taxonID	type	description	language	source
038987C3BA32FFF0B3A6FBA4FED8000C.taxon	materials_examined	Type species: Metacryptoseius persicus sp. nov.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA32FFF0B3A6FBA4FED8000C.taxon	diagnosis	Diagnosis. Dorsal shield of adult females and males not covering whole body, with 26 pairs of setae and 22 pairs of pore-like structures. Adult female sternal shield fused with endopodal shield between coxae I / II and II / III, with three pairs of setae and two pairs of lyrifissures; minute metasternal shields bearing sternal setae st 4 and lyrifissures iv 3 present. Adult male with sternogenital shield slightly narrowed posteriorly and not fused with anal shield, with five pairs of setae and three pairs of lyrifissures. Chaetotaxy of opisthogaster reduced to five pairs of setae (JV 1, JV 2, JV 5, ZV 2 and ZV 3). Gnathotectum of both sexes triangular with anterior margin smooth to slightly dentate. Deutosternal groove with six rows of denticles. Sexual dimorphism visible in the shape of corniculi and chelicerae: corniculi stout, in male rounded apically and in female acute; fixed cheliceral digit longer than movable digit in females.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA32FFF0B3A6FBA4FED8000C.taxon	description	Description. Female. Dorsal idiosoma. Dorsal shield with anterolateral incisions reaching level of setae z 2 to j 3. Dorsum with 32 pairs of setae: setae j 1 - j 6, J 1, J 3 - J 5, z 1, z 2, z 4, z 5, Z 2 - Z 5, s 2, s 4, s 5, S 3 - S 5, r 2 and r 3 on dorsal shield (s 2, r 2 and r 3 on laterodosal elements); setae s 6, r 5, R 1, R 3 - R 5 on soft cuticle. Dorsal shield with 22 pairs of discernible pore-like structures (eight podonotal, 14 opisthonotal), of which seven (three podonotal, four opisthonotal) superficially appear secretory (gland pores) and 15 (five podonotal, ten opisthonotal) non-secretory (lyrifissures); marginal poroid Rp (= idR 3) between setae R 3 - R 4 and two pairs of lyrifissures on soft cuticle. Ve n t r a l idiosoma. Tritosternum with narrow base and two free pilose laciniae; presternal region weakly sclerotized; sternal shield entire, with three pairs of setae and two pairs of lyrifissures (iv 1, iv 2), fused with endopodal shields between coxae I / II and II / III; fourth pair of sternal setae and lyrifissures on minute metasternal plates (Figs 10, 11). Endopodal shields well developed between coxae III and IV; parapodal shields present. Genital shield with hyaline, smooth convex anterior margin; posterior margin of genital shield broadly rounded; setae st 5 on lateral borders of shield, paragenital lyrifissures iv 5 on soft cuticle alongside genital shield. Inguinal region with a pair of nearly oval metapodal platelets and gland pores gv 2 posterior to coxae IV, sometimes with platelets behind posterior margin of genital shield. Anal shield relatively small, pear-shaped or nearly subtriangular; anus located on anterior half of shield, with relatively small anal opening (euanal setae and lyrifissures on anal valves absent); adanal gland pores gv 3 on lateral edges of shield; cribrum normally developed. Chaetotaxy of opisthogaster reduced to five pairs of simple setae (larval setae JV 1, JV 2, JV 5, ZV 2 and deutonymphal setae ZV 3) on soft cuticle around anal shield, and four pairs of lyrifissures. Peritrematal plates with one gland pore gp 1 and 2 lyrifissures ip (Fig. 10), lacking any posterior extension, poststigmatic gland pore on soft cuticle. Peritremes well developed, wide, extending from stigmata between coxae III and IV to anterior border of coxae II or between coxae I and II. Spermatheca with distinct connecting ducts and narrow calyx, solenostome on base of leg III. Gnathosoma (Fig. 7). Gnathotectum triangular, with denticulate or smooth anterior margin (Fig. 8); hypostome with three pairs of smooth setae; deutosternal groove with six moderately narrow rows of denticles, all rows connected; gnathobase with a pair of simple capitular setae. Fixed cheliceral digit multidentate, pilus dentilis short and setiform; dorsal seta small, smooth and near dorso-proximal lyrifissure; with large transversal antiaxial lyrifissure; movable digit with fringed hyaline process at its base. Corniculi stout and acute at tip; internal malae slender, pointed, finely fringed, extending beyond anterior border of palp femur; salivary stylets thin and rounded apically; labrum prominent, blade-like, reaching to palp tibia, with pilose surface. Palpi with normal setation and setal ontogeny as described for Gamasina by Evans (1964), including only one seta on palptrochanter of protonymph; palp genu with six setae in adults; palpal setae simple; palptarsal apotele two-tined. Legs. Legs I-IV of moderate length, with well developed paired claws and rounded pulvilli inserted on well developed pretarsi. Ventral face of coxae I with two coxal gland pores. Chaetotaxy: coxae 2 - 2 - 2 - 1; trochanters 6 - 5 - 5 - 5; femora (2 3 / 1 2 / 3 2) – (2 3 / 1 2 / 2 1) – (1 2 / 1 1 / 1 1) – (1 2 / 1 1 / 1 1); genua (1 3 / 1 2 / 2 2) – (2 3 / 1 2 / 1 2) – (1 2 / 1 2 / 1 1) – (1 2 / 1 2 / 1 1) and tibiae (1 3 / 1 2 / 2 2) – (2 2 / 1 2 / 1 2) – (1 1 / 1 2 / 1 1) – (1 1 / 1 2 / 1 1). Some ventral setae on femur and genu elongated. Seta av - 1 on tarsi II – IV enlarged and thick, tri-dentate at the tip (as in Fig. 29 a). Male. Dorsal idiosoma. Dorsal chaetotaxy, adenotaxy, poroidotaxy and shape as for female. Ve n t r a l idiosoma. Tritosternal base moderately narrow and shorter than that of female, laciniae free and pilose. Genital opening arising on anterior margin of sternogenital shield. Sternogenital shield fused with endopodal shields between coxae, with five pairs of setae and three pairs of lyrifissures, the anterior lyrifissure slit-like, second and third oval-shaped; margins heavily sclerotized and frame-like, especially between coxae III – IV; paragenital lyrifissures iv 5 on soft cuticle beside posterolateral corners of shield (Fig. 21). Five pairs of opisthogastric setae, anal shield and cribrum similar to those of female. Parapodal, metapodal and peritrematal shields, ventral platelets and peritremes similar to those of female. Gnathosoma. Gnathotectum similar to female; fixed cheliceral digit with only one tooth and a short setiform pilus dentilis; movable digit with two teeth; hyaline process apparently absent; spermatodactyl truncate distally (Figs 13, 24). Corniculi thumb-like, with rounded tip (Fig. 23). Other characters similar to the female. Legs. Pretarsi and chaetotaxy similar to those of female. Legs of moderate length, shorter than those of female. Femur and tarsus (Fig. 18) of leg II with one modified spur-like ventral setae; other leg setae (apart from av - 1 on tarsi II – IV, thickened, bi- or tri-dentate) simple, but many dorsal and lateral setae of femur, genu and tibia thickened. Deutonymph. Dorsal idiosoma. Dorsal shield without lateral incisions, covering the entire dorsum, usually with 23 pairs of setae. Setae s 6, R 1, R 3 - R 5, r 2, r 3, r 5, s 6 on soft cuticle. Dorsal idiosomal complement of lyrifissures and gland-pores similar to those of female. Ve n t r a l idiosoma. Sternal shield with four pairs of setae and three pairs of lyrifissures; setae st 5 and paragenital lyrifissures on soft cuticle at level of posterior margin of shield; endopodal shields weekly developed or absent (Fig. 5). Opisthogaster with five pairs of ventral setae (JV 1, JV 2, JV 5, ZV 2 and ZV 3). Peritrematal shields anteriorly free, not fused with dorsal shield, one lyrifissure (ip 1) on antero-dorsal edge of peritrematal shield; ip 2 and gland pore gp 1 between coxae II and III on soft cuticle; poststigmatic gland pores gp 2 on soft cuticle, exopodal plates behind coxae IV well delineated, with gland pores gv 2 posterior to them. Anal shield, glands gv 3 and cribrum similar to adult female. Gnathosoma. Gnathotectum, salivary stylets, corniculi, deutosternal structures and palpi similar to those of female. Movable chela bi-dentate and longer than fixed chela with small teeth near the tip. Legs. Pretarsi, claws, chaetotaxy of legs I-IV and modified av - 1 on legs II-IV similar to those of female. Protonymph. Dorsal idiosoma. Sclerotized podonotal and opisthonotal shields covering most of dorsal surface, and 27 pairs of setae; 11 pairs on podonotal shield, five pairs (r 2, r 3, r 5, s 6, S 5 and R 1) on lateral integument and nine on opisthonotal shield. Body dorsum with 18 pairs of pore-like structures, of which 11 (two on podonotal shield, one on soft cuticle, eight on pygidial shield) non-secretory (lyrifissures) and six (three on podonotal shield, three on opisthonotal shield) appear to be secretory (gland pores). Ve n t r a l idiosoma. Sternal shield poorly sclerotized with three pairs of setae; setae st 5 inserted on soft cuticle at level of coxae IV. Glands gv 2 present. Anal shield, adanal glands, and cribrum similar in those of deutonymph. Opisthogaster with four pairs of larval setae on soft cuticle around anal shield. Peritrematal shield as long as peritremes, short; peritrematal gland gp 1 present between