identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0389BB02A52D7250FF0A0F30532BEB67.text	0389BB02A52D7250FF0A0F30532BEB67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias clathrata subsp. parsonsi Davenport & Grimaldi 2019	<div><p>Delias clathrata parsonsi subsp. nov.</p><p>(Figs 1-4)</p><p>urn:lsid:zoobank.org:act: AABCD02F-B12C-4731-BED4-148E1F53C45D</p><p>Holotype: ♂ NMSUK, Papua New Guinea, Enga Province, Mt Hagen, Maropi Creek, 2550m, 22.12.201 6, N. Grimaldi.</p><p>Paratypes: (7 ♂♂, 3 ♀♀) 1♂ NG, Mt Hagen, Maropi Creek, 2550m, 24.12.2016 ; 1♂ CD, Mt Hagen Maropi Creek, 2550m, 24.12.2016 ; 1♂ OP, Mt Hagen, Maropi Creek, 2550m, 24.12.2016 ; 1♂ NG, Mt Hagen, Pap Creek, 2650m, 24.12.2016 ; 1♂ NG, Mt Hagen, Kumul Lodge, 2850m, 26.12.2016 ; 1♂ NG, Mt Hagen, Maropi Creek, 2550m, 07.09.2017 ; 1♂ PS, Pap Ck, Hagen Ra. 2800m, 12.xi.1973; 1♀ NG, Mt Hagen, Maropi Creek, 2550m, 24.12.201 6 ; 2♀♀ NG, Mt Hagen, Maropi Creek, 2550m, 08.09.201 7 .</p><p>Diagnosis: The subspecies is distinguished from previously described races of clathrata by the grey brown coloration of the pale areas, the narrow vein markings and smaller yellow basal spot on the hindwing underside.</p><p>Description: Male (Figs 1-2) Wingspan 55mm, forewing length 30mm. Upperside: forewing ground color white, black costal border partly absorbing black discocellular bar, black border along termen widening toward apex, indented on inner edge at cubital veins, 3 creamy-white subapical spots. Upperside of hindwing is white with a thin black terminal border.</p><p>Underside: forewing white with black costal and terminal borders narrower than in the nominate subspecies, discal cell incompletely filled with black, connected to termen on vein R3. Three orange-yellow subapical spots and two smaller terminal spots. Underside of hindwing: pale grey-brown with a triangular black sub-basal area from costa toward top of inner margin, crossing discal cell at sub-basal level and including a yellow basal spot that is anteriorly white and divided from the costal margin by a pale streak. Median area dark grey to sepia, divided by narrow, pale grey vein markings. The size and shade of the median patch is variable and more diffuse in flight-worn individuals. Black postmedian spot in space M2-M3 that interrupts the pale marking of vein M 2 in approximately 50% of individuals. Terminal area pale grey, variable in width, shading to white at tornus in some specimens. Anal area with yellow and black scaling. Female (Fig 4) Wingspan 50mm, forewing length 28mm. Upperside: forewing ground color white, black costal border partly absorbing broad black dc bar. Black border along termen narrower and less serrated than in the nominate subspecies, widening toward apex, inner edge dusted with grey, three creamy-white subapical spots. Hindwing white with narrower black terminal border than the nominate.</p><p>Underside: forewing white with black costal and terminal borders, discal cell incompletely filled with black extending to the termen on vein R3. Three orange-yellow subapical spots and 2 smaller terminal spots. Underside of hindwing is pale coffee-brown with triangular black subbasal area and yellow basal spot as in the male. Median area dark sepia brown, wider and</p><p>more diffused than in other races and divided by narrower vein markings. Black median spot in space M2-M3. Anal area with more pronounced yellow scaling than male.</p><p>Distribution: Mt Hagen, Western Highland Province, Papua New Guinea. Records of the species from the Wabag, Enga Province and Tari, Southern Highlands Province require confirmation.</p><p>Etymology: The subspecies is named after Michael Parsons who first mentioned and illustrated the new taxon in his seminal work ‘The Butterflies of Papua New Guinea. Their Systematics and Biology’ (Parsons, 1998).</p><p>Discussion: The population of D. clathrata on Mt Wilhelm exhibits a variety of phenotypes that intergrade between the nominate subspecies and ssp limata from Wau. These suggest that limata should be considered a synonym of the nominate subspecies.</p><p>The population from Gumine in south Simbu province also has a variable phenotype, the hindwing base colour ranging from white to coffee brown. The darkest of these resemble ssp. nov. parsonsi but are distinguished on the hindwing underside by the grey colour and elongated shape of the hindwing basal streak, more sharply defined median patch and broader white vein markings. The female has not been recorded and the identity of the population remains uncertain. Though less than 40km from Mt Wilhelm, Gumine is situated at the eastern extreme of the Kubor Range that extends westward to the Mt Hagen massif, and may therefore be a hybrid zone.</p></div>	https://treatment.plazi.org/id/0389BB02A52D7250FF0A0F30532BEB67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0B255024E924.text	0389BB02A52B7257FF0A0B255024E924.