM / ZSM); 1 male, Tahsuehshan Forest Recreation Area, 2000 m, 25. V. 1997, leg. Gy. M. László & G. László, slide ZSM Arct. 2019 - 976 Volynkin (Coll. MWM / ZSM); MIAOLI: 5 males, 6 females, Taiwan, Prov. Miaoli, 20 km E of Tungshih, 1335 m, 18. X. 1995 (121 ° 03 ‘ E, 24 ° 19 ‘ N), leg. T. Csovari & P. Steger (Coll. MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB1824C0AA065B4FF43B9DC.taxon	discussion	Remark. In the original description of Asura obsoleta Form alikangiae, Strand (1917) cited six specimens (five females and one male) from three localities (Kosempo, Alikang and Karapin), which are syntypes. Inoue (1988) upgraded this taxon to the species level and illustrated a female syntype. Here we designate this specimen (Fig. 19) as lectotype and fix the species’ type locality as “ Karapin ” (Taiwan, Chiayi County, Chaoliping).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB1824C0AA065B4FF43B9DC.taxon	diagnosis	Diagnosis. Forewing length is 10 – 11 mm in males and 11 – 14 mm in females. Huangilene alikangiae differs clearly from other congeners by its medial transverse line of forewing being nearly straight or slightly wavy, whereas in other Huangilene species that is strongly wavy. The male genital capsule of H. alikangiae is also characteristic due to the broad distal membranous lobe of valva having a dentate dorsal margin distally and the small lobes of anellus bearing very small denticles. The aedeagus vesica of H. alikangiae is characterized by the large dorsal medial diverticulum, and the narrow and elongate ventral medial diverticulum bearing a rather small cornutus. The female genitalia of H. alikangiae can be distinguished from those of other congeners by the large lateral protrusion of corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFB1824C0AA065B4FF43B9DC.taxon	distribution	Distribution. Taiwan Island (Daniel 1952, as Lyclene (Asura) undulosa, part; Inoue 1988; Chang 1989; Wang 1994).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBF824C0AA06018FC78BA5B.taxon	description	(Figs 23 – 25, 46, 47, 61, 62)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBF824C0AA06018FC78BA5B.taxon	materials_examined	Material examined. JAPAN: 1 male, Shiratani, Yakushima, 25. VII. 1974, H. Inoue, slide ZSM Arct. 2020 - 148 Volynkin (Coll. MWM / ZSM); 1 female, Kosugidani, Yakushima, 4. VIII. 1954, K. Ishizuka, slide ZSM Arct. 2020 - 149 Volynkin (Coll. MWM / ZSM); 2 males, Yuwandake, 11. VIII. 1961, Amami-Oshima, A. Kawazoe / Inoue Coll. B. M. 1992 - 71 / unique numbers NHMUK 010918603, NHMUK 010604733, slide NHMUK 010316346 Volynkin (Coll. NHMUK); 1 female, Yuwandake, 7. VIII. 1961, Amami-Oshima, A. Kawazoe / Inoue Coll. B. M. 1992 - 71 / NHMUK 010316347, slide NHMUK 010316347 Volynkin (Coll. NHMUK).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBF824C0AA06018FC78BA5B.taxon	diagnosis	Diagnosis. Forewing length is 8 – 10 mm in males and 10 – 11 mm in females. Huangilene alikangiae intermedia differs externally from the nominate subspecies by its thinner forewing markings with almost reduced medial transverse line. The genitalia of the two subspecies display no remarkable differences.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBF824C0AA06018FC78BA5B.taxon	distribution	Distribution. Japan: the Ryukyu Islands (Marumo 1923; Inoue 1988).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBF824D0AA0629AFAAFBC7D.taxon	diagnosis	Diagnosis. The male genitalia of the species-group are characterized by the relatively narrow medial costal process consisting of two trigonal lobes with blunt tips, of which the distal one is larger and serrulate; the elongate and serrulate distal section of costa (Fig. 51), the large membranous lobe of valva, the relatively short, hook-like curved distal saccular process, the aedeagus lacking distal sclerotized protrusions to the vesica, the strongly elongate medial ventral diverticulum of vesica bearing a robust cornutus with a massive, elongate base, which is equal in length or longer than the cornutus itself. The female genitalia are characterized by the narrow and elongate corpus bursae with a lateral longitudinal cluster of shortly spinulose scobination and the