identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0389B43E1178540AFF6CFC48FC378122.text	0389B43E1178540AFF6CFC48FC378122.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heleobia Stimpson 1865	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Heleobia Stimpson, 1865</p>
            <p> Type species:  Paludestrina culminea d’Orbigny, 1840 by original designation. </p>
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	https://treatment.plazi.org/id/0389B43E1178540AFF6CFC48FC378122	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Collado, Gonzalo A.;Valladares, Moisés A.;Méndez, Marco A.	Collado, Gonzalo A., Valladares, Moisés A., Méndez, Marco A. (2016): A new species of Heleobia (Caenogastropoda: Cochliopidae) from the Chilean Altiplano. Zootaxa 4137 (2): 277-280, DOI: 10.11646/zootaxa.4137.2.8
0389B43E11785409FF6CFB31FB1C8428.text	0389B43E11785409FF6CFB31FB1C8428.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heleobia carcotensis	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heleobia carcotensis sp. nov.</p>
            <p>Figure 2</p>
            <p>Holotype. (MZUC – UCCC 43802, Fig. 2A). Collected by G.A. Collado from Spring 1 in the Carcote saltpan, Chile (28 November 2011). Shell measurements SL 4.4 mm, SW 2.0 mm, AL 1.7 mm, AW 1.1 mm.</p>
            <p>Paratypes. (MZUC – UCCC 43803–43807, Fig. 2B–F). Snails from Spring 1 in the Carcote saltpan, Chile, collected with the holotype by G.A. Collado. Shell measurements: (n= 5): SL: 4.52 ± 0.35 (4.20–4.90); SW: 1.98 ± 0.11 (1.90– 2.10); AL: 1.80 ± 0.20 (1.60–2.10); AW: 1.16 ± 0.15 (0.99–1.30); SW/SL: 0.44 ± 0.01 (0.43–0.45); AL/SL: 0.40 ± 0.01 (0.38–0.42); AW/AL: 0.64 ± 0.05 (0.59–0.70).</p>
            <p>Etymology. Specific name refers to the Carcote saltpan.</p>
            <p>Description. Shell elongate conic, periostracum light brown, umbilicus absent. Teleoconch with six shell whorls and fine axial striae. Aperture ovate, slightly triangular or strongly angled adapically; parietal margin of the lip slightly thickened, outer lip thin. Protoconch (Fig. 2G) slightly differentiated from teleoconch.</p>
            <p>Operculum paucispiral, thin, ovate, grayish around a central light brown region, nucleus eccentric (Fig. 2H–I). Radula shown in Fig. 2 J–O. Median cusps of central radular teeth elongate, distally pointed, lateral cusps five-six (n= 20, 10 teeth from voucher V1–2, five from V1–8, five from V1–8), one basal cusp (n= 30, 10 teeth from each of the vouchers), basal tongue of central radular teeth V-shaped, lateral teeth with two-three cusps (n= 18, 10 teeth from voucher V1–2, six from V1–8, two from V1–10) on inner and three-five cusps (n= 30, 10 teeth from voucher V1–2, 10 from V1–5, 10 from V1–8) on outer side, central cusp larger and pointed. Inner marginal teeth having 19–22 cusps (n= 10, six from voucher V1–2, four from V1–8), outer marginal teeth with 29–42 cusps (n= 3, voucher V1–2).</p>
            <p>Foot gray, propodium white, head brown-black with less pigment centrally, snout black, distal lips white. Tentacles black with a brow, central axial band and a gray horizontal band near the base (Fig. 2P–R). Penis gray with a row of three - five apocrine glands (n= 3) along the outer edge (Fig. 2S–U); distal portion with a terminal papilla.</p>
            <p>Distribution and habitat. Spring 1 in the Carcote saltpan (3688 m above sea level). This saltpan is a closed basin in the Chilean Altiplano. Spring 1 is a thermal water body (21°C) that forms a rectangular pool that discharges to the central zone of the saltpan (Fig. 1). Snails were found on rocks and aquatic vegetation.</p>
            <p> Remarks. The morphology of the penis of this new species conforms to  Heleobia as currently diagnosed (Hershler &amp; Thompson 1992). The shell morphology differs from regional congeners in the form of the aperture (strongly angled adapically) and in the general ground plan of the penis [see Hubendick (1955), Collado et al. (2011a), Collado et al. (2013) and Collado (2015) for comparisons].  Heleobia carcotensis appears to be closely related to the  Heleobia snails from Colpa, Parinacota and Isluga in the Chilean Altiplano and  H. opachensis from the Atacama Desert. </p>
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	https://treatment.plazi.org/id/0389B43E11785409FF6CFB31FB1C8428	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Collado, Gonzalo A.;Valladares, Moisés A.;Méndez, Marco A.	Collado, Gonzalo A., Valladares, Moisés A., Méndez, Marco A. (2016): A new species of Heleobia (Caenogastropoda: Cochliopidae) from the Chilean Altiplano. Zootaxa 4137 (2): 277-280, DOI: 10.11646/zootaxa.4137.2.8
