taxonID	type	description	language	source
0389935EFB7FFFE5FF04A720F888FA06.taxon	materials_examined	Type materials. Holotype (UMUT-RM 32483), Usujiri, Hakodate, Hokkaido, Japan, 10.8 x 8.2 x 3.8 mm. Selected paratypes: Paratype 1 (NSMT-Mo 78956); Paratype 2 (NSMT-Mo 78957); Paratype 3 (UMUT-RM 32492); Paratype 4 (UMUT-RM 32471), Paratype 5 (UMUT-RM 32496); Paratype 6 (UMUT-RM 32494); Paratype 7 (UMUT- RM 32479). Remaining paratypes are also deposited at UMUT (UMUT RM 3232467 - 32505).	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	etymology	Etymology. This species is dedicated to Prof. Seiji Goshima, Faculty of Fisheries Sciences, Hokkaido University, Japan, who has greatly contributed to biological studies of the new species (Niu & Fuji 1989; Niu et al. 1992, 1994 a, b, 1997, 1998). Shell (Figures 2, 3). Shell moderately thick, aperture oval. Apex antero-dorsally directed and positioned at anterior fourth. Anterior and ventral slopes nearly straight, posterior slope convex. Exterior surface sculptured by crowded, rough, regularly arranged radial riblets. Exterior colour light brown with 12 brown radial rays. Interior margin alternating light brown and dark brown; intermediate area white; central area except apex stained dark chocolate brown; apex area white.	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	description	Animal. Dorsal part of mantle margin fringed with light brown. Head and visceral mass dark beige. Ventral surface creamy white. Radula (Figure 4). First lateral teeth relatively short with pointed cusp. Second lateral teeth long with slightly rounded cusp. Third lateral teeth broad with beak-like cusps. Marginal teeth small, rounded.	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	materials_examined	Type locality. Usujiri, Hakodate, Hokkaido, Japan (41 ° 56 ' 11 " N, 140 ° 57 ' 00 " E).	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	distribution	Distribution. Northern part of Japan from Erimo, Hokkaido to Karakuwa, Miyagi on the Pacific coast and Monbetsu, Hokkaido to Tsuruoka, Yamagata, Vostok Bay, Russia in the Japan Sea. Habitat. This species lives in the middle to lower intertidal zone, and is often seen among the small barnacle Chthamalus challengeri and on the purplish mussel Septifer virgatus.	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	discussion	Remarks. Although the new species is morphologically similar to L. kogamogai, it is distinguished by having rough, fine and regularly-arranged radial riblets (Fig. 3). The radulae differ in terms of the morphology of the lateral teeth. Lottia goshimai n. sp. has short first lateral teeth, a long and slightly rounded cusp on the second lateral teeth, and slightly beak-like third lateral teeth (Figs. 4 A – D). Lottia kogamogai has long and rounded first lateral teeth, a broad and pointed cusp on the second lateral teeth, and short, broad third lateral teeth (Figs. 4 E – H). Intraspecific variation was observed within L. goshimai n. sp. and L. kogamogai in shell and radular morphogy. The individuals living on the shell of Septifer virgatus have taller, darker shells and shorter second lateral teeth than those from rocks (Figs. 2, 4). The reproductive cycle of the species (as Collisella heroldi) was