taxonID	type	description	language	source
0389BB02A52D7250FF0A0F30532BEB67.taxon	description	(Figs 1 - 4) urn: lsid: zoobank. org: act: AABCD 02 F-B 12 C- 4731 - BED 4 - 148 E 1 F 53 C 45 D	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	materials_examined	Holotype: ♂ NMSUK, Papua New Guinea, Enga Province, Mt Hagen, Maropi Creek, 2550 m, 22.12.201 6, N. Grimaldi. Paratypes: (7 ♂♂, 3 ♀♀) 1 ♂ NG, Mt Hagen, Maropi Creek, 2550 m, 24.12.2016; 1 ♂ CD, Mt Hagen Maropi Creek, 2550 m, 24.12.2016; 1 ♂ OP, Mt Hagen, Maropi Creek, 2550 m, 24.12.2016; 1 ♂ NG, Mt Hagen, Pap Creek, 2650 m, 24.12.2016; 1 ♂ NG, Mt Hagen, Kumul Lodge, 2850 m, 26.12.2016; 1 ♂ NG, Mt Hagen, Maropi Creek, 2550 m, 07.09.2017; 1 ♂ PS, Pap Ck, Hagen Ra. 2800 m, 12. xi. 1973; 1 ♀ NG, Mt Hagen, Maropi Creek, 2550 m, 24.12.201 6; 2 ♀♀ NG, Mt Hagen, Maropi Creek, 2550 m, 08.09.201 7.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	diagnosis	Diagnosis: The subspecies is distinguished from previously described races of clathrata by the grey brown coloration of the pale areas, the narrow vein markings and smaller yellow basal spot on the hindwing underside.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	description	Description: Male (Figs 1 - 2) Wingspan 55 mm, forewing length 30 mm. Upperside: forewing ground color white, black costal border partly absorbing black discocellular bar, black border along termen widening toward apex, indented on inner edge at cubital veins, 3 creamy-white subapical spots. Upperside of hindwing is white with a thin black terminal border. Underside: forewing white with black costal and terminal borders narrower than in the nominate subspecies, discal cell incompletely filled with black, connected to termen on vein R 3. Three orange-yellow subapical spots and two smaller terminal spots. Underside of hindwing: pale grey-brown with a triangular black sub-basal area from costa toward top of inner margin, crossing discal cell at sub-basal level and including a yellow basal spot that is anteriorly white and divided from the costal margin by a pale streak. Median area dark grey to sepia, divided by narrow, pale grey vein markings. The size and shade of the median patch is variable and more diffuse in flight-worn individuals. Black postmedian spot in space M 2 - M 3 that interrupts the pale marking of vein M 2 in approximately 50 % of individuals. Terminal area pale grey, variable in width, shading to white at tornus in some specimens. Anal area with yellow and black scaling. Female (Fig 4) Wingspan 50 mm, forewing length 28 mm. Upperside: forewing ground color white, black costal border partly absorbing broad black dc bar. Black border along termen narrower and less serrated than in the nominate subspecies, widening toward apex, inner edge dusted with grey, three creamy-white subapical spots. Hindwing white with narrower black terminal border than the nominate. Underside: forewing white with black costal and terminal borders, discal cell incompletely filled with black extending to the termen on vein R 3. Three orange-yellow subapical spots and 2 smaller terminal spots. Underside of hindwing is pale coffee-brown with triangular black subbasal area and yellow basal spot as in the male. Median area dark sepia brown, wider and more diffused than in other races and divided by narrower vein markings. Black median spot in space M 2 - M 3. Anal area with more pronounced yellow scaling than male.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	distribution	Distribution: Mt Hagen, Western Highland Province, Papua New Guinea. Records of the species from the Wabag, Enga Province and Tari, Southern Highlands Province require confirmation.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	etymology	Etymology: The subspecies is named after Michael Parsons who first mentioned and illustrated the new taxon in his seminal work ‘ The Butterflies of Papua New Guinea. Their Systematics and Biology’ (Parsons, 1998).	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52D7250FF0A0F30532BEB67.taxon	discussion	Discussion: The population of D. clathrata on Mt Wilhelm exhibits a variety of phenotypes that intergrade between the nominate subspecies and ssp limata from Wau. These suggest that limata should be considered a synonym of the nominate subspecies. The population from Gumine in south Simbu province also has a variable phenotype, the hindwing base colour ranging from white to coffee brown. The darkest of these resemble ssp. nov. parsonsi but are distinguished on the hindwing underside by the grey colour and elongated shape of the hindwing basal streak, more sharply defined median patch and broader white vein markings. The female has not been recorded and the identity of the population remains uncertain. Though less than 40 km from Mt Wilhelm, Gumine is situated at the eastern extreme of the Kubor Range that extends westward to the Mt Hagen massif, and may therefore be a hybrid zone.