insertion of dorsal setae r 3 and r 5. Gnathosoma. Gnathotectum triangular, anterolaterally serrate; cheliceral fixed digit bi-dentate, movable digit reduced and edentate; other gnathosomal structures similar to those of deutonymph, except palpi with normal protonymphal complement of setae (see Evans 1964), including only one seta on trochanter. Legs. Legs I-IV with well-developed pretarsi and claws; protonymphal complement of coxal setae: 2 - 2 - 2 - 1; trochanters: 4 - 4 - 4 - 4; femur: (2 2 / 1 2 / 1 2) – (1 2 / 1 2 / 1 1) – (0 2 / 1 1 / 1 0) – (1 2 / 0 1 / 0 0); genu: (1 2 / 1 2 / 1 1) – (1 2 / 0 2 / 0 1) – (1 2 / 0 2 / 0 1) – (1 2 / 0 1 / 0 0); tibiae: (1 2 / 1 2 / 1 1) – (1 1 / 1 2 / 1 1) – (1 1 / 1 2 / 1 1) – (1 1 / 1 2 / 1 1). Seta av - 1 on tarsi II to IV modified as in deutonymph and adults. Notes. Metacryptoseius differs from other genera of the subfamily with six setae on palp genu, tibia I with three ventral setae, genu III with two ventral setae, femur I with four ventral setae, J 1 present and peritrematal shield reduced, in having dorsal setae J 3 and Z 5, metasternal shields bearing setae st 4 and iv 3 and deutosternal groove with six rows of denticles. The related genus Cryptoseius lacks metasternal shields, iv 3 is on sternal shield, setae J 3 and Z 5 absent and has eight rows of denticles in the deutosternal groove. Scamaphis Karg 1979, with 27 pairs of setae on dorsal shield (z 6 present, r 5 on the shield), lacks setae J 1 and J 2 and laterodorsal elements, has a deutosternum with five rows of denticles, and the shape of gnathotectum is different.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA32FFF0B3A6FBA4FED8000C.taxon	etymology	Etymology. The genus name Metacryptoseius refers to the similarity of this genus to the related genus Cryptoseius.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA30FFFBB3A6F9D1FF010334.taxon	diagnosis	Diagnosis. Dorsal shield elongate-oval with irregular lateral and posterior margins, slightly narrowed posteriorly; dorsal setal length of adults and nymphs slightly variable; setae JV 5 nearly five times longer than J 5 and Z 5 in female; s 2 approximately equal in length with z 2 and slightly longer than j 3; j 2 three times longer than z 1; dorsal characters of adult male like those of female, except JV 5, that is shorter, nearly twice longer than J 5 and other setae subequal in length. Sternal setae st 2 shorter than distance to st 3. Peritreme terminates between coxae I and II or anterior to margin of coxae II. Anal shield pear-shaped.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA30FFFBB3A6F9D1FF010334.taxon	description	Description. Female. Idiosoma. Length: 620 – 753, width: 483 – 516 (n = 5). Dorsal idiosoma (Fig. 6). Dorsal shield 644 – 647 long, 390 – 411 wide, with striations antero-laterally, elongate-oval, with irregular margins and 26 pairs of simple and smooth setae: setae z 2 (45) and s 2 (41) subequal in length; j 2 (53) approximately three times longer than z 1 (20); j 2> j 3 (28 – 34 long); opisthonotal central setae are shortest (16 – 18 long); longest dorsal setae not more than three times longer than J 5 (16 long); setae on soft cuticle, r 5 (ca 42), s 6 (47), R 1 (29), R 3 (42), R 4 (38), R 5 (42 – 50). Anterolateral incisions of dorsal shield reaching to level of setae j 3. Dorsal poroidotaxy and adenotaxy typical of genus. Ventral idiosoma (Fig. 10). Tritosternal base 33 long, laciniae 90 long; sternal shield granulated and reticulate-striate anteriorly (174 long, 130 wide at level of setae st 2), with posterior margin almost straight, bearing setae st 1 (41), st 2 (52), st 3 (51) and two pairs of lyrifissures, first pair oriented diagonally, but not always stable; st 4 (55) and third pair of sternal lyrifissures on minute metasternal plates (Fig. 11). Genital shield (160 long, 101 wide at level of genital setae) as defined in genus, with convex posterior margin; st 5 58 long. Opisthogaster with five pairs of simple setae: JV 1 (48), JV 2 (46), JV 5 (70), ZV 2 (36) and ZV 3 (30). Peritremes 257 long. Anal shield narrow, pear-shaped, with rounded anterior margin (132 long, 101 wide); para-anal setae 32 long, postanal 49 long. Gnathosoma. Gnathotectum triangular, with slightly serrate margin (Fig. 8); subcapitulum typical for genus (Fig. 7). Hypostomal (h 1, h 2, h 3) and capitular setae simple and smooth, 23, 39, 20 and 20 long respectively. Corniculi wide and stout, 36 – 38 long. Chelicerae 174 long and 37 wide, with movable digit (63 long from tip to base) longer than fixed digit (44 long from tip to