examined by Niu & Fuji (1989). According to this study, the reproductive season is August to October, and size at maturity is about 5 mm. The Lottia kogamogai complex is clearly divided according to geographical distribution of each species: Lottia goshimai n. sp., L. kogamogai and L. luchuana are limited to northern Japan, southern Japan and tropical Japan respectively (Fig. 5). Although L. tenuisculpta and L. lindbergi have a wide distribution along temperate Japan (Sasaki & Okutani 1994 b), these two species were not analyzed intensively in this paper. Specie locality ID in Fig. 1 GenBank Voucher Lottia kogamogai Sa aki & Okutani, 1994 Banđa, Tateyama, Chiba Pref., Japan (type locality) * K 320 Y LC 137940 UMUT-RM 32416 Banđa, Tateyama, Chiba Pref., Japan (type locality) * K 321 Y LC 137941 UMUT-RM 32417 Banđa, Tateyama, Chiba Pref., Japan (type locality) * K 322 Y LC 137942 UMUT-RM 32418 Cape Unoko, Kitaibaraki, Ibaraki Pref., Japan K 583 S LC 138001 UMUT-RM 32419 Cape Ooarai, Ooarai, Ibaraki Pref., Japan K 571 U LC 137997 UMUT-RM 32420 Cape Ooarai, Ooarai, Ibaraki Pref., Japan * K 572 U LC 137998 UMUT-RM 32421 Cape Inubou, Cho hi, Chiba Pref., Japan K 578 V LC 138000 UMUT-RM 32422 Cape Yahata, Kat uura, Chiba Pref., Japan K 489 W LC 137991 UMUT-RM 32423 Cape Yahata, Kat uura, Chiba Pref., Japan K 490 W LC 137992 UMUT-RM 32424 Cape Yahata, Kat uura, Chiba Pref., Japan K 566 W LC 137996 UMUT-RM 32425 Joga hima, Miura, Kanagawa Pref., Japan * K 336 Z LC 137945 UMUT-RM 32426 Joga hima, Miura, Kanagawa Pref., Japan * K 337 Z LC 137946 UMUT-RM 32427 Joga hima, Miura, Kanagawa Pref., Japan K 338 Z LC 137947 UMUT-RM 32428 Joga hima, Miura, Kanagawa Pref., Japan K 351 Z LC 137948 UMUT-RM 32429 Joga hima, Miura, Kanagawa Pref., Japan * K 503 Z LC 137994 UMUT-RM 32430 Mitsui hi, Manazuru, Kanagawa Pref., Japan K 316 a LC 137938 UMUT-RM 32431 Yokata Port, Toyama, Toyama Pref., Japan K 33 L LC 137916 UMUT-RM 32432 Yokata Port, Toyama, Toyama Pref., Japan K 249 L LC 137935 UMUT-RM 32433 Yokata Port, Toyama, Toyama Pref., Japan K 251 L LC 137936 UMUT-RM 32434 Akagurizaki, Ooi, Fukui Pref., Japan K 244 N LC 137933 UMUT-RM 32435 Akagurizaki, Ooi, Fukui Pref., Japan K 246 N LC 137934 UMUT-RM 32436 Morozaki, Minamichita, Aichi Pref., Japan K 29 b LC 137915 UMUT-RM 32437 Morozaki, Minamichita, Aichi Pref., Japan K 208 b LC 137930 UMUT-RM 32438 Rinkai, Shirahama, Wakayama Pref., Japan K 84 e LC 137919 UMUT-RM 32439 Ku himoto, Wakayama Pref., Japan K 3 đ LC 137909 UMUT-RM 32440 Ku himoto, Wakayama Pref., Japan K 4 đ LC 137910 UMUT-RM 32441 …… continued on the next page Specie locality ID in Fig. 1 GenBank Voucher Ku himoto, Wakayama Pref., Japan K 145 đ LC 137923 UMUT-RM 32442 Ku himoto, Wakayama Pref., Japan K 146 đ LC 137924 UMUT-RM 32443 Ku himoto, Wakayama Pref., Japan K 152 đ LC 137925 UMUT-RM 32444 Ku himoto, Wakayama Pref., Japan * K 157 đ LC 137926 UMUT-RM 32445 Urađome, Iwami, Tottori Pref., Japan * K 423 O LC 137979 UMUT-RM 32446 Cape Hane, Kochi Pref., Japan K 66 g LC 137917 UMUT-RM 32447 Go hikinohama, U a, Kochi Pref., Japan K 258 h LC 137937 UMUT-RM 32448 Omuronohama, Okit u, Kochi Pref., Japan K 28 i LC 137914 UMUT-RM 32449 Tat uku hi, Kochi Pref., Japan * K 21 j LC 137912 UMUT-RM 32450 Tat