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0B255024E924.taxon	description	(Figs 23 - 24)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0B255024E924.taxon	discussion	This species is commonly found in sympatry with D. hallstromi at elevations between 2,500 – 3,000 meters in central PNG. The female has rarely been collected and is not illustrated by Parsons (1998) or Yagishita (1993). It is distinguished from the male by a paler yellow ground colour and more diffused black margins on the upperside wings, and by orange coloration on the underside of the abdomen.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0CD851E3EB54.taxon	description	(Figs 17 - 18)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0CD851E3EB54.taxon	distribution	Delias hagenensis was described from material collected at Pap Creek, Walya, 3000 m in August 1973 and has since been recorded from Mt Hagen, Laiagam and Porgera (Parsons, 1998) and Tari (Lachlan, 2000). On Mt Hagen it is a common species however until now the female has not been recorded or illustrated in a publication.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0CD851E3EB54.taxon	description	Description of female: Broadly similar to male but distinguished on the upperside forewing by the suffusion of grey scales in the basal area and beyond the discocellular vein, (uniformly black in the male), and more prominent white subapical spots. The basal area of the hindwing has a slightly blue tinge in fresh specimens and contains yellow streaks at the costal and anal border. On the underside, the forewing discocellular bar is broader than in males and the white subapical spots are larger, particularly in space M 1 – M 2.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0A3D51E9EA33.taxon	description	(Figs 19 - 22)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0A3D51E9EA33.taxon	distribution	Delias hallstromi is a common species in central PNG however the female is rarely encountered. Parsons (1998) illustrates a specimen in the ANIC from Marifunga in the Eastern Highlands. On Mt Hagen, females were frequently seen at locations 1 and 2 feeding on flowering plants. Individual variation is seen in the width of the discocellular bar on the forewing underside and the extent of white markings on the forewing upperside.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7256FF0A08CD566AED15.taxon	description	(Figs 25 - 28)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7256FF0A08CD566AED15.taxon	discussion	The type locality of Delias mira roepkei Sanford & Bennett, 1955 is Mt Hagen. The taxon was raised to species level by Van Mastrigt (2000) who mentioned the presence of two forms in the Hagen population; one with broad black borders on the upperside, the other with narrower borders and paler yellow hindwing patch on the underside that he speculated might represent a separate species. Males collected at the Murmur Pass locations exhibit a continuum of individual variations in the upperside border depth and underside hindwing markings and are not readily dividable into two separate groups. Specimens from Mt Wilhelm (Simbu Province) and the Eastern Highlands are also variable and, on average, have a more peach-colored anal patch on the hindwing underside. Van Mastrigt’s decision to elevate roepkei to species level was based by the supposed sympatry of both roepkei and mira in parts of the Owen Stanley Mountains. Pending genetic analysis, the original taxonomy is adopted in this paper. Among a number of typically white coloured females, two individuals with yellow-colored uppersides have been recorded. (Figs 28 - 29). Similar yellow female forms have been recorded in a number of species of the D. iltis ̡ D. aroae and D. eichorni groups.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7256FF0A0CC2532AEB58.taxon	description	(Figs 29 - 30)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7256FF0A0CC2532AEB58.taxon	distribution	A new record for the Mt Hagen area. The species is widespread though less common in PNG than in Indonesian Papua and has previously been recorded from Mt Wilhelm and Tari (ssp. kerowagiensis Yagishita, 1993), the Owen Stanley Mts (ssp. owenstanleyi Schröder & Treadaway, 1994) the west Bismark Mts and Porgera in Enga province (CD).	