large, sack-like appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBE824D0AA064B8FA96BB6C.taxon	description	(Figs 26 – 29, 48, 49, 51, 63)	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBE824D0AA064B8FA96BB6C.taxon	materials_examined	Type material. Holotype (Figs 26, 48): male, “ N-Vietnam, 1600 m, Mt. Fan-si-pan (Nord), Cha-pa, Primärurwald [primary forest] 22.17 ’’ N 103.44 ’’ E, 25. – 30. III. 1995, leg. V. Sinjaev [Sinyaev] & A. Schintlmeister ”, slide ZSM Arct. 2019 - 640 Volynkin (Coll. MWM / ZSM). Paratypes (12 males and 4 females in total). VIETNAM: 1 male, the same data as in the holotype, slide ZSM Arct. 2019 - 417 Volynkin (Coll. MWM / ZSM); 1 female, North Vietnam, Mt. Fan-si-pan, Cha pa, 2400 m, NN (22 ° 15 ’ N 103 ° 46 ’ E) 8 – 29. V. 1993, leg. Sinyaev & Simonov, slide ZSM Arct. 272 / 2017 Volynkin (Coll. MWM / ZSM); 1 male, N Vietnam / Tonkin, Mt. Fan-si-pan (North), Cha-pa, cloud forest (22 ° 15 ’ N 103 ° 45 ’ E) 2 – 4. III. 1995, 2240 m, leg. Dr. R. Brechlin, slide ZSM Arct. 268 / 2017 Volynkin (Coll. MWM / ZSM); 6 males, 2 females, North Vietnam, Tam Dao, (secondary forest), 60 km NW Hanoi, 1200 m, (21 ° 34 ’ N, 105 ° 20 ’ E), 1 – 5. V. 1993, leg. Sinyaev & Simonov, slides ZSM Arct. 269 / 2017 (male), 271 / 2017 (female) Volynkin (Coll. MWM / ZSM); 1 male, North Vietnam, Cuc Phuong, 60 km SW Hanoi, 400 m, (20 ° 15 ’ N, 105 ° 20 ’ E), 18. XI. – 3. XII / [19] 92, leg. Sinyaev & Simonov (Coll. MWM / ZSM); 2 males, North Vietnam, Tam Dao, 60 km NW Hanoi, 950 m, (21 ° 34 ’ N, 105 ° 20 ’ E), 17. X. 1994, leg. V. Sinyaev, slides ZSM Arct. 2019 - 638, 2019 - 639 Volynkin (Coll. MWM / ZSM); 1 male, 1 female, South Vietnam, Bao Loc (secondary forest), Rung Cat Tien, 1500 m (11 ° 32 ’ N, 107 ° 48 ’ E), 10 – 20. XII. 1992, leg. Sinyaev & Simonov, slide ZSM Arct. 270 / 2017 (male) Volynkin (Coll. MWM / ZSM); 1 male, 13. VIII. 2019, Central Vietnam, Th ừa Thiên-Hu ế Province, Annamite Mts, B ạch Mã National Park, 1250 m, 16 ° 11 ’ 34 ’’ N 107 ° 51 ’ 12 ’’ E, edge of tropical forest, Ye. A. Derzhinsky leg., slide AV 5898 Volynkin (Coll. CAV).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBE824D0AA064B8FA96BB6C.taxon	diagnosis	Diagnosis. Forewing length is 8 – 10 mm in males and 10 – 10.5 mm in females. The new species differs externally from its congeners by the more strongly dentate postmedial transverse line of forewing. The male genital capsule of H. apoklinousa sp. n. is very characteristic and can be easily distinguished from those of other Huangilene species by the distally broadened valva, bilobate and not dentate medial costal process, elongate and serrulate distal section of costa, large distal membranous lobe of valva bearing a longitudinal weakly serrulate area protruding to the distal lobe of the medial costal process, and short, hook-like curved distal saccular process. The distal section of aedeagus of the new species is evenly serrulate and lacks protrusions to the vesica. The vesica structure of H. apoklinousa sp. n. is also unique within the genus and is characterized by its not scobinated subbasal ventral diverticulum, strongly elongate ventral medial diverticulum bearing a very robust spine-like cornutus having a very long base protruding inside the diverticulum, and short, narrow and scobinated medial ventral and distal diverticula. The female genitalia of H. apoklinousa sp. n. differ clearly from those of other congeners by their narrow and elongate corpus bursae having a lateral longitudinal cluster of shortly spinulose scobination (absent in other Huangilene species), and the large, sack-like appendix bursae (whereas in other Huangilene species that is short and conical).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
0389D55CFFBE824D0AA064B8FA96BB6C.taxon	distribution	Distribution. Vietnam (Lào Cai, Ninh Bình, Vĩnh Phúc, Th ừa Thiên-Hu ế and Lâm Đ ồng Provinces).	en	Volynkin, Anton V., Černý, Karel (2020): Huangilene, a new genus for the Miltochrista alikangiae (Strand, 1917) species-group, with descriptions of three new species from Indochina, continental China and Taiwan (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4869 (1): 91-111, DOI: 10.11646/zootaxa.4869.1.4