the dorsal lyrifissure) (Fig. 9). Legs. Leg chaetotaxy typical for genus; av - 1 (18 long) thickened and three dentate apically; legs I – IV lengths, 461, 404, 413 and 532 respectively. Male. Idiosoma. Length: 609 – 639, width: 425 – 446 (n = 3). Dorsal idiosoma (Fig. 12). Dorsal shield 593 – 620 long, 360 – 382 wide. Dorsal characters similar to female, but lengths of setae vary, setae S 4 are off shield; podonotal setae 23 – 32 long (except z 1 = 12, j 1 = 19, z 2 = 46> s 2 = 32); opisthonotal setae 14 – 27 long; J 5 14 long. Ventral idiosoma. Tritosternal base shorter than that of female (19 long), laciniae 94 long; sternogenital shield (303 long, 120 wide at level of st 2 and 70 at level of st 5) with anterior half reticulate, remainder evenly granulate, except area between setae st 5 which is finely punctate, with setae st 1, st 2, st 3, st 4, st 5 thickened (38, 37, 39, 34, 37 long respectively) and three pairs of lyrifissures; genital opening 31 wide. Peritremes longer than those of female (270 long). Ventral adenotaxy and poroidotaxy as in female; JV 5 (36 – 39 long) shorter than those of female, almost twice J 5; ventral setae JV 1 (30) and JV 2 (33) longer than ZV 2 (24) and ZV 3 (13 long); anal shield pear-shaped, 130 long and 111 wide. Gnathosoma. Curniculi with rounded tips (Fig. 14); chelicera 157 long, 40 width; fixed chela 29 long, movable chela 52 long; spermatodactyl 18 long (Fig. 13). Legs. Chaetotaxy typical of genus. Tarsal seta av - 1 (10 long) approximately conical, bifid; tarsal seta pv - 1 with basal third broad and rest very slender (Fig. 15). Length of legs I-IV, 385, 385, 400 and 470 respectively. Deutonymph. Idiosoma. Length: 471 – 523, width: 323 – 349 (n = 4). Dorsal idiosoma (Fig. 4). Dorsal shield (465 – 483 length, 259 – 271 width) finely punctate, with 24 pairs of setae; setae j 1 - j 6 18 - 25 long; setae z 1 11 long, z 2 16 - 22, z 4, z 5, s 2 and s 4 16 and s 5 19 long; soft cuticle with setae r 2 = r 3 = r 5 = s 6 (26 long) in podonotal region and setae R 1 (10 - 19), R 3 - R 5 (16) and sometimes S 4 (16) in opisthonotal region; setae on opisthonotal region 12 - 13 long (setae J and Z) and 20 long (setae S). Ventral idiosoma (Fig. 5). Tritosternal base ca 39 long, laciniae ca 65 long; sternal shield with four pairs of setae (st 1 = st 2 = st 3, 35 - 36 long and st 4 25 long), three pairs of lyrifissures; setae st 5 (22 long) and paragenital lyrifissures iv 5 inserted on soft cuticle at level of posterior margin of the shield. Endopodal shields present between coxae II-III and coxae III-IV present. Opisthogaster with five pairs of ventral setae JV 1 (29), JV 2 (22), JV 5 (28), ZV 2 (18) and ZV 3 (16). Peritreme extending to posterior margin of coxae I, 247 long; peritrematal shield originating opposite middle of coxae I, with ip 1 on anterior edge of peritrematal shield and glands gp 1 situated mediad to peritreme, between coxae II and III; poststigmatic glands gp 2 on soft cuticle. Anal shield subtriangular (81 long, 101 wide). Gnathosoma. Gnathotectum, subcapitulum and deutosternum typical of genus; corniculi with pointed tips and wide bases; movable digit of chelicerae 45 long, fixed digit (from dorsal lyrifissure) 32 long. Legs. Pretarsi, claws, and chaetotaxy of legs I-IV as those of adult female. Length of legs I-IV: 361, 318, 333 and 376 respectively. Seta av - 1 (14 long) on tarsi II-IV modified like in female. Protonymph. Idiosoma. Length: 510, width: 293. Dorsal idiosoma (Fig. 1). Dorsum with sclerotized podonotal (294 long, 258 wide) and opisthonotal (161 long, 242 wide) shields with punctate cuticle. Central opisthonotal setae ca 10 long; other dorsal setae longer (14 – 19 long). Ventral idiosoma (Fig. 2). Tritosternum with smooth base (46 long) and free pilose laciniae (61 long). Sternal shield with indistinct outline; setae st 1, st 2, st 3 smooth, 20 – 22 long; st 5 14 long; ventral setae with larval complement of smooth setae JV 1, JV 2, JV 5 and ZV 2 (14 – 20 long); pores gv 2 close to posterior margin of coxa IV. Anal shield (74 long, 56 wide) pear-shaped, with rounded anterior margins; circum-anal setae as other ventral setae. Peritreme (43 long) and peritrematal shield short and extending to posterior margin of coxae III or anterior margin of coxae IV. Gnathosoma. Gnathotectum triangular, serrate (Fig. 3); length of chelicera 93, width 32; fixed chela 24 long; movable chela 35 long. Legs. Leg chaetotaxy typical of genus. Seta av - 1 (10 long) on tarsi II – IV modified. Length of legs I, II, III and IV 285, 242, 251 and 279 respectively.