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias gilliardi Sanford & Bennett 1955	<div><p>Delias gilliardi Sanford &amp; Bennett, 1955</p><p>(Figs 23-24)</p><p>This species is commonly found in sympatry with D. hallstromi at elevations between 2,500 – 3,000 meters in central PNG. The female has rarely been collected and is not illustrated by Parsons (1998) or Yagishita (1993). It is distinguished from the male by a paler yellow ground colour and more diffused black margins on the upperside wings, and by orange coloration on the underside of the abdomen.</p></div>	https://treatment.plazi.org/id/0389BB02A52B7257FF0A0B255024E924	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0CD851E3EB54.text	0389BB02A52B7257FF0A0CD851E3EB54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias hagenensis Morinaka, Van Mastrigt & Sibatani 1993	<div><p>Delias hagenensis Morinaka, Van Mastrigt &amp; Sibatani, 1993</p><p>(Figs 17-18)</p><p>Delias hagenensis was described from material collected at Pap Creek, Walya, 3000m in August 1973 and has since been recorded from Mt Hagen, Laiagam and Porgera (Parsons, 1998) and Tari (Lachlan, 2000). On Mt Hagen it is a common species however until now the female has not been recorded or illustrated in a publication.</p><p>Description of female: Broadly similar to male but distinguished on the upperside forewing by the suffusion of grey scales in the basal area and beyond the discocellular vein, (uniformly black in the male), and more prominent white subapical spots. The basal area of the hindwing has a slightly blue tinge in fresh specimens and contains yellow streaks at the costal and anal border. On the underside, the forewing discocellular bar is broader than in males and the white subapical spots are larger, particularly in space M 1 – M 2.</p></div>	https://treatment.plazi.org/id/0389BB02A52B7257FF0A0CD851E3EB54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7257FF0A0A3D51E9EA33.text	0389BB02A52B7257FF0A0A3D51E9EA33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias hallstromi Sanford & Bennett 1955	<div><p>Delias hallstromi Sanford &amp; Bennett, 1955</p><p>(Figs 19-22)</p><p>Delias hallstromi is a common species in central PNG however the female is rarely encountered. Parsons (1998) illustrates a specimen in the ANIC from Marifunga in the Eastern Highlands. On Mt Hagen, females were frequently seen at locations 1 and 2 feeding on flowering plants. Individual variation is seen in the width of the discocellular bar on the forewing underside and the extent of white markings on the forewing upperside.</p></div>	https://treatment.plazi.org/id/0389BB02A52B7257FF0A0A3D51E9EA33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52B7256FF0A08CD566AED15.text	0389BB02A52B7256FF0A08CD566AED15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias mira subsp. roepkei Sanford & Bennett 1955	<div><p>Delias mira roepkei Sanford &amp; Bennett, 1955 stat. rev.</p><p>(Figs 25-28)</p><p>The type locality of Delias mira roepkei Sanford &amp; Bennett, 1955 is Mt Hagen. The taxon was raised to species level by Van Mastrigt (2000) who mentioned the presence of two forms in the Hagen population; one with broad black borders on the upperside, the other with narrower borders and paler yellow hindwing patch on the underside that he speculated might represent a separate species. Males collected at the Murmur Pass locations exhibit a continuum of</p><p>individual variations in the upperside border depth and underside hindwing markings and are not readily dividable into two separate groups.</p><p>Specimens from Mt Wilhelm (Simbu Province) and the Eastern Highlands are also variable and, on average, have a more peach-colored anal patch on the hindwing underside.</p><p>Van Mastrigt’s decision to elevate roepkei to species level was based by the supposed sympatry of both roepkei and mira in parts of the Owen Stanley Mountains. Pending genetic analysis, the original taxonomy is adopted in this paper.</p><p>Among a number of typically white coloured females, two individuals with yellow-colored uppersides have been recorded. (Figs 28-29). Similar yellow female forms have been recorded in a number of species of the D. iltis ̡ D. aroae and D. eichorni groups.</p></div>	https://treatment.plazi.org/id/0389BB02A52B7256FF0A08CD566AED15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7256FF0A0CC2532AEB58.text	0389BB02A52A7256FF0A0CC2532AEB58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias hapalina Jordan 1930	<div><p>Delias hapalina Jordan, 1930</p><p>(Figs 29-30)</p><p>A new record for the Mt Hagen area. The species is widespread though less common in PNG than in Indonesian Papua and has previously been recorded from Mt Wilhelm and Tari ( ssp. kerowagiensis Yagishita, 1993), the Owen Stanley Mts ( ssp. owenstanleyi Schröder &amp; Treadaway, 1994) the west Bismark Mts and Porgera in Enga province (CD).</p><p>Male specimens from Mt Hagen are distinguishable from topotypical kerowagiensis by the narrower black median bands on the hindwing underside and from the Bismark and Porgera populations which lack red median bands on the hindwing. The identity and distribution of potential subspecies of D. hapalina in PNG will be reviewed after comparative study of these populations.