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7256FF0A0CC2532AEB58.taxon	description	Male specimens from Mt Hagen are distinguishable from topotypical kerowagiensis by the narrower black median bands on the hindwing underside and from the Bismark and Porgera populations which lack red median bands on the hindwing. The identity and distribution of potential subspecies of D. hapalina in PNG will be reviewed after comparative study of these populations.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7255FF0A0A0E577EEEF1.taxon	description	(Figs 9 - 16)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7255FF0A0A0E577EEEF1.taxon	discussion	The subspecific identity of D. leucias from the central mountains of Papua New Guinea has a confusing history, having been assigned to the nominate race (type locality: Mt Goliath) by D’Abrera (1978), and subsequently to subspecies huonensis by later authors such as Parsons (1998), Yagishita et al. (1993) and Van Mastrigt (2003). Lachlan (2000) reports collection of Delias leucias leucias at Ok Tedi, Western Province and D. leucias huonensis at Tari, Southern Highlands. Gotts & Ginn (2004) described a new subspecies, D. leucias torini ̡ from the Snow Mts, west of the nominate range, and discussed variation in D. leucias across New Guinea, mentioning the differences between the Kerowagi (Mt Wilhelm) population and the nominate subspecies from the Star Mountains. They considered the taxon D. weiskei sayuriae Okano, 1989, recorded only from Mt Wilhelm, to be a variation of leucias rather than a subspecies of D. weiskei Ribbe, 1900. D. leucias huonensis Talbot, 1928 was described from a single male collected by Keysser in the isolated Rawlinson Mountains on the Huon peninsula in north east New Guinea. The race remains rare in collections and apparently in nature; William Brandt did not record the species during his expedition to the Rawlinson Mts in 1968. Specimens from Mt Hagen are similar to those from Mt Wilhelm & the Eastern Highlands, both can be distinguished in the majority of individuals from the nominate subspecies - by the complete separation of the pale hindwing patch from the white costal border by the red and black subapical bands, and a more orange coloration of the forewing underside - and from ssp huonensis by the narrower black margins on the both surfaces and the more elongated hind wing patch. Although these characters are consistent in the majority of specimens from central PNG, populations of the nominate race from various locations within Indonesian Papua show clinal development of the same features on a west - east axis. Delias leucias from central PNG is considered to represent a recognizable geographic form of the nominate subspecies.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A5297255FF0A0FE653CCECA0.taxon	description	(Figs 31 - 32)	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A5297255FF0A0FE653CCECA0.taxon	discussion	The type locality of D. callista porquaiensis Yagishita, 2003 is stated to be ’ Porquia, PNG’ however no such place name is gazetted in PNG and it is assumed that it is a misspelling of Porgera, the site of a large gold mine in western Enga Province, approximately 80 km west of Mt Hagen. Male specimens from Porgera and the Ok Tedi area agree with the original description of porquaiensis and are characterized by rich yellow coloration, broad black submarginal bands and triangular shaped black terminal vein markings on the hindwing underside. Males of D. callista from Mt Hagen consistently differ from this phenotype by having a paler shade of yellow on the hindwing underside, narrow subterminal bands and terminal vein markings. A similar form has been recorded from Gumine (IFTA). Females from Mt Hagen have a series of 5 white apical spots on the forewing upperside and 4 – 5 white subterminal spots on the hindwing that are absent or poorly developed in more western areas. The taxonomic status of D. callista populations in PNG and western Papua is uncertain and will be reviewed in a further publication.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A5297255FF0A0A5150CAEB88.taxon	discussion	The presence of this species in close proximity to, but at lower altitude than, its sister species D. gilliardi ̡ is of interest. A single male was recorded by a local collector at Aiyamp creek, on the lower western slopes of the Mt Hagen massif. The holotype and other specimens were collected by Phil Saywer in July 1975 at ‘ Poketamanda’ = Mt Nose, near Wabag, Enga Province, approximately 30 km west of Mt Hagen.	en	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