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA30FFFBB3A6F9D1FF010334.taxon	materials_examined	Material examined. Holotype. Female. NE Iran, North Khorasan province, Esfarayen (36 º 55 ' 29 " N, 57 º 44 ' 08 " E), altitude 1,471 m, ex Scarabaeus transcaspicus, 26 June 2007 [deposited in Acarological Collection, Department of Entomology, Faculty of Agriculture, Tarbiat Modares University, Tehran, Iran (AETMU)]. Paratypes. 27 females, 7 males, 14 deutonymphs, 1 protonymph, same locality and host data as holotype (deposited in AETMU); 2 females, 1 male, 1 deutonymph, same data as holotype [deposited in Museum of Zoology, Navarra University, Pamplona, Spain (MZUNAV)]; 2 females: NE Iran, Razavi Khorasan province, Sarakhs, Bazangan Lake (36 º 18 ' 549 " N, 60 º 29 ' 061 " E), altitude 858 m, ex Copris hispanus, 5 May 2007 (deposited in AETMU); 31 females, 5 males and 2 deutonymphs, same locality data as previous specimens, but from Scarabaeus transcaspicus (deposited in AETMU); 1 female, 1 male, with same locality and host data as latter paratypes [deposited in Museum of Zoology, Tehran University, Karaj, Iran (MZTU)].	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA30FFFBB3A6F9D1FF010334.taxon	etymology	Etymology. The species epithet “ persicus ” refers to the Latin name of Iran, the country of origin of this species.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3BFFFEB3A6FA19FA67044C.taxon	diagnosis	Diagnosis. Adult dorsum with 32 pairs of setae, variable in length; setae j 2, z 2, s 2, s 6, r 2, r 3, r 5, R 1, R 3 - R 5 and ventral JV 5 conspicuously longer, eight to ten times longer than J 5 in female, and more than four times in male; other setae short, opisthonotal shield setae shorter setae; seta z 2 longer than s 2 and about four times longer than j 3; j 2 more than six times longer than seta z 1. Sternal setae st 2 longer than distances between their bases and that of st 3; male setae on dorsal shield (except j 1, j 2, z 2) longer than those of female. Peritreme extends to anterior margin of coxa II. Anal shield subtriangular, with anterior margin slightly convex.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3BFFFEB3A6FA19FA67044C.taxon	description	Description. Female. Idiosoma. Length: 564 – 580, width: 370 – 406 (n = 5). Dorsal idiosoma (Fig. 16). Dorsal shield ovoid, 549 – 561 long, 326 – 361 wide; with striations antero-laterally, otherwise covered with fine dense punctations; lateral incisions reaching level of setae z 2 or s 2. Dorsal setae j 2 (84), z 2 (98), s 2 (71), s 6 (72), r 2 (81), r 3 (79), r 5 (76) and R 5 (78) long; other dorsal setae on shield short (13 – 24), R 1 and R 3 (35 – 39), R 4 (56) and sometimes S 4 (11) on soft cuticle. Dorsal adenotaxy and poroidotaxy typical of genus. Ventral idiosoma (Fig. 20). Tritosternum with smooth base (35) and pilose laciniae (70 long). Sternal shield (157 long, 111 wide) finely granulated but anteriorly reticulate-striate, anterior and posterior borders irregular; with three pairs of setae st 1, st 2 and st 3 (74, 67, 67 long respectively) and two pairs of lyrifissures, first pair almost positioned longitudinally; st 4 (72) and third pair of sternal lyrifissures inserted on minute metasternal plates (17 long, 11 wide) flanked by endopodal shields between coxae III and IV. Genital shield 157 long, 91 wide posterior to base of genital setae, st 5 (57). Five pairs of simple setae inserted on opisthogaster (JV 1, JV 2, JV 5, ZV 2 and ZV 3 48, 30, 113, 42, and 39 long respectively). Anal shield subtriangular, 106 long, 102 wide, with slightly convex anterior margin; postanal seta (52) longer than para-anal setae (30). Peritremes (239 long) extending from stigmata located between coxae III – IV to anterior margin of coxae II. Gnathosoma. Hypostomal setae h 1, h 2, h 3 and capitular setae simple and smooth (21, 32, 17, 20 long respectively). Fixed cheliceral digit (40 long) with a large median tooth and 2 small teeth, 1 subterminal and 1 proximal. Pilus dentilis setiform and fine; movable digit (52 long) with 1 median and 1 smaller apical tooth. Length and width of chelicera 140 and 40 respectively. Legs. Leg chaetotaxy typical of genus; av - 1 (17 long) thickened and tri-dentate terminally. Length of legs I – IV, 377, 342, 343 and 425 respectively. Male. Idiosoma. Length: 628 – 646, width: 400 – 423 (n = 4). Dorsal idiosoma (Fig. 17). Dorsal shield 596 – 606 long, 342 – 361 wide, with shape, adenotaxy and poroidotaxy similar to those of female, except for lateral incisions which may not reach level of setae