</p></div>	https://treatment.plazi.org/id/0389BB02A52A7256FF0A0CC2532AEB58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A52A7255FF0A0A0E577EEEF1.text	0389BB02A52A7255FF0A0A0E577EEEF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias leucias subsp. leucias Jordan 1912	<div><p>Delias leucias leucias Jordan, 1912</p><p>(Figs 9-16)</p><p>The subspecific identity of D. leucias from the central mountains of Papua New Guinea has a confusing history, having been assigned to the nominate race (type locality: Mt Goliath) by D’Abrera (1978), and subsequently to subspecies huonensis by later authors such as Parsons (1998), Yagishita et al. (1993) and Van Mastrigt (2003). Lachlan (2000) reports collection of Delias leucias leucias at Ok Tedi, Western Province and D. leucias huonensis at Tari, Southern Highlands. Gotts &amp; Ginn (2004) described a new subspecies, D. leucias torini̡ from the Snow Mts, west of the nominate range, and discussed variation in D. leucias across New Guinea, mentioning the differences between the Kerowagi (Mt Wilhelm) population and the nominate subspecies from the Star Mountains. They considered the taxon D. weiskei sayuriae Okano, 1989, recorded only from Mt Wilhelm, to be a variation of leucias rather than a subspecies of D. weiskei Ribbe, 1900 .</p><p>D. leucias huonensis Talbot, 1928 was described from a single male collected by Keysser in the isolated Rawlinson Mountains on the Huon peninsula in north east New Guinea. The race remains rare in collections and apparently in nature; William Brandt did not record the species during his expedition to the Rawlinson Mts in 1968.</p><p>Specimens from Mt Hagen are similar to those from Mt Wilhelm &amp; the Eastern Highlands, both can be distinguished in the majority of individuals from the nominate subspecies - by the complete separation of the pale hindwing patch from the white costal border by the red and black subapical bands, and a more orange coloration of the forewing underside - and from ssp</p><p>huonensis by the narrower black margins on the both surfaces and the more elongated hind wing patch. Although these characters are consistent in the majority of specimens from central PNG, populations of the nominate race from various locations within Indonesian Papua show clinal development of the same features on a west - east axis. Delias leucias from central PNG is considered to represent a recognizable geographic form of the nominate subspecies.</p></div>	https://treatment.plazi.org/id/0389BB02A52A7255FF0A0A0E577EEEF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A5297255FF0A0FE653CCECA0.text	0389BB02A5297255FF0A0FE653CCECA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias callista Jordan 1930	<div><p>Delias callista Jordan, 1930</p><p>(Figs 31-32)</p><p>The type locality of D. callista porquaiensis Yagishita, 2003 is stated to be ’Porquia, PNG’ however no such place name is gazetted in PNG and it is assumed that it is a misspelling of Porgera, the site of a large gold mine in western Enga Province, approximately 80km west of Mt Hagen. Male specimens from Porgera and the Ok Tedi area agree with the original description of porquaiensis and are characterized by rich yellow coloration, broad black submarginal bands and triangular shaped black terminal vein markings on the hindwing underside.</p><p>Males of D. callista from Mt Hagen consistently differ from this phenotype by having a paler shade of yellow on the hindwing underside, narrow subterminal bands and terminal vein markings. A similar form has been recorded from Gumine (IFTA). Females from Mt Hagen have a series of 5 white apical spots on the forewing upperside and 4 – 5 white subterminal spots on the hindwing that are absent or poorly developed in more western areas. The taxonomic status of D. callista populations in PNG and western Papua is uncertain and will be reviewed in a further publication.</p></div>	https://treatment.plazi.org/id/0389BB02A5297255FF0A0FE653CCECA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
0389BB02A5297255FF0A0A5150CAEB88.text	0389BB02A5297255FF0A0A5150CAEB88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Delias carstenziana subsp. inexpecta Morinaka & Nakazawa 1999	<div><p>Delias carstenziana inexpecta Morinaka &amp; Nakazawa, 1999</p><p>The presence of this species in close proximity to, but at lower altitude than, its sister species D. gilliardi ̡ is of interest. A single male was recorded by a local collector at Aiyamp creek, on the lower western slopes of the Mt Hagen massif. The holotype and other specimens were collected by Phil Saywer in July 1975 at ‘Poketamanda’ = Mt Nose, near Wabag, Enga Province, approximately 30km west of Mt Hagen.</p></div>	https://treatment.plazi.org/id/0389BB02A5297255FF0A0A5150CAEB88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Davenport, Chris;Grimaldi, Nicolas	Davenport, Chris, Grimaldi, Nicolas (2019): A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae). Sugapa Digital 11 (2): 53-67, DOI: 10.19269/sugapa2019.11(2).01