uku hi, Kochi Pref., Japan K 174 j LC 137927 UMUT-RM 32451 T uou, Nagato, Yamaguchi Pref., Japan * K 364 P LC 137949 UMUT-RM 32452 T uou, Nagato, Yamaguchi Pref., Japan K 366 P LC 137950 UMUT-RM 32453 T uou, Nagato, Yamaguchi Pref., Japan K 367 P LC 137951 UMUT-RM 32454 I ozakihana, Shingu, Fukuoka Pref., Japan * K 70 k LC 137918 UMUT-RM 32455 Ura hiro, Nobeoka, Miyazaki Pref., Japan K 334 o LC 137943 UMUT-RM 32456 Ura hiro, Nobeoka, Miyazaki Pref., Japan K 335 o LC 137944 UMUT-RM 32457 Cape Sata, Minamio umi, Kago hima Pref., Japan K 26 m LC 137913 UMUT-RM 32458 Cape Sata, Minamio umi, Kago hima Pref., Japan K 230 m LC 137931 UMUT-RM 32459 Cape Sata, Minamio umi, Kago hima Pref., Japan K 231 m LC 137932 UMUT-RM 32460 Hamakanaya, Futt u, Chiba Pref. Japan L 945 X LC 138005 UMUT-RM 32461 Kaino, Kihoku, Mie Pref., Japan L 158 c LC 138003 UMUT-RM 32462 Kaino, Kihoku, Mie Pref., Japan L 159 c LC 138004 UMUT-RM 32463 Kamogaura, Wajima, I hikawa Pref., Japan L 1147 M LC 138007 UMUT-RM 32464 Kamogaura, Wajima, I hikawa Pref., Japan L 1148 M LC 138008 UMUT-RM 32465 Kamogaura, Wajima, I hikawa Pref., Japan L 1149 M LC 138009 UMUT-RM 32466 Lottia goshimai n. sp. Oumu, Monbet u, Hokkaiđo, Japan K 434 A LC 137983 UMUT-RM 32467 Oumu, Monbet u, Hokkaiđo, Japan K 439 A LC 137984 UMUT-RM 32468 Cape Erimo, Erimo, Hokkaiđo, Japan K 522 E LC 137995 UMUT-RM 32469 …… continued on the next page Specie locality ID in Fig. 1 GenBank Voucher …… continued on the next page Specie locality ID in Fig. 1 GenBank Voucher A amu hi, Aomori Pref., Japan * K 482 I LC 137987 UMUT-RM 32497 A amu hi, Aomori Pref., Japan * K 483 I LC 137988 UMUT-RM 32498 A amu hi, Aomori Pref., Japan * K 484 I LC 137989 UMUT-RM 32499 Ot uchi, Iwate Pref., Japan * K 205 Q LC 137928 UMUT-RM 32500 Ot uchi, Iwate Pref., Japan K 206 Q LC 137929 UMUT-RM 32501 Ot uchi, Iwate Pref., Japan K 496 Q LC 137993 UMUT-RM 32502 Karakuwa, Ke ennuma, Miyagi Pref., Japan * K 318 R LC 137939 NSMT-Mo 78956 Karakuwa, Ke ennuma, Miyagi Pref., Japan K 429 R LC 137980 UMUT-RM 32503 Karakuwa, Ke ennuma, Miyagi Pref., Japan K 431 R LC 137981 UMUT-RM 32504 Imaizumi, T uruoka, Yamagata Pref., Japan K 574 J LC 137999 UMUT-RM 32505 Lottia tenuisculupta Sa aki & Okutani, 1994 Taka hima, Otaru, Hokkaiđo, Japan * K 369 B LC 137953 UMUT-RM 32506 Urađome, Iwami, Tottori Pref., Japan K 417 O LC 137978 UMUT-RM 32507 Kamekawahamađa, Beppu, Ohita Pref., Japan K 433 n LC 137982 UMUT-RM 32508	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
0389935EFB7FFFE5FF04A720F888FA06.taxon	description	Lottia lindbergi Sa aki & Okutani, 1994 Kaminokuni, Hiyama, Hokkaiđo, Japan * K 388 C LC 137965 UMUT-RM 32509 Hirai o, Hitachinaka, Ibaraki Pref., Japan L 173 T LC 137971 UMUT-RM 32510 Tat uku i, Kochi Pref., Japan K 20 j LC 137911 UMUT-RM 32511 Lottia luchuana (Pil bry, 1901) Ka ari, Amami, Kago hima Pref., Japan L 965 p LC 138006 UMUT-RM 32512 Ogimi, Okinawa, Okinawa, Japan L 131 q LC 138002 UMUT-RM 32513 Onna, Okinawa, Japan * K 486 r LC 137990 UMUT-RM 32514	en	Nakayama, Ryo, Sasaki, Takenori, Nakano, Tomoyuki (2017): Molecular analysis reveals a new cryptic species in a limpet Lottia kogamogai (Patellogastropoda: Lottiidae) from Japan. Zootaxa 4277 (2): 237-251, DOI: 10.11646/zootaxa.4277.2.4