z 2; setae s 2, r 2, r 3 37 – 39 long; the longest setae on dorsal shield are j 2 (63) and z 2 (66); length of other dorsal shield setae relatively longer than those of female: podonotal setae 33 – 37 long (except j 1 and z 1 12 long), opisthonotal setae 17 – 25 (J 5 = 17) long; dorsal setae on soft cuticle: r 5 (37), s 6 (29) and R setae (17 – 23) long. Ve n t r a l idiosoma (Fig. 21). Tritosternal base 19 long, laciniae 84 long; sternogenital shield (273 long, 118 wide at level of st 2 and 62 wide at level of st 5), with setae st 1 (36), st 2 (34), st 3 (40), st 4 (34) and st 5 (38) thickened (Fig. 22 a) as posterior setae on coxae I – III, reticulate anteriorly, rest evenly granulate, except for area between st 5, which is densely punctate. Genital opening on anterior margin of shield, 38 wide. Ventral idiosomal adenotaxy and poroidotaxy as in female; JV 5 (81) longest, almost five times longer than J 5, JV 1 and JV 2 (Fig. 22 b) (28 and 31 respectively) longer than ZV 2 (24) and ZV 3 (13). Peritremes longer than those of female, extending to posterior border of coxae I (250 long). Anal shield as for female (105 long, 100 wide); para-anal setae (27) slightly shorter than post-anal seta (31). Ventral platelets present. Gnathosoma. Hypostomal (h 1, h 2, h 3) and capitular setae simple and smooth, 23, 36, 17 and 18 long respectively. Chelicera 140 long, 40 wide; movable chela (50) two times longer than fixed chela (24) (Fig. 24); spermatodactyl 18 long; hyaline process indistinct. Legs. Seta av - 1 (12 long) and pv - 1 on tarsus II – IV modified (Fig. 18); length of legs approximately similar to those of female, leg I – IV: 385, 354, 346 and 438 respectively. Deutonymph. Idiosoma. Length: 517 – 537, width: 308 – 354 (n = 3). Dorsal idiosoma (Fig. 25). Dorsal shield ovoid, 462 – 485 long, 243 – 264 wide, with 24 pairs of setae; dorsal setae j 1, j 2, z 1, z 2, S 2 and J 5, 19, 35, 11, 34, 22 and 9 long respectively, other dorsal setae 9 – 17 long; lateral setae r 2, r 3, r 5, s 6, R 1, R 3, R 4 and R 5 on soft cuticle and 32, 34, 17, 38, 30, 26, 27 and 30 long respectively. Dorsal adenotaxy and poroidotaxy similar to those of female. Ve n t r a l idiosoma. Tritosternal base (30 long) and laciniae (90 long) typical of genus. Sternal shield with endopodal extensions between coxae I – II but not between II – III and III – IV, and with four pairs of setae st 1, st 2, st 3, st 4 (28, 29, 29, 21 long respectively) and three pairs of lyrifissures; setae st 5 (21) and paragenital lyrifissures inserted on soft cuticle at level of posterior margin of shield. Opisthogaster with five pairs of ventral setae JV 1, JV 2, JV 5, ZV 2 and ZV 3 (21, 23, 48, 19 and 14 long respectively). Peritremes 146 long. Anal shield subtriangular, 83 long, 94 wide. Gnathosoma. Gnathotectum, chelicerae, salivary stylets, corniculi, deutosternal structures and palpi similar to those of adults. Chelicera 127 long, 44 width; fixed chela (34) shorter than movable chela (44). Hypostomal setae h 1, h 2, h 3 and capitular setae 23, 34, 17 and 18 long respectively. Legs. Pretarsi, claws, and chaetotaxy of legs I – IV as in female; seta av - 1 (13 long) modified. Length of legs I – IV 352, 318, 327 and 377 respectively.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3BFFFEB3A6FA19FA67044C.taxon	materials_examined	Material examined. Holotype. Female. N Iran, Mazandaran province, Galugah region, ex Scarabaeus transcaspicus (deposited in AETMU). Paratypes. 8 females, 4 males, 1 deutonymph, same data as holotype (deposited in AETMU); 2 females, 1 male, same data as holotype (deposited in MZUNAV); 1 female, same data as holotype (deposited in MZTU); 14 females, 1 male, 1 deutonymph, E Iran, Razavi Khorasan province, Torbat Heydarie, Feiz-Abad (34 º 57 ' 23 " N, 58 º 45 ' 18 " E), altitude 895 m, ex Scarabaeus transcaspicus, 5 May 2007 (deposited in AETMU); 5 females, E Iran, Razavi Khorasan province, Sarakhs, Bazangan Lake (36 º 18 ' 33 " N, 60 º 29 ' 37 " E), altitude 858 m, ex Scarabaeus transcaspicus, 5 May 2007 (deposited in AETMU).	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3BFFFEB3A6FA19FA67044C.taxon	etymology	Etymology. The species epithet “ khorasanicus ” refers to the province in which specimens of this species were collected.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3BFFFEB3A6FA19FA67044C.taxon	discussion	Remarks. Differences between this species and related species are highlighted in the following key to females, in which M. guyimingi (Ma) comb. nov. is also included:	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFFEB3A6FD11FCA805CA.taxon	materials_examined	Type species: Cryptoseius petrova Makarova, 1998: 115.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFFEB3A6FD11FCA805CA.taxon	diagnosis	Diagnosis. Dorsal shield of adult not covering whole body, with 22 – 23 pairs of setae: setae j 1 - j 6, J 2, J 4 - J 5, z 1, z 2, z 4, z 5, Z 3 - Z 4, s 2, s 4, s 5, S 3 - S 5, r 2 and r 3 (s 2, r 2 and r 3 on laterodorsal elements); setae s 6, r 5, R 1, R 3 - R 5 on soft cuticle. Female with sternal setae st 4 on soft cuticle and lyrifissures iv 3 on posterior margin of sternal shield. Deutosternal groove with eight rows of denticles.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFE3B3A6FBACFB4300C4.taxon	diagnosis	Diagnosis. Adult female and male with 28 – 29 pairs of dorsal setae (17 podonotal and 11 – 12 opisthonotal), 19 – 20 pairs inserted on dorsal shield, three pairs on laterodosal elements (s 2, r 2 and r 3) and six pairs on lateral membranous integument; setae j 3, j 4, j 6, z 2, z 4, s 4, s 5, s 6, S 3, S 4 and S 5 clearly longer, can be 12 times longer than J 5; seta JV 5 more than 10 times longer than J 5; setae z 2 approximately four times longer than setae s 2; dorsum of male similar to that of female. Opisthonotal setae J 2, J 3, Z 2 and Z 5 absent. Peritreme extending from stigmata between coxae III – IV to middle region of coxae II.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFE3B3A6FBACFB4300C4.taxon	description	Description. Female. Idiosoma. Length: 623 – 674, width: 414 – 476 (n = 5). Dorsal idiosoma (Fig. 27). Dorsal shield ovoid (517 – 535 long, 302 – 324 wide) with posterior margin rounded; shield punctate, with a few antero-lateral striate; anterolateral incisions of dorsal shield reach beyond level of z 2. Dorsal shield with 23 pairs of simple, smooth setae, heterogeneous in length; setae j 3 (127 – 137 long), j 4 (112 – 129), j 6 (87 – 109), z 2 (115), z 4 (141), s 4 (146), s 5 (145), s 6 (124), S 2 (136), S 3 (141) and S 5 (122) elongated; j 1 (27), j 2 (39), j 5 (17), J 1 (present or absent) = J 4 = J 5 (11), z 1 (13), z 5 (12), Z 3 and Z 4 as long as setae J series; s 2 (22), r 2 (56) and r 3 (45) on laterodosal elements; setae r 5 (38 – 54), s 6, R 1 (38) and R 3 - R 5 (66 – 69) on lateral soft cuticle. Dorsal shield with 20 pairs of pore-like structures (eight podonotal, 12 opisthonotal), of which six (three podonotal, three opisthonotal) appear secretory (gland pores) and 14 (five podonotal, nine opisthonotal) non-secretory (lyrifissures); poroid Rp 3, idS 4 and idS 5 on soft cuticle. Ventral idiosoma (Fig. 31). Tritosternum typical of genus (base 40 long, laciniae 90). Sternal shield (142 long, 136 wide at level of st 2, 175 between coxae II and III) with a dense fine granulation; anterior and posterior margins irregular, three pairs of setae (st 1 and st 2 37 long and st 3 51) and three pairs of lyrifissures (the first and second diagonally oriented, third on posterior margin horizontally oriented) on the shield; st 4 (54) on soft cuticle, flanked by endopodal shields between coxae III and IV. Genital shield 144 long, 80 wide at level st 5 (64) located laterally on shield; positions of iv 5 and gv 2 as for genus. Opisthogaster with five pairs of simple setae, JV 1 (42), JV 2 (51), JV 5 (129), ZV 2 (43) and ZV 3 (32). Anal shield pear-shaped, 106 long, ca 83 wide, para-anal setae 18 long, postanal seta 31; cribrum well developed. Metapodal and parapodal plates present. Peritrematal plates with gland pore gp 1 and one lyrifissure ip 1 (Fig. 31); poststigmatic gland pores gp 2 inserted on soft cuticle; peritremes extending to middle of coxae II, 172 long. Spermathecal apparatus without distinctly sclerotized structures, but usually with a discernible tube from solenostome on base of legs III to a narrow calyx. Gnathosoma. Subcapitulum and gnathotectum (Fig. 28 a) typical of genus. Hypostomal setae simple and smooth, h 1 (25), h 2 (37), h 3 (18), and (20) long. Chelicera 148 long; movable chela 50 long and fixed chela 36; fixed digit with a large median and two small teeth, one subterminal and other one proximal. Pilus dentilis setiform and fine; dorsal seta small and near proximal lyrifissure; movable digit with one median tooth, with fringed hyaline process at its base. Legs. Leg chaetotaxy typical of genus. Ventral modified setae av - 1 on tarsi II – IV (Fig. 29 a) 16 long. Length of legs as follows: leg I, 452 long; II, 397; III, 416 and IV, 541. Male. Idiosoma. Dorsal shield: 465 – 503 long, 261 – 300 wide (n = 2). Dorsal idiosoma (Fig. 30). Dorsal shield (460 – 471 long, 248 – 269 wide) densely covered with punctations. Dorsal chaetotaxy, adenotaxy and poroidotaxy similar to those of female. Lateral incisions reduced, reaching base of setae r 2 or r 3. Ventral idiosoma (Fig. 32). Base of tritosternum 19 long, laciniae pilose and free 64 long; sternogenital shield 250 long, 85 wide at level of setae st 2 and 64 at level of setae st 5, anteriorly densely covered with fine granulations, posteriorly, between setae st 5, with dense punctuations, extending beyond the posterior border of coxae IV and fused with endopodal elements between coxae I – II, II – III and III – IV; setae st 1 and st 2 (20 long), st 3 (22), st 4 (26) and st 5 (33) thickened; genital opening on the shield (21 wide), anterior to setae st 1. Ventral idiosomal adenotaxy and poroidotaxy similar to those of female; JV 5 (102) is longest ventral setae, JV 1 (25), JV 2 (23), ZV 2 (20) and ZV 3 (15); anal shield pear-shaped, 92 long, 62 wide. Parapodal and metapodal plates present. Gnathosoma. Gnathotectum triangular, slightly dentate (Fig. 28 b); subcapitulum typical of genus. Length of hypostomal and capitular setae: h 1, 20; h 2, 33; h 3, 13 and c, 16. Chelicera 115 long and 35 wide, movable chelae with two teeth (43 long) and fixed chela with a median tooth and 35 long; spermatodactyl 20 long; hyaline process indistinct. Legs. Legs chaetotaxy typical of genus. Length of legs I – IV: 317, 300, 304 and 402 respectively. Seta av - 1 on tarsi II – IV modified and nearly conical, 11 long (Fig. 29 b). Deutonymph. Idiosoma. Length: 411 – 458, width: 288 – 326 (n = 2). Dorsal idiosoma (Fig. 26). Dorsal shield (395 – 427 length, 200 – 231 width) with 21 – 22 pairs of setae; setae j 1, j 2, j 4, j 5 and j 6 with 10 – 18 long, j 3 longer (26); setae z 1, z 2, z 5, s 2, s 4 and s 5, respectively, 11, 31, 11, 13, 45 and 62 long; soft cuticle with r 2, r 3, r 5, s 6 (15, 15, 19, 57 long respectively) in podonotal region and R 1, R 3 - R 5, on opisthonotal region (18, 16, 14, 61 long respectively). Dorsal idiosomal complement of lyrifissures and gland-pores similar to those of female. Ventral idiosoma. Tritosternal base 29 long; laciniae 83 long; sternal shield (188 long, 63 width at level of setae st 2), with four pairs of setae (st 1 = 18, st 2 = 21, st 3 = 21, st 4 = 21) and three pairs of lyrifissures; setae st 5 (19) and paragenital lyrifissures inserted on soft cuticle at level of posterior margin of sternal shield. Endopodal shields between coxae II – III and III – IV, and parapodal present. Peritremes 120 long. Anal shield (81 long and 74 wide) pear-shaped, para-anal setae 19 and postanal seta 24 long; adanal gland pores inserted as in adults; cribrum well developed. Opisthogaster with five pairs of ventral setae: JV 1, JV 2, ZV 2, ZV 3 (15 – 18 long) and JV 5 (61). Gnathosoma. Chelicera 107 long and 32 wide; movable chela 37 and fixed chela 28 long. Hypostomal setae h 1 (18), h 2 (38), h 3 (12) and capitular setae (13) smooth and simple. Legs. Pretarsi, claws, and chaetotaxy of legs I – IV similar to adult female; av - 1 thickened and tri-dentate, 12 long; length of legs I – IV, 300, 269, 258 and 335, respectively.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFE3B3A6FBACFB4300C4.taxon	materials_examined	Material examined. Holotype. Female: NE Iran, Razavi Khorasan province, Torbat Heydarie, Feiz-Abad (34 º 57 ' 23 " N, 58 º 45 ' 18 " E), altitude 895 m, ex Scarabaeus transcaspicus, 5 May 2007 (deposited in AETMU). Paratypes. 27 females, 3 males, 5 deutonymphs, same data as holotype (deposited in AETMU); 2 females, 1 male, same data as holotype (deposited in MZUNAV); 1 female, same data as holotype (deposited in MZTU).	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
038987C3BA3EFFE3B3A6FBACFB4300C4.taxon	etymology	Etymology. The species is named in honour of the famous philosopher, astronomer, mathematician and poet of Iran, Omar Khayyam Neishaburi. Notes. Cryptoseius khayyami differs from the type species C. petrovae Makarova, 1998 in the number of long dorsal setae in the j series in adults, four pairs in C. petrovae (j 3 - j 6) and three pairs in C. khayyami sp. nov. (j 3, j 4 and j 6) and in the relative length of several dorsal setae in deutonymphs: r 5> R 1, R 1 = 1 / 3 s 6, S 4 <S 5 = ½ S 3 in C. petrovae and r 5 <R 1, R 1 = ½ s 6, S 4 = S 5 = S 3 in C. khayyami sp. nov.	en	Kazemi, Shahrooz, Moraza, Mar A L., Kamali, Karim, Saboori, Alireza (2008): A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa 1852: 1-20, DOI: 10.5281/zenodo.183438
